chr4 : 164,152,197 – 164,153,103
906 bp 102 TFs 0 linked genes
This 906 bp open chromatin element has no linked target genes and is bound by 102 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:164,147,197 – 164,158,103
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
102 transcription factors
Source
Cell type
AR 1 dataset
ChIP LNCaP-abl GSE39459.AR.LNCaP-abl 165 bp overlap
BARX2 3 datasets
Motif DE_48h DE_48h-BARX2_MA1471.2 9 bp overlap
Motif DE_60h DE_60h-BARX2_MA1471.2 9 bp overlap
Motif DE_72h DE_72h-BARX2_MA1471.2 9 bp overlap
BCL6B 3 datasets
Motif DE_48h DE_48h-BCL6B_MA0731.1 17 bp overlap
Motif DE_60h DE_60h-BCL6B_MA0731.1 17 bp overlap
Motif DE_72h DE_72h-BCL6B_MA0731.1 17 bp overlap
Cebpa 3 datasets
ChIP BLaER1 ENCFF680YXW 292 bp overlap
ChIP BLaER1 ENCFF798NMV 251 bp overlap
ChIP BLaER1 ENCFF844FIP 364 bp overlap
EGR1 1 dataset
Motif DE_60h DE_60h-EGR1_MA0162.5 10 bp overlap
EGR4 1 dataset
Motif DE_60h DE_60h-EGR4_MA0733.2 11 bp overlap
ERF::FOXO1 2 datasets
Motif DE_60h DE_60h-ERFFOXO1_MA1936.2 12 bp overlap
Motif DE_72h DE_72h-ERFFOXO1_MA1936.2 12 bp overlap
EWSR1-FLI1 2 datasets
Motif DE_60h DE_60h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_72h DE_72h-EWSR1-FLI1_MA0149.1 18 bp overlap
FOXA1 6 datasets
Motif DE_48h DE_48h-FOXA1_MA0148.5 8 bp overlap
Motif DE_60h DE_60h-FOXA1_MA0148.5 8 bp overlap
Motif DE_60h DE_60h-FOXA1_MA0148.5 8 bp overlap
Motif DE_72h DE_72h-FOXA1_MA0148.5 8 bp overlap
Motif DE_72h DE_72h-FOXA1_MA0148.5 8 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 299 bp overlap
FOXA2 6 datasets
ChIP Caco-2 GSE66218.FOXA2.Caco-2 117 bp overlap
ChIP DE DE-FOXA2-1 677 bp overlap
ChIP DE DE-FOXA2-2 681 bp overlap
Motif DE_48h DE_48h-FOXA2_MA0047.4 8 bp overlap
Motif DE_60h DE_60h-FOXA2_MA0047.4 8 bp overlap
Motif DE_72h DE_72h-FOXA2_MA0047.4 8 bp overlap
FOXA3 3 datasets
Motif DE_48h DE_48h-FOXA3_MA1683.2 7 bp overlap
Motif DE_60h DE_60h-FOXA3_MA1683.2 7 bp overlap
Motif DE_72h DE_72h-FOXA3_MA1683.2 7 bp overlap
FOXB1 5 datasets
Motif DE_48h DE_48h-FOXB1_MA0845.1 11 bp overlap
Motif DE_60h DE_60h-FOXB1_MA0845.1 11 bp overlap
Motif DE_60h DE_60h-FOXB1_MA0845.1 11 bp overlap
Motif DE_72h DE_72h-FOXB1_MA0845.1 11 bp overlap
Motif DE_72h DE_72h-FOXB1_MA0845.1 11 bp overlap
FOXC1 3 datasets
Motif DE_48h DE_48h-FOXC1_MA0032.2 11 bp overlap
Motif DE_60h DE_60h-FOXC1_MA0032.2 11 bp overlap
Motif DE_72h DE_72h-FOXC1_MA0032.2 11 bp overlap
FOXC2 4 datasets
Motif DE_48h DE_48h-FOXC2_MA0846.2 11 bp overlap
Motif DE_60h DE_60h-FOXC2_MA0846.2 11 bp overlap
Motif DE_60h DE_60h-FOXC2_MA0846.2 11 bp overlap
Motif DE_72h DE_72h-FOXC2_MA0846.2 11 bp overlap
FOXD1 3 datasets
Motif DE_48h DE_48h-FOXD1_MA0031.2 7 bp overlap
Motif DE_60h DE_60h-FOXD1_MA0031.2 7 bp overlap
Motif DE_72h DE_72h-FOXD1_MA0031.2 7 bp overlap
FOXD2 3 datasets
Motif DE_48h DE_48h-FOXD2_MA0847.4 11 bp overlap
Motif DE_60h DE_60h-FOXD2_MA0847.4 11 bp overlap
Motif DE_72h DE_72h-FOXD2_MA0847.4 11 bp overlap
FOXD3 1 dataset
Motif DE_60h DE_60h-FOXD3_MA0041.3 14 bp overlap
FOXF2 3 datasets
Motif DE_48h DE_48h-FOXF2_MA0030.2 9 bp overlap
Motif DE_60h DE_60h-FOXF2_MA0030.2 9 bp overlap
Motif DE_72h DE_72h-FOXF2_MA0030.2 9 bp overlap
FOXG1 3 datasets
Motif DE_48h DE_48h-FOXG1_MA0613.1 8 bp overlap
Motif DE_60h DE_60h-FOXG1_MA0613.1 8 bp overlap
Motif DE_72h DE_72h-FOXG1_MA0613.1 8 bp overlap
FOXH1 3 datasets
Motif DE_48h DE_48h-FOXH1_MA0479.2 8 bp overlap
Motif DE_60h DE_60h-FOXH1_MA0479.2 8 bp overlap
Motif DE_72h DE_72h-FOXH1_MA0479.2 8 bp overlap
FOXI1 3 datasets
Motif DE_48h DE_48h-FOXI1_MA0042.2 7 bp overlap
Motif DE_60h DE_60h-FOXI1_MA0042.2 7 bp overlap
Motif DE_72h DE_72h-FOXI1_MA0042.2 7 bp overlap
FOXK1 3 datasets
Motif DE_48h DE_48h-FOXK1_MA0852.3 7 bp overlap
Motif DE_60h DE_60h-FOXK1_MA0852.3 7 bp overlap
Motif DE_72h DE_72h-FOXK1_MA0852.3 7 bp overlap
FOXK2 3 datasets
Motif DE_48h DE_48h-FOXK2_MA1103.3 7 bp overlap
Motif DE_60h DE_60h-FOXK2_MA1103.3 7 bp overlap
Motif DE_72h DE_72h-FOXK2_MA1103.3 7 bp overlap
FOXL1 3 datasets
Motif DE_48h DE_48h-FOXL1_MA0033.2 7 bp overlap
Motif DE_60h DE_60h-FOXL1_MA0033.2 7 bp overlap
Motif DE_72h DE_72h-FOXL1_MA0033.2 7 bp overlap
FOXN3 4 datasets
Motif DE_48h DE_48h-FOXN3_MA1489.1 8 bp overlap
Motif DE_60h DE_60h-FOXN3_MA1489.1 8 bp overlap
Motif DE_60h DE_60h-FOXN3_MA1489.1 8 bp overlap
Motif DE_72h DE_72h-FOXN3_MA1489.1 8 bp overlap
FOXO4 3 datasets
Motif DE_48h DE_48h-FOXO4_MA0848.1 7 bp overlap
Motif DE_60h DE_60h-FOXO4_MA0848.1 7 bp overlap
Motif DE_72h DE_72h-FOXO4_MA0848.1 7 bp overlap
FOXO6 3 datasets
Motif DE_48h DE_48h-FOXO6_MA0849.1 7 bp overlap
Motif DE_60h DE_60h-FOXO6_MA0849.1 7 bp overlap
Motif DE_72h DE_72h-FOXO6_MA0849.1 7 bp overlap
FOXP1 3 datasets
Motif DE_48h DE_48h-FOXP1_MA0481.4 7 bp overlap
Motif DE_60h DE_60h-FOXP1_MA0481.4 7 bp overlap
Motif DE_72h DE_72h-FOXP1_MA0481.4 7 bp overlap
FOXP2 7 datasets
Motif DE_48h DE_48h-FOXP2_MA0593.2 9 bp overlap
Motif DE_48h DE_48h-FOXP2_MA0593.2 9 bp overlap
Motif DE_60h DE_60h-FOXP2_MA0593.2 9 bp overlap
Motif DE_60h DE_60h-FOXP2_MA0593.2 9 bp overlap
Motif DE_60h DE_60h-FOXP2_MA0593.2 9 bp overlap
Motif DE_72h DE_72h-FOXP2_MA0593.2 9 bp overlap
Motif DE_72h DE_72h-FOXP2_MA0593.2 9 bp overlap
FOXP3 3 datasets
Motif DE_48h DE_48h-FOXP3_MA0850.1 7 bp overlap
Motif DE_60h DE_60h-FOXP3_MA0850.1 7 bp overlap
Motif DE_72h DE_72h-FOXP3_MA0850.1 7 bp overlap
FOXP4 3 datasets
Motif DE_48h DE_48h-FOXP4_MA2117.1 7 bp overlap
Motif DE_60h DE_60h-FOXP4_MA2117.1 7 bp overlap
Motif DE_72h DE_72h-FOXP4_MA2117.1 7 bp overlap
FOXS1 3 datasets
Motif DE_48h DE_48h-FOXS1_MA2118.1 8 bp overlap
Motif DE_60h DE_60h-FOXS1_MA2118.1 8 bp overlap
Motif DE_72h DE_72h-FOXS1_MA2118.1 8 bp overlap
Foxf1 3 datasets
Motif DE_48h DE_48h-Foxf1_MA1606.2 7 bp overlap
Motif DE_60h DE_60h-Foxf1_MA1606.2 7 bp overlap
Motif DE_72h DE_72h-Foxf1_MA1606.2 7 bp overlap
Foxj2 3 datasets
Motif DE_48h DE_48h-Foxj2_MA0614.1 8 bp overlap
Motif DE_60h DE_60h-Foxj2_MA0614.1 8 bp overlap
Motif DE_72h DE_72h-Foxj2_MA0614.1 8 bp overlap
Foxj3 3 datasets
Motif DE_48h DE_48h-Foxj3_MA0851.2 9 bp overlap
Motif DE_60h DE_60h-Foxj3_MA0851.2 9 bp overlap
Motif DE_72h DE_72h-Foxj3_MA0851.2 9 bp overlap
Foxl2 5 datasets
Motif DE_48h DE_48h-Foxl2_MA1607.2 10 bp overlap
Motif DE_60h DE_60h-Foxl2_MA1607.2 10 bp overlap
Motif DE_60h DE_60h-Foxl2_MA1607.2 10 bp overlap
Motif DE_72h DE_72h-Foxl2_MA1607.2 10 bp overlap
Motif DE_72h DE_72h-Foxl2_MA1607.2 10 bp overlap
Foxo1 3 datasets
Motif DE_48h DE_48h-Foxo1_MA0480.3 7 bp overlap
Motif DE_60h DE_60h-Foxo1_MA0480.3 7 bp overlap
Motif DE_72h DE_72h-Foxo1_MA0480.3 7 bp overlap
Foxo3 3 datasets
Motif DE_48h DE_48h-Foxo3_MA0157.4 7 bp overlap
Motif DE_60h DE_60h-Foxo3_MA0157.4 7 bp overlap
Motif DE_72h DE_72h-Foxo3_MA0157.4 7 bp overlap
GATA2 3 datasets
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
GATA4 8 datasets
ChIP DE DE-GATA4-1 515 bp overlap
ChIP DE DE-GATA4-2 568 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 268 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 565 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 246 bp overlap
GATA6 9 datasets
ChIP Caco-2_PROLIF GSE23436.GATA6.Caco-2_PROLIF 139 bp overlap
ChIP DE DE-GATA6-1 496 bp overlap
ChIP DE DE-GATA6-2 626 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 294 bp overlap
ChIP foregut GSE117136.GATA6.foregut 431 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 400 bp overlap
GRHL2 3 datasets
Motif DE_48h DE_48h-GRHL2_MA1105.3 8 bp overlap
Motif DE_60h DE_60h-GRHL2_MA1105.3 8 bp overlap
Motif DE_72h DE_72h-GRHL2_MA1105.3 8 bp overlap
Gata3 3 datasets
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HNF4A 1 dataset
ChIP Caco-2_PROLIF GSE23436.HNF4A.Caco-2_PROLIF 79 bp overlap
HOXC12 3 datasets
Motif DE_48h DE_48h-HOXC12_MA0906.2 10 bp overlap
Motif DE_60h DE_60h-HOXC12_MA0906.2 10 bp overlap
Motif DE_72h DE_72h-HOXC12_MA0906.2 10 bp overlap
HOXD11 3 datasets
Motif DE_48h DE_48h-HOXD11_MA0908.2 9 bp overlap
Motif DE_60h DE_60h-HOXD11_MA0908.2 9 bp overlap
Motif DE_72h DE_72h-HOXD11_MA0908.2 9 bp overlap
IRF3 1 dataset
Motif DE_60h DE_60h-IRF3_MA1418.2 17 bp overlap
IRF5 1 dataset
Motif DE_60h DE_60h-IRF5_MA1420.1 14 bp overlap
KLF1 1 dataset
Motif DE_60h DE_60h-KLF1_MA0493.3 8 bp overlap
KLF10 1 dataset
Motif DE_60h DE_60h-KLF10_MA1511.2 9 bp overlap
KLF11 1 dataset
Motif DE_60h DE_60h-KLF11_MA1512.2 10 bp overlap
KLF14 1 dataset
Motif DE_60h DE_60h-KLF14_MA0740.2 9 bp overlap
KLF16 1 dataset
Motif DE_60h DE_60h-KLF16_MA0741.1 11 bp overlap
KLF17 1 dataset
Motif DE_60h DE_60h-KLF17_MA1514.2 14 bp overlap
KLF2 1 dataset
Motif DE_60h DE_60h-KLF2_MA1515.2 8 bp overlap
KLF4 1 dataset
Motif DE_60h DE_60h-KLF4_MA0039.5 8 bp overlap
KLF5 1 dataset
Motif DE_60h DE_60h-KLF5_MA0599.1 10 bp overlap
KLF9 1 dataset
Motif DE_60h DE_60h-KLF9_MA1107.3 11 bp overlap
LIN54 2 datasets
Motif DE_60h DE_60h-LIN54_MA0619.2 7 bp overlap
Motif DE_72h DE_72h-LIN54_MA0619.2 7 bp overlap
Lhx3 2 datasets
Motif DE_60h DE_60h-Lhx3_MA0135.2 12 bp overlap
Motif DE_72h DE_72h-Lhx3_MA0135.2 12 bp overlap
MEIS1 2 datasets
Motif DE_60h DE_60h-MEIS1_MA0498.3 5 bp overlap
Motif DE_72h DE_72h-MEIS1_MA0498.3 5 bp overlap
Nfat5 2 datasets
Motif DE_60h DE_60h-Nfat5_MA0606.3 8 bp overlap
Motif DE_72h DE_72h-Nfat5_MA0606.3 8 bp overlap
Nr2e3 3 datasets
Motif DE_48h DE_48h-Nr2e3_MA0164.2 6 bp overlap
Motif DE_60h DE_60h-Nr2e3_MA0164.2 6 bp overlap
Motif DE_72h DE_72h-Nr2e3_MA0164.2 6 bp overlap
ONECUT3 3 datasets
Motif DE_48h DE_48h-ONECUT3_MA0757.2 12 bp overlap
Motif DE_60h DE_60h-ONECUT3_MA0757.2 12 bp overlap
Motif DE_72h DE_72h-ONECUT3_MA0757.2 12 bp overlap
PATZ1 1 dataset
Motif DE_60h DE_60h-PATZ1_MA1961.2 11 bp overlap
PBX1 3 datasets
Motif DE_48h DE_48h-PBX1_MA0070.2 9 bp overlap
Motif DE_60h DE_60h-PBX1_MA0070.2 9 bp overlap
Motif DE_72h DE_72h-PBX1_MA0070.2 9 bp overlap
POU2F2 2 datasets
Motif DE_60h DE_60h-POU2F2_MA0507.3 13 bp overlap
Motif DE_72h DE_72h-POU2F2_MA0507.3 13 bp overlap
POU3F2 2 datasets
Motif DE_60h DE_60h-POU3F2_MA0787.1 12 bp overlap
Motif DE_72h DE_72h-POU3F2_MA0787.1 12 bp overlap
POU3F3 2 datasets
Motif DE_60h DE_60h-POU3F3_MA0788.1 13 bp overlap
Motif DE_72h DE_72h-POU3F3_MA0788.1 13 bp overlap
PPARA::RXRA 2 datasets
Motif DE_60h DE_60h-PPARARXRA_MA1148.2 17 bp overlap
Motif DE_72h DE_72h-PPARARXRA_MA1148.2 17 bp overlap
Prdm5 2 datasets
Motif DE_60h DE_60h-Prdm5_MA1999.2 11 bp overlap
Motif DE_72h DE_72h-Prdm5_MA1999.2 11 bp overlap
RBPJ 3 datasets
Motif DE_60h DE_60h-RBPJ_MA1116.2 6 bp overlap
Motif DE_60h DE_60h-RBPJ_MA1116.2 6 bp overlap
Motif DE_72h DE_72h-RBPJ_MA1116.2 6 bp overlap
RREB1 1 dataset
Motif DE_60h DE_60h-RREB1_MA0073.2 19 bp overlap
SOX10 1 dataset
Motif DE_60h DE_60h-SOX10_MA0442.3 6 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 267 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 292 bp overlap
SOX4 1 dataset
Motif DE_60h DE_60h-SOX4_MA0867.3 8 bp overlap
SP8 1 dataset
Motif DE_60h DE_60h-SP8_MA0747.2 11 bp overlap
SP9 1 dataset
Motif DE_60h DE_60h-SP9_MA1564.2 10 bp overlap
SRY 3 datasets
Motif DE_48h DE_48h-SRY_MA0084.2 7 bp overlap
Motif DE_60h DE_60h-SRY_MA0084.2 7 bp overlap
Motif DE_72h DE_72h-SRY_MA0084.2 7 bp overlap
STAT3 2 datasets
Motif DE_60h DE_60h-STAT3_MA0144.3 9 bp overlap
Motif DE_72h DE_72h-STAT3_MA0144.3 9 bp overlap
Sox11 1 dataset
Motif DE_60h DE_60h-Sox11_MA0869.3 8 bp overlap
Sox17 1 dataset
Motif DE_60h DE_60h-Sox17_MA0078.3 10 bp overlap
Sox5 1 dataset
Motif DE_60h DE_60h-Sox5_MA0087.3 8 bp overlap
Sox6 1 dataset
Motif DE_60h DE_60h-Sox6_MA0515.1 10 bp overlap
Sox7 1 dataset
Motif DE_60h DE_60h-Sox7_MA2095.1 10 bp overlap
TCF7L2 3 datasets
Motif DE_48h DE_48h-TCF7L2_MA0523.2 9 bp overlap
Motif DE_60h DE_60h-TCF7L2_MA0523.2 9 bp overlap
Motif DE_72h DE_72h-TCF7L2_MA0523.2 9 bp overlap
TEAD1 3 datasets
Motif DE_48h DE_48h-TEAD1_MA0090.4 9 bp overlap
Motif DE_60h DE_60h-TEAD1_MA0090.4 9 bp overlap
Motif DE_72h DE_72h-TEAD1_MA0090.4 9 bp overlap
TEAD2 3 datasets
Motif DE_48h DE_48h-TEAD2_MA1121.2 7 bp overlap
Motif DE_60h DE_60h-TEAD2_MA1121.2 7 bp overlap
Motif DE_72h DE_72h-TEAD2_MA1121.2 7 bp overlap
TEAD3 3 datasets
Motif DE_48h DE_48h-TEAD3_MA0808.1 8 bp overlap
Motif DE_60h DE_60h-TEAD3_MA0808.1 8 bp overlap
Motif DE_72h DE_72h-TEAD3_MA0808.1 8 bp overlap
TEAD4 3 datasets
Motif DE_48h DE_48h-TEAD4_MA0809.3 8 bp overlap
Motif DE_60h DE_60h-TEAD4_MA0809.3 8 bp overlap
Motif DE_72h DE_72h-TEAD4_MA0809.3 8 bp overlap
TP63 3 datasets
Motif DE_48h DE_48h-TP63_MA0525.2 18 bp overlap
Motif DE_60h DE_60h-TP63_MA0525.2 18 bp overlap
Motif DE_72h DE_72h-TP63_MA0525.2 18 bp overlap
TRPS1 3 datasets
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
VEZF1 1 dataset
Motif DE_60h DE_60h-VEZF1_MA1578.2 6 bp overlap
ZNF140 2 datasets
Motif DE_60h DE_60h-ZNF140_MA1589.2 19 bp overlap
Motif DE_72h DE_72h-ZNF140_MA1589.2 19 bp overlap
ZNF354A 2 datasets
Motif DE_60h DE_60h-ZNF354A_MA1978.2 20 bp overlap
Motif DE_72h DE_72h-ZNF354A_MA1978.2 20 bp overlap
ZNF384 3 datasets
Motif DE_48h DE_48h-ZNF384_MA1125.2 8 bp overlap
Motif DE_60h DE_60h-ZNF384_MA1125.2 8 bp overlap
Motif DE_72h DE_72h-ZNF384_MA1125.2 8 bp overlap
ZNF677 2 datasets
Motif DE_60h DE_60h-ZNF677_MA2101.1 12 bp overlap
Motif DE_72h DE_72h-ZNF677_MA2101.1 12 bp overlap
ZNF701 1 dataset
Motif DE_60h DE_60h-ZNF701_MA1987.2 17 bp overlap
ZNF740 1 dataset
Motif DE_60h DE_60h-ZNF740_MA0753.3 10 bp overlap
Zfp809 2 datasets
Motif DE_60h DE_60h-Zfp809_MA2125.1 9 bp overlap
Motif DE_72h DE_72h-Zfp809_MA2125.1 9 bp overlap
Zfx 3 datasets
Motif DE_48h DE_48h-Zfx_MA0146.3 10 bp overlap
Motif DE_60h DE_60h-Zfx_MA0146.3 10 bp overlap
Motif DE_72h DE_72h-Zfx_MA0146.3 10 bp overlap
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