chr5 : 169,528,729 169,529,472
743 bp 127 TFs 2 linked genes
This 743 bp open chromatin element is linked to SPDL1 and SLIT3 and is bound by 127 transcription factors.
Linked Genes
2 genes
Gene Expression Dist. to TSS Distance Link type
SPDL1 54.7 kb Distal Multiome
SLIT3 227.9 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr5:169,523,729 – 169,534,472
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
127 transcription factors
Source
Cell type
ATF2 4 datasets
Motif DE_36h DE_36h-ATF2_MA1632.2 10 bp overlap
Motif DE_48h DE_48h-ATF2_MA1632.2 10 bp overlap
Motif DE_60h DE_60h-ATF2_MA1632.2 10 bp overlap
Motif DE_72h DE_72h-ATF2_MA1632.2 10 bp overlap
Arid3a 1 dataset
Motif DE_60h DE_60h-Arid3a_MA0151.1 6 bp overlap
BACH2 4 datasets
Motif DE_36h DE_36h-BACH2_MA1470.2 19 bp overlap
Motif DE_48h DE_48h-BACH2_MA1470.2 19 bp overlap
Motif DE_60h DE_60h-BACH2_MA1470.2 19 bp overlap
Motif DE_72h DE_72h-BACH2_MA1470.2 19 bp overlap
BARX1 2 datasets
Motif DE_48h DE_48h-BARX1_MA0875.2 6 bp overlap
Motif DE_60h DE_60h-BARX1_MA0875.2 6 bp overlap
BRD4 1 dataset
ChIP HAP1 GSE108387.BRD4.HAP1 517 bp overlap
BSX 2 datasets
Motif DE_48h DE_48h-BSX_MA0876.2 6 bp overlap
Motif DE_60h DE_60h-BSX_MA0876.2 6 bp overlap
CEBPB 1 dataset
ChIP HFOB_DIFF GSE82295.CEBPB.HFOB_DIFF 272 bp overlap
Crx 1 dataset
Motif DE_60h DE_60h-Crx_MA0467.3 6 bp overlap
DLX1 2 datasets
Motif DE_48h DE_48h-DLX1_MA0879.3 6 bp overlap
Motif DE_60h DE_60h-DLX1_MA0879.3 6 bp overlap
DLX6 2 datasets
Motif DE_48h DE_48h-DLX6_MA0882.2 6 bp overlap
Motif DE_60h DE_60h-DLX6_MA0882.2 6 bp overlap
DUX4 5 datasets
Motif DE_36h DE_36h-DUX4_MA0468.1 11 bp overlap
Motif DE_48h DE_48h-DUX4_MA0468.1 11 bp overlap
Motif DE_60h DE_60h-DUX4_MA0468.1 11 bp overlap
Motif DE_72h DE_72h-DUX4_MA0468.1 11 bp overlap
ChIP WA01 GSE94322.DUX4.WA01 194 bp overlap
Dlx3 2 datasets
Motif DE_48h DE_48h-Dlx3_MA0880.2 6 bp overlap
Motif DE_60h DE_60h-Dlx3_MA0880.2 6 bp overlap
Dlx4 2 datasets
Motif DE_48h DE_48h-Dlx4_MA0881.2 6 bp overlap
Motif DE_60h DE_60h-Dlx4_MA0881.2 6 bp overlap
Dmbx1 1 dataset
Motif DE_60h DE_60h-Dmbx1_MA0883.2 10 bp overlap
EN2 2 datasets
Motif DE_48h DE_48h-EN2_MA0642.3 7 bp overlap
Motif DE_60h DE_60h-EN2_MA0642.3 7 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 199 bp overlap
ERF::SREBF2 5 datasets
Motif DE_36h DE_36h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_48h DE_48h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_48h DE_48h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_60h DE_60h-ERFSREBF2_MA1939.2 16 bp overlap
Motif DE_72h DE_72h-ERFSREBF2_MA1939.2 16 bp overlap
ERG 1 dataset
ChIP aortic-endothelial-cell_D40 GSE139377.ERG.aortic-endothelial-cell_D40 204 bp overlap
ETV2 1 dataset
Motif DE_48h DE_48h-ETV2_MA0762.2 9 bp overlap
FIGLA 4 datasets
Motif DE_36h DE_36h-FIGLA_MA0820.2 6 bp overlap
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
Motif DE_72h DE_72h-FIGLA_MA0820.2 6 bp overlap
FOS 4 datasets
Motif DE_36h DE_36h-FOS_MA1951.2 13 bp overlap
Motif DE_48h DE_48h-FOS_MA1951.2 13 bp overlap
Motif DE_60h DE_60h-FOS_MA1951.2 13 bp overlap
Motif DE_72h DE_72h-FOS_MA1951.2 13 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 355 bp overlap
FOXA2 3 datasets
ChIP DE DE-FOXA2-1 743 bp overlap
ChIP DE DE-FOXA2-2 642 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 341 bp overlap
FOXC1 1 dataset
Motif DE_48h DE_48h-FOXC1_MA0032.2 11 bp overlap
FOXC2 1 dataset
Motif DE_48h DE_48h-FOXC2_MA0846.2 11 bp overlap
FOXD2 1 dataset
Motif DE_48h DE_48h-FOXD2_MA0847.4 11 bp overlap
FOXE1 1 dataset
Motif DE_48h DE_48h-FOXE1_MA1487.3 12 bp overlap
FOXS1 1 dataset
Motif DE_48h DE_48h-FOXS1_MA2118.1 8 bp overlap
Foxl2 1 dataset
Motif DE_48h DE_48h-Foxl2_MA1607.2 10 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 612 bp overlap
ChIP DE DE-GATA4-2 712 bp overlap
ChIP foregut GSE117136.GATA4.foregut 555 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 594 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 482 bp overlap
GATA6 14 datasets
ChIP AGS GSE51705.GATA6.AGS 156 bp overlap
ChIP DE DE-GATA6-1 650 bp overlap
ChIP DE DE-GATA6-2 711 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 585 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 661 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 643 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 705 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 741 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 692 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 358 bp overlap
ChIP foregut GSE117136.GATA6.foregut 673 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 602 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 480 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 479 bp overlap
GBX1 2 datasets
Motif DE_48h DE_48h-GBX1_MA0889.2 7 bp overlap
Motif DE_60h DE_60h-GBX1_MA0889.2 7 bp overlap
GBX2 2 datasets
Motif DE_48h DE_48h-GBX2_MA0890.2 6 bp overlap
Motif DE_60h DE_60h-GBX2_MA0890.2 6 bp overlap
GRHL1 2 datasets
Motif DE_48h DE_48h-GRHL1_MA0647.2 10 bp overlap
Motif DE_60h DE_60h-GRHL1_MA0647.2 10 bp overlap
GSC 1 dataset
Motif DE_60h DE_60h-GSC_MA0648.2 6 bp overlap
GSC2 1 dataset
Motif DE_60h DE_60h-GSC2_MA0891.2 6 bp overlap
HESX1 2 datasets
Motif DE_48h DE_48h-HESX1_MA0894.2 6 bp overlap
Motif DE_60h DE_60h-HESX1_MA0894.2 6 bp overlap
HOXA7 2 datasets
Motif DE_48h DE_48h-HOXA7_MA1498.3 6 bp overlap
Motif DE_60h DE_60h-HOXA7_MA1498.3 6 bp overlap
Hand1 1 dataset
Motif DE_48h DE_48h-Hand1_MA2123.1 9 bp overlap
Hnf1A 1 dataset
Motif DE_48h DE_48h-Hnf1A_MA1991.2 10 bp overlap
IKZF1 1 dataset
Motif DE_48h DE_48h-IKZF1_MA1508.2 8 bp overlap
IKZF2 1 dataset
Motif DE_48h DE_48h-IKZF2_MA2326.1 6 bp overlap
IRF7 1 dataset
Motif DE_48h DE_48h-IRF7_MA0772.2 13 bp overlap
ISL2 1 dataset
Motif DE_48h DE_48h-ISL2_MA0914.2 6 bp overlap
Irf1 1 dataset
Motif DE_48h DE_48h-Irf1_MA0050.4 11 bp overlap
JUN 3 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 561 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 420 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 602 bp overlap
KDM1A 3 datasets
ChIP pancreatic-progenitor_GT GSE104840.KDM1A.pancreatic-progenitor_GT 350 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 213 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 413 bp overlap
LBX1 2 datasets
Motif DE_48h DE_48h-LBX1_MA0618.2 7 bp overlap
Motif DE_60h DE_60h-LBX1_MA0618.2 7 bp overlap
LBX2 2 datasets
Motif DE_48h DE_48h-LBX2_MA0699.2 6 bp overlap
Motif DE_60h DE_60h-LBX2_MA0699.2 6 bp overlap
LHX2 2 datasets
Motif DE_48h DE_48h-LHX2_MA0700.3 6 bp overlap
Motif DE_60h DE_60h-LHX2_MA0700.3 6 bp overlap
LHX6 1 dataset
Motif DE_48h DE_48h-LHX6_MA0658.2 8 bp overlap
LHX9 2 datasets
Motif DE_48h DE_48h-LHX9_MA0701.3 7 bp overlap
Motif DE_60h DE_60h-LHX9_MA0701.3 7 bp overlap
MGA::EVX1 2 datasets
Motif DE_48h DE_48h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_60h DE_60h-MGAEVX1_MA1960.2 11 bp overlap
MSX1 2 datasets
Motif DE_48h DE_48h-MSX1_MA0666.3 6 bp overlap
Motif DE_60h DE_60h-MSX1_MA0666.3 6 bp overlap
MSX2 2 datasets
Motif DE_48h DE_48h-MSX2_MA0708.3 6 bp overlap
Motif DE_60h DE_60h-MSX2_MA0708.3 6 bp overlap
Msx3 2 datasets
Motif DE_48h DE_48h-Msx3_MA0709.2 6 bp overlap
Motif DE_60h DE_60h-Msx3_MA0709.2 6 bp overlap
NANOG 2 datasets
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 393 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 301 bp overlap
NBN 1 dataset
ChIP MCF-10A GSE93038.NBN.MCF-10A 285 bp overlap
NFIA 2 datasets
Motif DE_48h DE_48h-NFIA_MA0670.2 6 bp overlap
Motif DE_60h DE_60h-NFIA_MA0670.2 6 bp overlap
NFIX 2 datasets
Motif DE_48h DE_48h-NFIX_MA0671.2 6 bp overlap
Motif DE_60h DE_60h-NFIX_MA0671.2 6 bp overlap
NKX2-2 2 datasets
Motif DE_48h DE_48h-NKX2-2_MA1645.2 8 bp overlap
Motif DE_60h DE_60h-NKX2-2_MA1645.2 8 bp overlap
NKX2-3 1 dataset
Motif DE_48h DE_48h-NKX2-3_MA0672.2 8 bp overlap
NKX2-4 1 dataset
Motif DE_48h DE_48h-NKX2-4_MA2003.2 8 bp overlap
NKX2-8 1 dataset
Motif DE_48h DE_48h-NKX2-8_MA0673.2 8 bp overlap
NR1D1 4 datasets
Motif DE_36h DE_36h-NR1D1_MA1531.2 14 bp overlap
Motif DE_48h DE_48h-NR1D1_MA1531.2 14 bp overlap
Motif DE_60h DE_60h-NR1D1_MA1531.2 14 bp overlap
Motif DE_72h DE_72h-NR1D1_MA1531.2 14 bp overlap
NR1D2 4 datasets
Motif DE_36h DE_36h-NR1D2_MA1532.2 15 bp overlap
Motif DE_48h DE_48h-NR1D2_MA1532.2 15 bp overlap
Motif DE_60h DE_60h-NR1D2_MA1532.2 15 bp overlap
Motif DE_72h DE_72h-NR1D2_MA1532.2 15 bp overlap
Nkx3-1 1 dataset
Motif DE_48h DE_48h-Nkx3-1_MA0124.3 7 bp overlap
Nkx3-2 1 dataset
Motif DE_48h DE_48h-Nkx3-2_MA0122.4 10 bp overlap
Nobox 2 datasets
Motif DE_48h DE_48h-Nobox_MA0125.2 6 bp overlap
Motif DE_60h DE_60h-Nobox_MA0125.2 6 bp overlap
Nr2F6 4 datasets
Motif DE_36h DE_36h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_48h DE_48h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_60h DE_60h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_72h DE_72h-Nr2F6_MA0728.1 15 bp overlap
ONECUT1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 409 bp overlap
OTX1 1 dataset
Motif DE_60h DE_60h-OTX1_MA0711.2 6 bp overlap
PDX1 3 datasets
ChIP hESC GSE58685.PDX1.hESC 156 bp overlap
ChIP hiPSC GSE125768.PDX1.hiPSC 456 bp overlap
ChIP pancreatic-progenitor GSE117136.PDX1.pancreatic-progenitor 288 bp overlap
PITX1 1 dataset
Motif DE_60h DE_60h-PITX1_MA0682.3 6 bp overlap
PITX2 1 dataset
Motif DE_60h DE_60h-PITX2_MA1547.2 8 bp overlap
PITX3 1 dataset
Motif DE_60h DE_60h-PITX3_MA0714.2 6 bp overlap
POU2F3 1 dataset
Motif DE_48h DE_48h-POU2F3_MA0627.3 9 bp overlap
POU5F1 1 dataset
Motif DE_48h DE_48h-POU5F1_MA1115.2 7 bp overlap
PRRX2 2 datasets
Motif DE_48h DE_48h-PRRX2_MA0075.4 7 bp overlap
Motif DE_60h DE_60h-PRRX2_MA0075.4 7 bp overlap
RAX 2 datasets
Motif DE_48h DE_48h-RAX_MA0718.2 6 bp overlap
Motif DE_60h DE_60h-RAX_MA0718.2 6 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 146 bp overlap
RHOXF1 1 dataset
Motif DE_60h DE_60h-RHOXF1_MA0719.2 6 bp overlap
RNF2 1 dataset
ChIP fibroblast GSE139053.RNF2.fibroblast 151 bp overlap
RUNX3 4 datasets
Motif DE_36h DE_36h-RUNX3_MA0684.3 8 bp overlap
Motif DE_48h DE_48h-RUNX3_MA0684.3 8 bp overlap
Motif DE_60h DE_60h-RUNX3_MA0684.3 8 bp overlap
Motif DE_72h DE_72h-RUNX3_MA0684.3 8 bp overlap
Rarb 5 datasets
Motif DE_36h DE_36h-Rarb_MA0857.1 16 bp overlap
Motif DE_48h DE_48h-Rarb_MA0857.1 16 bp overlap
Motif DE_48h DE_48h-Rarb_MA0858.1 17 bp overlap
Motif DE_60h DE_60h-Rarb_MA0857.1 16 bp overlap
Motif DE_72h DE_72h-Rarb_MA0857.1 16 bp overlap
SMAD2 1 dataset
Motif DE_48h DE_48h-SMAD2_MA1964.2 6 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 680 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 714 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 611 bp overlap
SMAD2_3 5 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 488 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 639 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 567 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 517 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 603 bp overlap
SMAD4 2 datasets
ChIP Caco-2 GSE112946.SMAD4.Caco-2 169 bp overlap
ChIP endoderm GSE29422.SMAD4.endoderm 215 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 409 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 301 bp overlap
SOX14 1 dataset
Motif DE_48h DE_48h-SOX14_MA1562.2 9 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 305 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 604 bp overlap
SOX8 1 dataset
Motif DE_48h DE_48h-SOX8_MA0868.3 7 bp overlap
SP5 1 dataset
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
SREBF1 4 datasets
Motif DE_36h DE_36h-SREBF1_MA0829.3 10 bp overlap
Motif DE_48h DE_48h-SREBF1_MA0829.3 10 bp overlap
Motif DE_60h DE_60h-SREBF1_MA0829.3 10 bp overlap
Motif DE_72h DE_72h-SREBF1_MA0829.3 10 bp overlap
SRY 1 dataset
Motif DE_48h DE_48h-SRY_MA0084.2 7 bp overlap
Sox5 1 dataset
Motif DE_48h DE_48h-Sox5_MA0087.3 8 bp overlap
Spz1 4 datasets
Motif DE_36h DE_36h-Spz1_MA0111.1 11 bp overlap
Motif DE_48h DE_48h-Spz1_MA0111.1 11 bp overlap
Motif DE_60h DE_60h-Spz1_MA0111.1 11 bp overlap
Motif DE_72h DE_72h-Spz1_MA0111.1 11 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 190 bp overlap
Tfcp2l1 1 dataset
Motif DE_60h DE_60h-Tfcp2l1_MA0145.2 14 bp overlap
Vdr 4 datasets
Motif DE_36h DE_36h-Vdr_MA0693.4 7 bp overlap
Motif DE_48h DE_48h-Vdr_MA0693.4 7 bp overlap
Motif DE_60h DE_60h-Vdr_MA0693.4 7 bp overlap
Motif DE_72h DE_72h-Vdr_MA0693.4 7 bp overlap
ZBTB24 1 dataset
Motif DE_48h DE_48h-ZBTB24_MA2330.1 10 bp overlap
ZEB1 4 datasets
Motif DE_36h DE_36h-ZEB1_MA0103.4 6 bp overlap
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
Motif DE_72h DE_72h-ZEB1_MA0103.4 6 bp overlap
ZFP14 1 dataset
Motif DE_48h DE_48h-ZFP14_MA1972.1 15 bp overlap
ZFP42 1 dataset
Motif DE_36h DE_36h-ZFP42_MA1651.2 13 bp overlap
ZNF135 1 dataset
Motif DE_48h DE_48h-ZNF135_MA1587.1 14 bp overlap
ZNF16 2 datasets
Motif DE_48h DE_48h-ZNF16_MA1654.2 21 bp overlap
Motif DE_72h DE_72h-ZNF16_MA1654.2 21 bp overlap
ZNF189 1 dataset
Motif DE_48h DE_48h-ZNF189_MA1725.2 9 bp overlap
ZNF213 1 dataset
Motif DE_36h DE_36h-ZNF213_MA2121.1 12 bp overlap
ZNF263 1 dataset
Motif DE_48h DE_48h-ZNF263_MA0528.3 7 bp overlap
ZNF28 1 dataset
ChIP HEK293T GSE78099.ZNF28.HEK293T 189 bp overlap
ZNF282 1 dataset
Motif DE_48h DE_48h-ZNF282_MA1154.2 15 bp overlap
ZNF341 1 dataset
Motif DE_48h DE_48h-ZNF341_MA1655.2 8 bp overlap
ZNF449 1 dataset
Motif DE_36h DE_36h-ZNF449_MA1656.2 10 bp overlap
Zic3 1 dataset
Motif DE_48h DE_48h-Zic3_MA0697.3 7 bp overlap