chr4 : 170,004,428 170,005,171
743 bp 112 TFs 4 linked genes
This 743 bp open chromatin element is linked to 4 target genes and is bound by 112 transcription factors.
Linked Genes
4 genes
Gene Expression Dist. to TSS Distance Link type
MFAP3L 21.7 kb Distal Multiome
AADAT 85.5 kb Distal Multiome
LINC01612 221.8 kb Distal Multiome
HPF1 246.8 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:169,999,428 – 170,010,171
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
112 transcription factors
Source
Cell type
BARX1 1 dataset
Motif DE_48h DE_48h-BARX1_MA0875.2 6 bp overlap
BCL11A 3 datasets
Motif DE_48h DE_48h-BCL11A_MA2324.1 7 bp overlap
Motif DE_60h DE_60h-BCL11A_MA2324.1 7 bp overlap
Motif DE_72h DE_72h-BCL11A_MA2324.1 7 bp overlap
BRD4 2 datasets
ChIP HCC1395_JQ1 GSE63581.BRD4.HCC1395_JQ1 203 bp overlap
ChIP cortical-interneuron GSE117508.BRD4.cortical-interneuron 227 bp overlap
BSX 1 dataset
Motif DE_48h DE_48h-BSX_MA0876.2 6 bp overlap
CHD7 1 dataset
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 198 bp overlap
DLX1 1 dataset
Motif DE_48h DE_48h-DLX1_MA0879.3 6 bp overlap
DLX6 1 dataset
Motif DE_48h DE_48h-DLX6_MA0882.2 6 bp overlap
Dlx2 1 dataset
Motif DE_48h DE_48h-Dlx2_MA0885.3 8 bp overlap
Dlx3 1 dataset
Motif DE_48h DE_48h-Dlx3_MA0880.2 6 bp overlap
Dlx4 1 dataset
Motif DE_48h DE_48h-Dlx4_MA0881.2 6 bp overlap
Dlx5 1 dataset
Motif DE_48h DE_48h-Dlx5_MA1476.3 8 bp overlap
Dmrt1 3 datasets
Motif DE_48h DE_48h-Dmrt1_MA1603.2 9 bp overlap
Motif DE_60h DE_60h-Dmrt1_MA1603.2 9 bp overlap
Motif DE_72h DE_72h-Dmrt1_MA1603.2 9 bp overlap
ELF1 1 dataset
Motif DE_48h DE_48h-ELF1_MA0473.4 9 bp overlap
ELK1::HOXA1 3 datasets
Motif DE_48h DE_48h-ELK1HOXA1_MA1931.1 14 bp overlap
Motif DE_60h DE_60h-ELK1HOXA1_MA1931.1 14 bp overlap
Motif DE_72h DE_72h-ELK1HOXA1_MA1931.1 14 bp overlap
EN2 1 dataset
Motif DE_48h DE_48h-EN2_MA0642.3 7 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 232 bp overlap
ESR1 1 dataset
ChIP primary-endometrium-cancer_E1_DSG GSE114737.ESR1.primary-endometrium-cancer_E1_DSG 626 bp overlap
ESR2 3 datasets
Motif DE_48h DE_48h-ESR2_MA0258.2 15 bp overlap
Motif DE_60h DE_60h-ESR2_MA0258.2 15 bp overlap
Motif DE_72h DE_72h-ESR2_MA0258.2 15 bp overlap
ETV1 1 dataset
ChIP COLO-800 GSE80443.ETV1.COLO-800 195 bp overlap
FOSL2 5 datasets
ChIP LPS141 GSE111253.FOSL2.LPS141 224 bp overlap
ChIP NPC_K755R-pos GSE122631.FOSL2.NPC_K755R-pos 308 bp overlap
ChIP NPC_R1159Q GSE122631.FOSL2.NPC_R1159Q 263 bp overlap
ChIP SK-N-SH ENCFF127ZDW 285 bp overlap
ChIP SK-N-SH ENCSR000BVB.FOSL2.SK-N-SH 170 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 479 bp overlap
ChIP DE DE-FOXA2-2 461 bp overlap
FOXM1 1 dataset
ChIP MDA-MB-231_THIOS GSE40762.FOXM1.MDA-MB-231_THIOS 169 bp overlap
GABPA 1 dataset
Motif DE_48h DE_48h-GABPA_MA0062.4 10 bp overlap
GATA4 2 datasets
ChIP DE DE-GATA4-1 569 bp overlap
ChIP DE DE-GATA4-2 628 bp overlap
GATA6 4 datasets
ChIP DE DE-GATA6-1 499 bp overlap
ChIP DE DE-GATA6-2 634 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 572 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 410 bp overlap
GBX1 1 dataset
Motif DE_48h DE_48h-GBX1_MA0889.2 7 bp overlap
GBX2 1 dataset
Motif DE_48h DE_48h-GBX2_MA0890.2 6 bp overlap
HESX1 1 dataset
Motif DE_48h DE_48h-HESX1_MA0894.2 6 bp overlap
HNRNPK 2 datasets
ChIP K-562 GSE120104.HNRNPK.K-562 184 bp overlap
ChIP K-562 ENCSR014RCS.HNRNPK.K-562 185 bp overlap
HOXA7 1 dataset
Motif DE_48h DE_48h-HOXA7_MA1498.3 6 bp overlap
HOXB2::ELK1 3 datasets
Motif DE_48h DE_48h-HOXB2ELK1_MA1957.1 14 bp overlap
Motif DE_60h DE_60h-HOXB2ELK1_MA1957.1 14 bp overlap
Motif DE_72h DE_72h-HOXB2ELK1_MA1957.1 14 bp overlap
ISL2 2 datasets
Motif DE_48h DE_48h-ISL2_MA0914.2 6 bp overlap
Motif DE_60h DE_60h-ISL2_MA0914.2 6 bp overlap
Ikzf3 1 dataset
Motif DE_48h DE_48h-Ikzf3_MA1992.2 9 bp overlap
JUND 1 dataset
ChIP SK-N-SH ENCSR000EIB.JUND.SK-N-SH 179 bp overlap
KLF1 3 datasets
Motif DE_48h DE_48h-KLF1_MA0493.3 8 bp overlap
Motif DE_60h DE_60h-KLF1_MA0493.3 8 bp overlap
Motif DE_72h DE_72h-KLF1_MA0493.3 8 bp overlap
KLF10 3 datasets
Motif DE_48h DE_48h-KLF10_MA1511.2 9 bp overlap
Motif DE_60h DE_60h-KLF10_MA1511.2 9 bp overlap
Motif DE_72h DE_72h-KLF10_MA1511.2 9 bp overlap
KLF14 3 datasets
Motif DE_48h DE_48h-KLF14_MA0740.2 9 bp overlap
Motif DE_60h DE_60h-KLF14_MA0740.2 9 bp overlap
Motif DE_72h DE_72h-KLF14_MA0740.2 9 bp overlap
KLF2 3 datasets
Motif DE_48h DE_48h-KLF2_MA1515.2 8 bp overlap
Motif DE_60h DE_60h-KLF2_MA1515.2 8 bp overlap
Motif DE_72h DE_72h-KLF2_MA1515.2 8 bp overlap
KLF4 3 datasets
Motif DE_48h DE_48h-KLF4_MA0039.5 8 bp overlap
Motif DE_60h DE_60h-KLF4_MA0039.5 8 bp overlap
Motif DE_72h DE_72h-KLF4_MA0039.5 8 bp overlap
KLF5 3 datasets
Motif DE_48h DE_48h-KLF5_MA0599.1 10 bp overlap
Motif DE_60h DE_60h-KLF5_MA0599.1 10 bp overlap
Motif DE_72h DE_72h-KLF5_MA0599.1 10 bp overlap
LBX1 1 dataset
Motif DE_48h DE_48h-LBX1_MA0618.2 7 bp overlap
LBX2 1 dataset
Motif DE_48h DE_48h-LBX2_MA0699.2 6 bp overlap
LHX2 1 dataset
Motif DE_48h DE_48h-LHX2_MA0700.3 6 bp overlap
LHX9 1 dataset
Motif DE_48h DE_48h-LHX9_MA0701.3 7 bp overlap
Lhx1 1 dataset
Motif DE_48h DE_48h-Lhx1_MA1518.3 10 bp overlap
MAZ 1 dataset
ChIP HEK293 GSE76494.MAZ.HEK293 190 bp overlap
MED1 1 dataset
ChIP dopaminergic-neuron_Dopamine_neurons GSE93905.MED1.dopaminergic-neuron_Dopamine_neurons 140 bp overlap
MGA::EVX1 3 datasets
Motif DE_48h DE_48h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_60h DE_60h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_72h DE_72h-MGAEVX1_MA1960.2 11 bp overlap
MSX1 1 dataset
Motif DE_48h DE_48h-MSX1_MA0666.3 6 bp overlap
MSX2 1 dataset
Motif DE_48h DE_48h-MSX2_MA0708.3 6 bp overlap
MYB 2 datasets
Motif DE_48h DE_48h-MYB_MA0100.4 6 bp overlap
Motif DE_60h DE_60h-MYB_MA0100.4 6 bp overlap
MYBL2 3 datasets
Motif DE_48h DE_48h-MYBL2_MA0777.1 15 bp overlap
Motif DE_60h DE_60h-MYBL2_MA0777.1 15 bp overlap
Motif DE_72h DE_72h-MYBL2_MA0777.1 15 bp overlap
MYF5 1 dataset
Motif DE_48h DE_48h-MYF5_MA1641.2 8 bp overlap
MYOG 1 dataset
Motif DE_48h DE_48h-MYOG_MA0500.3 8 bp overlap
Msx3 1 dataset
Motif DE_48h DE_48h-Msx3_MA0709.2 6 bp overlap
NEUROD1 2 datasets
ChIP D283-Med GSE92582.NEUROD1.D283-Med 575 bp overlap
ChIP D283-Med_shNEUROD1-1579 GSE92582.NEUROD1.D283-Med_shNEUROD1-1579 360 bp overlap
NFYB 1 dataset
Motif DE_48h DE_48h-NFYB_MA0502.3 9 bp overlap
NHLH1 1 dataset
Motif DE_48h DE_48h-NHLH1_MA0048.3 9 bp overlap
NHLH2 1 dataset
Motif DE_48h DE_48h-NHLH2_MA1529.2 16 bp overlap
Nkx3-2 2 datasets
Motif DE_48h DE_48h-Nkx3-2_MA0122.4 10 bp overlap
Motif DE_60h DE_60h-Nkx3-2_MA0122.4 10 bp overlap
Nobox 1 dataset
Motif DE_48h DE_48h-Nobox_MA0125.2 6 bp overlap
Olig2 1 dataset
Motif DE_48h DE_48h-Olig2_MA1997.2 6 bp overlap
PATZ1 3 datasets
Motif DE_48h DE_48h-PATZ1_MA1961.2 11 bp overlap
Motif DE_60h DE_60h-PATZ1_MA1961.2 11 bp overlap
Motif DE_72h DE_72h-PATZ1_MA1961.2 11 bp overlap
PLAG1 3 datasets
Motif DE_48h DE_48h-PLAG1_MA0163.1 14 bp overlap
Motif DE_60h DE_60h-PLAG1_MA0163.1 14 bp overlap
Motif DE_72h DE_72h-PLAG1_MA0163.1 14 bp overlap
POU6F2 1 dataset
Motif DE_60h DE_60h-POU6F2_MA0793.2 9 bp overlap
PRDM9 3 datasets
Motif DE_48h DE_48h-PRDM9_MA1723.2 20 bp overlap
Motif DE_60h DE_60h-PRDM9_MA1723.2 20 bp overlap
Motif DE_72h DE_72h-PRDM9_MA1723.2 20 bp overlap
PRRX2 1 dataset
Motif DE_48h DE_48h-PRRX2_MA0075.4 7 bp overlap
Ptf1A 2 datasets
Motif DE_48h DE_48h-Ptf1A_MA1619.2 8 bp overlap
Motif DE_48h DE_48h-Ptf1A_MA1620.2 8 bp overlap
RARA 3 datasets
Motif DE_48h DE_48h-RARA_MA0729.1 18 bp overlap
Motif DE_60h DE_60h-RARA_MA0729.1 18 bp overlap
Motif DE_72h DE_72h-RARA_MA0729.1 18 bp overlap
RAX 1 dataset
Motif DE_48h DE_48h-RAX_MA0718.2 6 bp overlap
RBPJ 2 datasets
ChIP GSC8-11_12d-das GSE74557.RBPJ.GSC8-11_12d-das 171 bp overlap
ChIP GSC8-11_12d-das GSE74557.RBPJ.GSC8-11_12d-das 173 bp overlap
RREB1 3 datasets
Motif DE_48h DE_48h-RREB1_MA0073.2 19 bp overlap
Motif DE_60h DE_60h-RREB1_MA0073.2 19 bp overlap
Motif DE_72h DE_72h-RREB1_MA0073.2 19 bp overlap
Rhox11 1 dataset
Motif DE_48h DE_48h-Rhox11_MA0629.2 9 bp overlap
SATB1 1 dataset
Motif DE_60h DE_60h-SATB1_MA1963.2 7 bp overlap
SIN3A 1 dataset
ChIP SK-N-SH ENCSR000BPB.SIN3A.SK-N-SH 169 bp overlap
SIX2 1 dataset
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 71 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 634 bp overlap
SMARCA4 5 datasets
ChIP 501-mel_SHCTR GSE61965.SMARCA4.501-mel_SHCTR 735 bp overlap
ChIP 501-mel_SHMITF GSE61965.SMARCA4.501-mel_SHMITF 274 bp overlap
ChIP 501-mel_SHMITF GSE61965.SMARCA4.501-mel_SHMITF 349 bp overlap
ChIP 501-mel_SHSOX10 GSE61965.SMARCA4.501-mel_SHSOX10 261 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 219 bp overlap
SOX14 3 datasets
Motif DE_48h DE_48h-SOX14_MA1562.2 9 bp overlap
Motif DE_60h DE_60h-SOX14_MA1562.2 9 bp overlap
Motif DE_72h DE_72h-SOX14_MA1562.2 9 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 447 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 502 bp overlap
SOX21 3 datasets
Motif DE_48h DE_48h-SOX21_MA0866.1 15 bp overlap
Motif DE_60h DE_60h-SOX21_MA0866.1 15 bp overlap
Motif DE_72h DE_72h-SOX21_MA0866.1 15 bp overlap
SOX8 3 datasets
Motif DE_48h DE_48h-SOX8_MA0868.3 7 bp overlap
Motif DE_60h DE_60h-SOX8_MA0868.3 7 bp overlap
Motif DE_72h DE_72h-SOX8_MA0868.3 7 bp overlap
Sox1 3 datasets
Motif DE_48h DE_48h-Sox1_MA0870.1 15 bp overlap
Motif DE_60h DE_60h-Sox1_MA0870.1 15 bp overlap
Motif DE_72h DE_72h-Sox1_MA0870.1 15 bp overlap
Sox11 3 datasets
Motif DE_48h DE_48h-Sox11_MA0869.3 8 bp overlap
Motif DE_60h DE_60h-Sox11_MA0869.3 8 bp overlap
Motif DE_72h DE_72h-Sox11_MA0869.3 8 bp overlap
Sox17 3 datasets
Motif DE_48h DE_48h-Sox17_MA0078.3 10 bp overlap
Motif DE_60h DE_60h-Sox17_MA0078.3 10 bp overlap
Motif DE_72h DE_72h-Sox17_MA0078.3 10 bp overlap
Sox5 3 datasets
Motif DE_48h DE_48h-Sox5_MA0087.3 8 bp overlap
Motif DE_60h DE_60h-Sox5_MA0087.3 8 bp overlap
Motif DE_72h DE_72h-Sox5_MA0087.3 8 bp overlap
Sox7 3 datasets
Motif DE_48h DE_48h-Sox7_MA2095.1 10 bp overlap
Motif DE_60h DE_60h-Sox7_MA2095.1 10 bp overlap
Motif DE_72h DE_72h-Sox7_MA2095.1 10 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 194 bp overlap
TCF12 1 dataset
ChIP SK-N-SH ENCFF147AHB 391 bp overlap
TFAP4 1 dataset
Motif DE_48h DE_48h-TFAP4_MA0691.1 10 bp overlap
Tbx6 1 dataset
Motif DE_48h DE_48h-Tbx6_MA1567.3 9 bp overlap
Tcf12 1 dataset
Motif DE_48h DE_48h-Tcf12_MA0521.3 6 bp overlap
Twist2 1 dataset
Motif DE_48h DE_48h-Twist2_MA0633.3 6 bp overlap
Wt1 3 datasets
Motif DE_48h DE_48h-Wt1_MA1627.2 10 bp overlap
Motif DE_60h DE_60h-Wt1_MA1627.2 10 bp overlap
Motif DE_72h DE_72h-Wt1_MA1627.2 10 bp overlap
ZBTB18 1 dataset
Motif DE_48h DE_48h-ZBTB18_MA0698.2 11 bp overlap
ZFP14 2 datasets
Motif DE_48h DE_48h-ZFP14_MA1972.1 15 bp overlap
Motif DE_60h DE_60h-ZFP14_MA1972.1 15 bp overlap
ZNF547 1 dataset
Motif DE_48h DE_48h-ZNF547_MA2334.1 13 bp overlap
ZNF652 3 datasets
Motif DE_48h DE_48h-ZNF652_MA1657.2 9 bp overlap
Motif DE_60h DE_60h-ZNF652_MA1657.2 9 bp overlap
Motif DE_72h DE_72h-ZNF652_MA1657.2 9 bp overlap
ZNF677 3 datasets
Motif DE_48h DE_48h-ZNF677_MA2101.1 12 bp overlap
Motif DE_60h DE_60h-ZNF677_MA2101.1 12 bp overlap
Motif DE_72h DE_72h-ZNF677_MA2101.1 12 bp overlap
ZNF680 2 datasets
Motif DE_48h DE_48h-ZNF680_MA1729.2 11 bp overlap
Motif DE_60h DE_60h-ZNF680_MA1729.2 11 bp overlap
ZNF707 2 datasets
Motif DE_48h DE_48h-ZNF707_MA1715.1 15 bp overlap
Motif DE_60h DE_60h-ZNF707_MA1715.1 15 bp overlap
ZNF740 3 datasets
Motif DE_48h DE_48h-ZNF740_MA0753.3 10 bp overlap
Motif DE_60h DE_60h-ZNF740_MA0753.3 10 bp overlap
Motif DE_72h DE_72h-ZNF740_MA0753.3 10 bp overlap
ZNF766 1 dataset
ChIP HEK293T GSE78099.ZNF766.HEK293T 104 bp overlap
ZNF770 1 dataset
Motif DE_48h DE_48h-ZNF770_MA2099.1 8 bp overlap