chr16 : 78,243,715 78,244,828
1,113 bp 121 TFs 2 linked genes
This 1.1 kb open chromatin element is linked to WWOX and MAF and is bound by 121 transcription factors.
Linked Genes
2 genes
Link type
Gene Expression Dist. to TSS Distance Link type
WWOX 145.2 kb Distal Multiome
MAF 1356.1 kb Distal Multiome+HiCAR
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr16:78,238,715 – 78,249,828
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
121 transcription factors
Source
Cell type
ASCL1 6 datasets
Motif DE_48h DE_48h-ASCL1_MA1100.3 8 bp overlap
Motif DE_48h DE_48h-ASCL1_MA1631.2 9 bp overlap
Motif DE_60h DE_60h-ASCL1_MA1100.3 8 bp overlap
Motif DE_60h DE_60h-ASCL1_MA1631.2 9 bp overlap
Motif DE_72h DE_72h-ASCL1_MA1100.3 8 bp overlap
Motif DE_72h DE_72h-ASCL1_MA1631.2 9 bp overlap
CDX2 2 datasets
ChIP Caco-2_DIFF GSE23436.CDX2.Caco-2_DIFF 262 bp overlap
ChIP Caco-2_PROLIF GSE23436.CDX2.Caco-2_PROLIF 408 bp overlap
CTCF 1 dataset
Motif DE_72h DE_72h-CTCF_MA1929.2 31 bp overlap
CTNNB1 1 dataset
ChIP LS180_125 GSE31939.CTNNB1.LS180_125 130 bp overlap
CUX2 2 datasets
Motif DE_60h DE_60h-CUX2_MA0755.2 9 bp overlap
Motif DE_72h DE_72h-CUX2_MA0755.2 9 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF093OYK 326 bp overlap
Crx 2 datasets
Motif DE_60h DE_60h-Crx_MA0467.3 6 bp overlap
Motif DE_72h DE_72h-Crx_MA0467.3 6 bp overlap
Dmbx1 1 dataset
Motif DE_72h DE_72h-Dmbx1_MA0883.2 10 bp overlap
ELF1 1 dataset
Motif DE_72h DE_72h-ELF1_MA0473.4 9 bp overlap
ELF3 1 dataset
Motif DE_72h DE_72h-ELF3_MA0640.3 9 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 757 bp overlap
ETV1 1 dataset
Motif DE_48h DE_48h-ETV1_MA0761.3 9 bp overlap
ETV5::FOXO1 2 datasets
Motif DE_72h DE_72h-ETV5FOXO1_MA1947.2 10 bp overlap
Motif DE_72h DE_72h-ETV5FOXO1_MA1947.2 10 bp overlap
EZH2 1 dataset
ChIP hepatocyte ENCFF118DKH 93 bp overlap
FIGLA 3 datasets
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
Motif DE_72h DE_72h-FIGLA_MA0820.2 6 bp overlap
FOSL2 1 dataset
ChIP NPC_K755R-pos GSE122631.FOSL2.NPC_K755R-pos 166 bp overlap
FOXA1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 383 bp overlap
FOXA2 4 datasets
ChIP Caco-2 GSE66218.FOXA2.Caco-2 250 bp overlap
ChIP DE DE-FOXA2-1 1113 bp overlap
ChIP DE DE-FOXA2-2 1113 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 293 bp overlap
FOXH1 1 dataset
Motif DE_48h DE_48h-FOXH1_MA0479.2 8 bp overlap
FOXL2 1 dataset
ChIP KGN_1512 GSE138496.FOXL2.KGN_1512 233 bp overlap
FOXP2 1 dataset
Motif DE_72h DE_72h-FOXP2_MA0593.2 9 bp overlap
GATA1 2 datasets
ChIP K-562_dCas9-LSD1 GSE132212.GATA1.K-562_dCas9-LSD1 194 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 194 bp overlap
GATA4 8 datasets
ChIP DE DE-GATA4-1 1110 bp overlap
ChIP DE DE-GATA4-2 1113 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 892 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 867 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 1019 bp overlap
GATA5 3 datasets
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
GATA6 18 datasets
ChIP AGS GSE51705.GATA6.AGS 136 bp overlap
ChIP Caco-2_PROLIF GSE23436.GATA6.Caco-2_PROLIF 254 bp overlap
ChIP DE DE-GATA6-1 1069 bp overlap
ChIP DE DE-GATA6-2 1113 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 304 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 339 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 825 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 814 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 1113 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 893 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 902 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 245 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 961 bp overlap
ChIP foregut GSE117136.GATA6.foregut 834 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 625 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 797 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 864 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 982 bp overlap
GSC 2 datasets
Motif DE_60h DE_60h-GSC_MA0648.2 6 bp overlap
Motif DE_72h DE_72h-GSC_MA0648.2 6 bp overlap
GSC2 2 datasets
Motif DE_60h DE_60h-GSC2_MA0891.2 6 bp overlap
Motif DE_72h DE_72h-GSC2_MA0891.2 6 bp overlap
Gfi1B 1 dataset
Motif DE_72h DE_72h-Gfi1B_MA0483.2 10 bp overlap
HIC1 1 dataset
ChIP HEK293 ENCSR803GYT.HIC1.HEK293 235 bp overlap
HIF1A 1 dataset
ChIP U2OS_trough_DMOG GSE85096.HIF1A.U2OS_trough_DMOG 226 bp overlap
HNF4A 4 datasets
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 194 bp overlap
ChIP Caco-2_PROLIF GSE23436.HNF4A.Caco-2_PROLIF 209 bp overlap
Motif DE_60h DE_60h-HNF4A_MA0114.5 9 bp overlap
Motif DE_72h DE_72h-HNF4A_MA0114.5 9 bp overlap
HNF4G 2 datasets
Motif DE_60h DE_60h-HNF4G_MA0484.3 9 bp overlap
Motif DE_72h DE_72h-HNF4G_MA0484.3 9 bp overlap
Hmx1 2 datasets
Motif DE_60h DE_60h-Hmx1_MA0896.2 9 bp overlap
Motif DE_72h DE_72h-Hmx1_MA0896.2 9 bp overlap
Hmx2 2 datasets
Motif DE_60h DE_60h-Hmx2_MA0897.2 15 bp overlap
Motif DE_72h DE_72h-Hmx2_MA0897.2 15 bp overlap
IKZF1 1 dataset
Motif DE_48h DE_48h-IKZF1_MA1508.2 8 bp overlap
ISL2 2 datasets
Motif DE_60h DE_60h-ISL2_MA0914.2 6 bp overlap
Motif DE_72h DE_72h-ISL2_MA0914.2 6 bp overlap
JUN 6 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 315 bp overlap
ChIP DE_D1 S13-DE-d1-JUN-exp1 264 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 279 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 901 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 848 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 339 bp overlap
KDM1A 2 datasets
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 949 bp overlap
ChIP pancreatic-progenitor_PP2-KDM1A-inh GSE104840.KDM1A.pancreatic-progenitor_PP2-KDM1A-inh 275 bp overlap
MAF 2 datasets
Motif DE_60h DE_60h-MAF_MA1520.2 13 bp overlap
Motif DE_72h DE_72h-MAF_MA1520.2 13 bp overlap
MAFA 2 datasets
Motif DE_60h DE_60h-MAFA_MA1521.2 13 bp overlap
Motif DE_72h DE_72h-MAFA_MA1521.2 13 bp overlap
MAX 2 datasets
ChIP Ishikawa ENCSR000BTY.MAX.Ishikawa 120 bp overlap
ChIP MCF-7 ENCSR000BUL.MAX.MCF-7 129 bp overlap
MEIS1 3 datasets
Motif DE_60h DE_60h-MEIS1_MA1639.2 9 bp overlap
Motif DE_72h DE_72h-MEIS1_MA0498.3 5 bp overlap
Motif DE_72h DE_72h-MEIS1_MA1639.2 9 bp overlap
Mafg 2 datasets
Motif DE_60h DE_60h-Mafg_MA0659.4 12 bp overlap
Motif DE_72h DE_72h-Mafg_MA0659.4 12 bp overlap
Msgn1 2 datasets
Motif DE_60h DE_60h-Msgn1_MA1524.3 10 bp overlap
Motif DE_72h DE_72h-Msgn1_MA1524.3 10 bp overlap
NFE2 1 dataset
Motif DE_72h DE_72h-NFE2_MA0841.2 10 bp overlap
NKX2-3 2 datasets
Motif DE_60h DE_60h-NKX2-3_MA0672.2 8 bp overlap
Motif DE_72h DE_72h-NKX2-3_MA0672.2 8 bp overlap
NKX2-8 2 datasets
Motif DE_60h DE_60h-NKX2-8_MA0673.2 8 bp overlap
Motif DE_72h DE_72h-NKX2-8_MA0673.2 8 bp overlap
Nkx3-2 2 datasets
Motif DE_60h DE_60h-Nkx3-2_MA0122.4 10 bp overlap
Motif DE_72h DE_72h-Nkx3-2_MA0122.4 10 bp overlap
ONECUT1 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.ONECUT1.pancreatic-progenitor_PP1 340 bp overlap
OSR1 2 datasets
Motif DE_60h DE_60h-OSR1_MA1542.2 8 bp overlap
Motif DE_72h DE_72h-OSR1_MA1542.2 8 bp overlap
OTX1 2 datasets
Motif DE_60h DE_60h-OTX1_MA0711.2 6 bp overlap
Motif DE_72h DE_72h-OTX1_MA0711.2 6 bp overlap
PBX1 2 datasets
Motif DE_60h DE_60h-PBX1_MA0070.2 9 bp overlap
Motif DE_72h DE_72h-PBX1_MA0070.2 9 bp overlap
PDX1 4 datasets
ChIP hESC GSE58685.PDX1.hESC 269 bp overlap
ChIP hiPSC GSE125768.PDX1.hiPSC 451 bp overlap
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 422 bp overlap
ChIP pancreatic-progenitor GSE117136.PDX1.pancreatic-progenitor 672 bp overlap
PITX1 2 datasets
Motif DE_60h DE_60h-PITX1_MA0682.3 6 bp overlap
Motif DE_72h DE_72h-PITX1_MA0682.3 6 bp overlap
PITX2 2 datasets
Motif DE_60h DE_60h-PITX2_MA1547.2 8 bp overlap
Motif DE_72h DE_72h-PITX2_MA1547.2 8 bp overlap
PITX3 2 datasets
Motif DE_60h DE_60h-PITX3_MA0714.2 6 bp overlap
Motif DE_72h DE_72h-PITX3_MA0714.2 6 bp overlap
PSIP1 1 dataset
ChIP T-REx-293 GSE117155.PSIP1.T-REx-293 298 bp overlap
RARA 1 dataset
ChIP hiPSC_D5 GSE132532.RARA.hiPSC_D5 233 bp overlap
RELB 1 dataset
Motif DE_72h DE_72h-RELB_MA1117.2 7 bp overlap
RFX7 2 datasets
Motif DE_60h DE_60h-RFX7_MA1554.2 8 bp overlap
Motif DE_72h DE_72h-RFX7_MA1554.2 8 bp overlap
RHOXF1 2 datasets
Motif DE_60h DE_60h-RHOXF1_MA0719.2 6 bp overlap
Motif DE_72h DE_72h-RHOXF1_MA0719.2 6 bp overlap
RUNX3 1 dataset
Motif DE_72h DE_72h-RUNX3_MA0684.3 8 bp overlap
RXR 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.RXR.pancreatic-progenitor_PP1 196 bp overlap
RXRA::VDR 2 datasets
Motif DE_48h DE_48h-RXRAVDR_MA0074.1 15 bp overlap
Motif DE_72h DE_72h-RXRAVDR_MA0074.1 15 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 1113 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 930 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 291 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 666 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 1026 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 1113 bp overlap
SMARCA4 3 datasets
ChIP Aska-SS GSE108025.SMARCA4.Aska-SS 464 bp overlap
ChIP SYO-1 GSE108025.SMARCA4.SYO-1 756 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 241 bp overlap
SMARCC1 4 datasets
ChIP Aska-SS GSE108025.SMARCC1.Aska-SS 848 bp overlap
ChIP DE_D1 S15-DE-d1-BAF155-exp1 243 bp overlap
ChIP DE_D1 S15-DE-d1-BAF155-exp1 226 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 211 bp overlap
SNAI1 3 datasets
Motif DE_48h DE_48h-SNAI1_MA1558.2 7 bp overlap
Motif DE_60h DE_60h-SNAI1_MA1558.2 7 bp overlap
Motif DE_72h DE_72h-SNAI1_MA1558.2 7 bp overlap
SNAI2 4 datasets
Motif DE_48h DE_48h-SNAI2_MA0745.3 8 bp overlap
Motif DE_60h DE_60h-SNAI2_MA0745.3 8 bp overlap
Motif DE_72h DE_72h-SNAI2_MA0745.3 8 bp overlap
ChIP keratinocyte_LacZ GSE55421.SNAI2.keratinocyte_LacZ 187 bp overlap
SNAI3 3 datasets
Motif DE_48h DE_48h-SNAI3_MA1559.2 9 bp overlap
Motif DE_60h DE_60h-SNAI3_MA1559.2 9 bp overlap
Motif DE_72h DE_72h-SNAI3_MA1559.2 9 bp overlap
SOX17 2 datasets
ChIP DE_D2 DED2-SOX17_Batch_II 600 bp overlap
ChIP DE_D2 DED2-SOX17_Batch_II 204 bp overlap
SOX17_M 2 datasets
ChIP DE_D2 DED2-SOX17-M_Batch_II 535 bp overlap
ChIP DE_D2 DED2-SOX17-M_Batch_II 231 bp overlap
SP5 2 datasets
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
Motif DE_72h DE_72h-SP5_MA1965.2 6 bp overlap
SRF 2 datasets
Motif DE_60h DE_60h-SRF_MA0083.3 16 bp overlap
Motif DE_72h DE_72h-SRF_MA0083.3 16 bp overlap
SS18 2 datasets
ChIP SYO-1_shCt GSE139053.SS18.SYO-1_shCt 423 bp overlap
ChIP SYO-1_shCt GSE139053.SS18.SYO-1_shCt 162 bp overlap
Six3 2 datasets
Motif DE_48h DE_48h-Six3_MA0631.2 11 bp overlap
Motif DE_72h DE_72h-Six3_MA0631.2 11 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 249 bp overlap
TAL1 1 dataset
ChIP K-562_enCRISPRi-LK GSE132212.TAL1.K-562_enCRISPRi-LK 214 bp overlap
TCF12 3 datasets
Motif DE_48h DE_48h-TCF12_MA1648.2 7 bp overlap
Motif DE_60h DE_60h-TCF12_MA1648.2 7 bp overlap
Motif DE_72h DE_72h-TCF12_MA1648.2 7 bp overlap
TCF3 3 datasets
Motif DE_48h DE_48h-TCF3_MA0522.4 7 bp overlap
Motif DE_60h DE_60h-TCF3_MA0522.4 7 bp overlap
Motif DE_72h DE_72h-TCF3_MA0522.4 7 bp overlap
TCF4 3 datasets
Motif DE_48h DE_48h-TCF4_MA0830.3 8 bp overlap
Motif DE_60h DE_60h-TCF4_MA0830.3 8 bp overlap
Motif DE_72h DE_72h-TCF4_MA0830.3 8 bp overlap
TEAD1 3 datasets
ChIP HEK293_siE1A GSE130135.TEAD1.HEK293_siE1A 165 bp overlap
ChIP HUCCT1 GSE68296.TEAD1.HUCCT1 161 bp overlap
ChIP pancreas_12D ERP008682.TEAD1.pancreas_12D 181 bp overlap
TEAD4 2 datasets
Motif DE_48h DE_48h-TEAD4_MA0809.3 8 bp overlap
ChIP hMSC-TERT4 GSE140782.TEAD4.hMSC-TERT4 205 bp overlap
TRIM28 1 dataset
ChIP hESC GSE115387.TRIM28.hESC 174 bp overlap
YY1AP1 3 datasets
ChIP PC-9_1DF GSE131687.YY1AP1.PC-9_1DF 532 bp overlap
ChIP PC-9_1DF_DMSO GSE131687.YY1AP1.PC-9_1DF_DMSO 307 bp overlap
ChIP PC-9_2DF GSE131687.YY1AP1.PC-9_2DF 236 bp overlap
ZBTB12 1 dataset
Motif DE_72h DE_72h-ZBTB12_MA1649.2 7 bp overlap
ZEB1 3 datasets
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
Motif DE_72h DE_72h-ZEB1_MA0103.4 6 bp overlap
ZFP14 1 dataset
Motif DE_72h DE_72h-ZFP14_MA1972.1 15 bp overlap
ZFX 1 dataset
ChIP PrEC GSE102616.ZFX.PrEC 252 bp overlap
ZIC1 1 dataset
Motif DE_72h DE_72h-ZIC1_MA0696.1 14 bp overlap
ZIC2 1 dataset
ChIP HEK293 ENCFF033NQQ 238 bp overlap
ZIC4 1 dataset
Motif DE_72h DE_72h-ZIC4_MA0751.2 14 bp overlap
ZNF175 1 dataset
Motif DE_48h DE_48h-ZNF175_MA2332.1 9 bp overlap
ZNF214 2 datasets
Motif DE_60h DE_60h-ZNF214_MA1975.2 13 bp overlap
Motif DE_72h DE_72h-ZNF214_MA1975.2 13 bp overlap
ZNF416 2 datasets
Motif DE_60h DE_60h-ZNF416_MA1979.2 10 bp overlap
Motif DE_72h DE_72h-ZNF416_MA1979.2 10 bp overlap
ZNF460 3 datasets
Motif DE_60h DE_60h-ZNF460_MA1596.1 16 bp overlap
Motif DE_72h DE_72h-ZNF460_MA1596.1 16 bp overlap
Motif DE_72h DE_72h-ZNF460_MA1596.1 16 bp overlap
ZNF664 2 datasets
ChIP HEK293 ENCFF343XSW 385 bp overlap
ChIP HEK293 ENCSR714LZQ.ZNF664.HEK293 301 bp overlap
ZNF682 2 datasets
Motif DE_60h DE_60h-ZNF682_MA1599.2 11 bp overlap
Motif DE_72h DE_72h-ZNF682_MA1599.2 11 bp overlap
ZNF766 2 datasets
Motif DE_60h DE_60h-ZNF766_MA2098.1 9 bp overlap
Motif DE_72h DE_72h-ZNF766_MA2098.1 9 bp overlap
ZNF770 1 dataset
ChIP HEK293 GSE76494.ZNF770.HEK293 212 bp overlap
ZNF784 2 datasets
Motif DE_48h DE_48h-ZNF784_MA1717.2 8 bp overlap
Motif DE_72h DE_72h-ZNF784_MA1717.2 8 bp overlap
ZSCAN4 2 datasets
Motif DE_48h DE_48h-ZSCAN4_MA1155.1 15 bp overlap
Motif DE_72h DE_72h-ZSCAN4_MA1155.1 15 bp overlap
Zfx 2 datasets
Motif DE_60h DE_60h-Zfx_MA0146.3 10 bp overlap
Motif DE_72h DE_72h-Zfx_MA0146.3 10 bp overlap
Zic1::Zic2 2 datasets
Motif DE_72h DE_72h-Zic1Zic2_MA1628.2 7 bp overlap
Motif DE_72h DE_72h-Zic1Zic2_MA1628.2 7 bp overlap
Zic2 2 datasets
Motif DE_72h DE_72h-Zic2_MA1629.2 9 bp overlap
Motif DE_72h DE_72h-Zic2_MA1629.2 9 bp overlap
Zic3 2 datasets
Motif DE_72h DE_72h-Zic3_MA0697.3 7 bp overlap
Motif DE_72h DE_72h-Zic3_MA0697.3 7 bp overlap