chr20 : 56,003,967 56,004,403
436 bp 77 TFs 1 linked gene
This 436 bp open chromatin element is linked to CBLN4 and is bound by 77 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
CBLN4 1.1 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr20:55,998,967 – 56,009,403
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
77 transcription factors
Source
Cell type
AATF 1 dataset
ChIP NALM-6 GSE93626.AATF.NALM-6 120 bp overlap
AGO1 2 datasets
ChIP Hep-G2 ENCSR555ZMV.AGO1.Hep-G2 183 bp overlap
ChIP HepG2 ENCFF277EOU 436 bp overlap
AR 1 dataset
ChIP LNCaP_DHTTHZ1 GSE125245.AR.LNCaP_DHTTHZ1 116 bp overlap
ARNT 1 dataset
ChIP 501-mel GSE95280.ARNT.501-mel 310 bp overlap
BCOR 3 datasets
ChIP WA01 GSE104690.BCOR.WA01 436 bp overlap
ChIP WA01_KDM2B_KO GSE104690.BCOR.WA01_KDM2B_KO 168 bp overlap
ChIP WA01_RNF2-R_Doxycyclin GSE104690.BCOR.WA01_RNF2-R_Doxycyclin 303 bp overlap
BRD4 5 datasets
ChIP HEK293_sgNT GSE129407.BRD4.HEK293_sgNT 75 bp overlap
ChIP MCF-10A ERP003925.BRD4.MCF-10A 293 bp overlap
ChIP MDA-MB-157 ERP003925.BRD4.MDA-MB-157 254 bp overlap
ChIP T-47D ERP003925.BRD4.T-47D 350 bp overlap
ChIP retina_AB1-RB GSE86981.BRD4.retina_AB1-RB 317 bp overlap
CBFB 1 dataset
ChIP ME-1 GSE46044.CBFB.ME-1 416 bp overlap
CHD1 1 dataset
ChIP WA01 ENCSR000AQK.CHD1.WA01 191 bp overlap
CTCF 13 datasets
ChIP GSC23 GSE139416.CTCF.GSC23 376 bp overlap
ChIP SK-N-SH ENCSR000DXQ.CTCF.SK-N-SH 99 bp overlap
ChIP T-47D_NaCl-isotonic-triptolide GSE111923.CTCF.T-47D_NaCl-isotonic-triptolide 288 bp overlap
ChIP T-47D_Y537S GSE148277.CTCF.T-47D_Y537S 404 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 233 bp overlap
ChIP anterior-temporal-cortex_fetal GSE116825.CTCF.anterior-temporal-cortex_fetal 299 bp overlap
ChIP cortical-interneuron_KCl-neg_JQ1-neg GSE117508.CTCF.cortical-interneuron_KCl-neg_JQ1-neg 192 bp overlap
ChIP cortical-interneuron_KCl-pos_JQ1-pos GSE117508.CTCF.cortical-interneuron_KCl-pos_JQ1-pos 207 bp overlap
ChIP keratinocyte ENCSR000DWX.CTCF.keratinocyte 134 bp overlap
ChIP neural ENCSR822CEA.CTCF.neural 339 bp overlap
ChIP right lobe of liver ENCFF011NDG 346 bp overlap
ChIP thyroid gland ENCFF300RYK 335 bp overlap
ChIP thyroid-gland ENCSR033KMZ.CTCF.thyroid-gland 250 bp overlap
E2F1 1 dataset
ChIP LNCaP_shRB GSE94958.E2F1.LNCaP_shRB 250 bp overlap
E2F6 1 dataset
ChIP WA01 ENCSR000BSI.E2F6.WA01 243 bp overlap
EHMT2 1 dataset
ChIP Rh41 GSE118666.EHMT2.Rh41 283 bp overlap
ELF1 1 dataset
ChIP ME-1 GSE46044.ELF1.ME-1 214 bp overlap
EPAS1 1 dataset
ChIP 501-mel GSE95280.EPAS1.501-mel 357 bp overlap
ERG 2 datasets
ChIP CD34_NR29 GSE23730.ERG.CD34_NR29 297 bp overlap
ChIP ME-1 GSE46044.ERG.ME-1 210 bp overlap
ESR1 11 datasets
ChIP Ishikawa GSE99905.ESR1.Ishikawa 416 bp overlap
ChIP MCF-7 GSE48930.ESR1.MCF-7 242 bp overlap
ChIP MCF-7 GSE148277.ESR1.MCF-7 233 bp overlap
ChIP MCF-7_PaPE-1 GSE93510.ESR1.MCF-7_PaPE-1 331 bp overlap
ChIP MCF-7_PaPE-1_OA GSE93510.ESR1.MCF-7_PaPE-1_OA 379 bp overlap
ChIP MCF-7_RUNX2_DOX GSE86538.ESR1.MCF-7_RUNX2_DOX 203 bp overlap
ChIP MCF-7_Y537S_DMSO GSE148277.ESR1.MCF-7_Y537S_DMSO 394 bp overlap
ChIP MCF-7_aldosterone GSE99626.ESR1.MCF-7_aldosterone 332 bp overlap
ChIP MCF-7_estradiol_45min_ChIP-and-reChIP GSE99626.ESR1.MCF-7_estradiol_45min_ChIP-and-reChIP 269 bp overlap
ChIP T-47D GSE148277.ESR1.T-47D 346 bp overlap
ChIP T-47D_Y537S_E2 GSE148277.ESR1.T-47D_Y537S_E2 235 bp overlap
EZH2 11 datasets
ChIP A-1847_CARM1-KO GSE95643.EZH2.A-1847_CARM1-KO 106 bp overlap
ChIP H1 ENCFF232NZA 436 bp overlap
ChIP LNCaP-abl GSE39459.EZH2.LNCaP-abl 416 bp overlap
ChIP THP-1 GSE135024.EZH2.THP-1 61 bp overlap
ChIP astrocyte ENCSR000ARR.EZH2.astrocyte 312 bp overlap
ChIP epithelial_mammary ENCSR000ARE.EZH2.epithelial_mammary 176 bp overlap
ChIP hESC GSE113817.EZH2.hESC 401 bp overlap
ChIP mammary epithelial cell ENCFF224GAI 250 bp overlap
ChIP neural progenitor cell ENCFF018MKA 282 bp overlap
ChIP neural progenitor cell ENCFF472NFV 436 bp overlap
ChIP prostate-cancer_plko GSE107780.EZH2.prostate-cancer_plko 358 bp overlap
FOXA1 1 dataset
ChIP breast-cancer_heregulin GSE101407.FOXA1.breast-cancer_heregulin 67 bp overlap
FOXO1 1 dataset
ChIP CD34 GSE80773.FOXO1.CD34 180 bp overlap
GABPA 1 dataset
ChIP WA01 ENCSR000BIW.GABPA.WA01 146 bp overlap
HDAC2 1 dataset
ChIP WA01 ENCSR000AVB.HDAC2.WA01 192 bp overlap
HIF3A 1 dataset
ChIP Hep-3B2-1-7 GSE129491.HIF3A.Hep-3B2-1-7 359 bp overlap
HNRNPLL 2 datasets
ChIP Hep-G2 ENCSR804HMZ.HNRNPLL.Hep-G2 83 bp overlap
ChIP Hep-G2 GSE120104.HNRNPLL.Hep-G2 68 bp overlap
HOXA3 1 dataset
ChIP Hep-G2 ENCSR999JKC.HOXA3.Hep-G2 150 bp overlap
JARID2 2 datasets
ChIP AMIPS6 GSE48516.JARID2.AMIPS6 364 bp overlap
ChIP EDOMIPS2 GSE48516.JARID2.EDOMIPS2 436 bp overlap
KDM4A 2 datasets
ChIP WA01 ENCSR000AVC.KDM4A.WA01 203 bp overlap
ChIP WA01 ENCSR000AVC.KDM4A.WA01 86 bp overlap
KDM5B 3 datasets
ChIP Hep-G2 ENCSR227MRE.KDM5B.Hep-G2 162 bp overlap
ChIP MCF-7 GSE46055.KDM5B.MCF-7 392 bp overlap
ChIP T-47D GSE46055.KDM5B.T-47D 134 bp overlap
KDM6B 1 dataset
ChIP CUTLL1 GSE56694.KDM6B.CUTLL1 345 bp overlap
KLF4 1 dataset
ChIP MCF-7 ENCFF948KTQ 114 bp overlap
MECOM 1 dataset
ChIP SKH1_CEBPA-ER GSE102697.MECOM.SKH1_CEBPA-ER 199 bp overlap
MNX1 1 dataset
ChIP Hep-G2 ENCSR108MKR.MNX1.Hep-G2 327 bp overlap
MTF1 1 dataset
Motif DE_12h DE_12h-MTF1_MA0863.1 14 bp overlap
MXI1 1 dataset
ChIP SK-N-SH ENCSR000EIA.MXI1.SK-N-SH 153 bp overlap
MYC 3 datasets
ChIP U2OS_Doxy GSE44672.MYC.U2OS_Doxy 118 bp overlap
ChIP U2OS_Doxy GSE77356.MYC.U2OS_Doxy 127 bp overlap
ChIP U2OS_SHRCTR GSE77356.MYC.U2OS_SHRCTR 135 bp overlap
NANOG 1 dataset
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 328 bp overlap
NFAT5 1 dataset
ChIP Hep-G2 ENCSR565BVI.NFAT5.Hep-G2 150 bp overlap
NFKB1 1 dataset
ChIP HEK293T_2KR GSE129618.NFKB1.HEK293T_2KR 201 bp overlap
NR2F2 1 dataset
ChIP MCF-7_E2 GSE132432.NR2F2.MCF-7_E2 436 bp overlap
Nrf1 1 dataset
Motif DE_12h DE_12h-Nrf1_MA0506.3 12 bp overlap
POU2F1 2 datasets
ChIP T-47D GSE148277.POU2F1.T-47D 408 bp overlap
ChIP T-47D_D538G GSE148277.POU2F1.T-47D_D538G 300 bp overlap
POU5F1 1 dataset
ChIP WA01_3IL ERP004238.POU5F1.WA01_3IL 387 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 333 bp overlap
RAD21 4 datasets
ChIP H1 ENCFF698EWO 241 bp overlap
ChIP liver ENCFF522JHE 337 bp overlap
ChIP neural ENCSR198ZYJ.RAD21.neural 367 bp overlap
ChIP neural cell ENCFF564MOT 393 bp overlap
REST 35 datasets
ChIP GM12878 ENCFF235NGC 231 bp overlap
ChIP GM12878 ENCSR000BGF.REST.GM12878 147 bp overlap
ChIP GM12878 ENCSR000BQS.REST.GM12878 119 bp overlap
ChIP H1 ENCFF203SWY 376 bp overlap
ChIP H1 ENCFF429RUE 234 bp overlap
ChIP HEK293 ENCFF073DOT 407 bp overlap
ChIP HEK293 ENCSR896UBV.REST.HEK293 223 bp overlap
ChIP HL-60 ENCFF589LOF 267 bp overlap
ChIP HL-60 ENCSR000BTF.REST.HL-60 304 bp overlap
ChIP Hep-G2 ENCSR000BOT.REST.Hep-G2 285 bp overlap
ChIP Hep-G2 ENCSR000BJL.REST.Hep-G2 311 bp overlap
ChIP HepG2 ENCFF122AWR 291 bp overlap
ChIP HepG2 ENCFF800JSL 180 bp overlap
ChIP Ishikawa ENCFF456OHV 371 bp overlap
ChIP Ishikawa ENCSR000BUU.REST.Ishikawa 193 bp overlap
ChIP K-562 ENCSR000ATM.REST.K-562 436 bp overlap
ChIP K-562 ENCSR000BMW.REST.K-562 130 bp overlap
ChIP LNCaP GSE119385.REST.LNCaP 315 bp overlap
ChIP MCF-7 ENCFF893RRD 345 bp overlap
ChIP MCF-7 ENCSR000BSP.REST.MCF-7 241 bp overlap
ChIP PANC-1 ENCSR000BJO.REST.PANC-1 314 bp overlap
ChIP PANC-1 ENCSR000BPK.REST.PANC-1 181 bp overlap
ChIP PFSK-1 ENCFF845VHA 321 bp overlap
ChIP PFSK1 ENCSR000BOX.REST.PFSK1 264 bp overlap
ChIP Panc1 ENCFF629OJO 285 bp overlap
ChIP SK-N-SH ENCFF861MKH 175 bp overlap
ChIP SK-N-SH ENCSR000BJJ.REST.SK-N-SH 263 bp overlap
ChIP WA01 ENCSR000BHM.REST.WA01 374 bp overlap
ChIP WA01 ENCSR663WAR.REST.WA01 412 bp overlap
ChIP colorectal-cancer_shCDH1 GSE112555.REST.colorectal-cancer_shCDH1 436 bp overlap
ChIP colorectal-cancer_shCTRL_dissociated GSE112555.REST.colorectal-cancer_shCTRL_dissociated 328 bp overlap
ChIP colorectal-cancer_shCTRL_intact GSE112555.REST.colorectal-cancer_shCTRL_intact 436 bp overlap
ChIP liver ENCFF240FWT 299 bp overlap
ChIP liver ENCSR867WPH.REST.liver 397 bp overlap
ChIP liver ENCSR893QWP.REST.liver 313 bp overlap
RNF2 3 datasets
ChIP H1 ENCFF239FFS 74 bp overlap
ChIP WA01 ENCSR784VUY.RNF2.WA01 203 bp overlap
ChIP WA01 GSE104690.RNF2.WA01 417 bp overlap
RORC 2 datasets
ChIP HCC70 GSE126380.RORC.HCC70 436 bp overlap
ChIP HCC70_XY018 GSE126380.RORC.HCC70_XY018 391 bp overlap
RUNX1 1 dataset
ChIP Jurkat GSE85524.RUNX1.Jurkat 290 bp overlap
RXR 1 dataset
ChIP TSU-1621MT GSE60477.RXR.TSU-1621MT 198 bp overlap
SIN3A 1 dataset
ChIP WA01 ENCSR000EBO.SIN3A.WA01 159 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 256 bp overlap
SMARCA4 3 datasets
ChIP 22Rv1 GSE115615.SMARCA4.22Rv1 436 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 230 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 213 bp overlap
SMARCC1 2 datasets
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4 174 bp overlap
ChIP ESC S25-ESC-d0-BAF155-exp1 205 bp overlap
SMC1A 1 dataset
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.SMC1A.primary-epidermal-keratinocyte_diff_d0 281 bp overlap
SOX2 1 dataset
ChIP RENVM GSE49404.SOX2.RENVM 182 bp overlap
SREBP2 2 datasets
ChIP HCC70 GSE126380.SREBP2.HCC70 403 bp overlap
ChIP HCC70_XY018 GSE126380.SREBP2.HCC70_XY018 391 bp overlap
SUZ12 5 datasets
ChIP Aska-SS GSE108025.SUZ12.Aska-SS 436 bp overlap
ChIP H1 ENCFF881NFR 436 bp overlap
ChIP LNCaP GSE39459.SUZ12.LNCaP 245 bp overlap
ChIP Lu-130 GSE99312.SUZ12.Lu-130 152 bp overlap
ChIP WA01 ENCSR000ATS.SUZ12.WA01 153 bp overlap
TAL1 1 dataset
ChIP ME-1 GSE46044.TAL1.ME-1 397 bp overlap
TARDBP 2 datasets
ChIP Hep-G2 ENCSR145CXH.TARDBP.Hep-G2 162 bp overlap
ChIP Hep-G2 GSE120104.TARDBP.Hep-G2 157 bp overlap
TBP 2 datasets
ChIP H1 ENCFF859IIO 377 bp overlap
ChIP WA01 ENCSR000ECB.TBP.WA01 304 bp overlap
TFAP2A 1 dataset
Motif DE_12h DE_12h-TFAP2A_MA0872.1 13 bp overlap
TRIM25 1 dataset
ChIP BT-549 GSE79588.TRIM25.BT-549 338 bp overlap
TRIM28 1 dataset
ChIP HCT-116 GSE72622.TRIM28.HCT-116 72 bp overlap
TSHZ2 2 datasets
ChIP SK-N-SH ENCFF182EBB 381 bp overlap
ChIP SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour ENCSR282NLQ.TSHZ2.SK-N-SH_all-trans-retinoic-acid_6-uM_48-hour 242 bp overlap
Tfcp2l1 1 dataset
Motif DE_12h DE_12h-Tfcp2l1_MA0145.2 14 bp overlap
WDR5 1 dataset
ChIP SMMC-7721 GSE126982.WDR5.SMMC-7721 436 bp overlap
ZBTB48 1 dataset
ChIP U2OS_ZBTB48-KO GSE96776.ZBTB48.U2OS_ZBTB48-KO 385 bp overlap
ZBTB7A 1 dataset
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 272 bp overlap
ZNF454 1 dataset
Motif DE_12h DE_12h-ZNF454_MA1712.2 17 bp overlap
ZNF549 1 dataset
Motif DE_12h DE_12h-ZNF549_MA1728.2 8 bp overlap
ZNF669 1 dataset
Motif DE_12h DE_12h-ZNF669_MA1985.1 15 bp overlap
ZNF90 1 dataset
ChIP HEK293T GSE78099.ZNF90.HEK293T 108 bp overlap
ZNF93 1 dataset
Motif DE_12h DE_12h-ZNF93_MA1721.2 14 bp overlap