chr4 : 18,043,988 18,044,582
594 bp 112 TFs 0 linked genes
This 594 bp open chromatin element has no linked target genes and is bound by 112 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:18,038,988 – 18,049,582
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
112 transcription factors
Source
Cell type
ALX3 1 dataset
Motif DE_60h DE_60h-ALX3_MA0634.2 6 bp overlap
ARGFX 1 dataset
Motif DE_60h DE_60h-ARGFX_MA1463.2 8 bp overlap
ATF2 1 dataset
Motif DE_60h DE_60h-ATF2_MA1632.2 10 bp overlap
CRX 4 datasets
ChIP retina_Hu20 GSE137311.CRX.retina_Hu20 389 bp overlap
ChIP retina_Hu21 GSE137311.CRX.retina_Hu21 353 bp overlap
ChIP retina_Hu3 GSE137311.CRX.retina_Hu3 366 bp overlap
ChIP retina_Hu6 GSE137311.CRX.retina_Hu6 540 bp overlap
CTCF 3 datasets
ChIP retina_Hu22 GSE137311.CTCF.retina_Hu22 481 bp overlap
ChIP retina_Hu25 GSE137311.CTCF.retina_Hu25 478 bp overlap
ChIP retina_Hu6 GSE137311.CTCF.retina_Hu6 520 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF460KDD 251 bp overlap
DRGX 1 dataset
Motif DE_60h DE_60h-DRGX_MA1481.2 6 bp overlap
DUX4 2 datasets
Motif DE_60h DE_60h-DUX4_MA0468.1 11 bp overlap
ChIP WA01 GSE94322.DUX4.WA01 228 bp overlap
DUXA 1 dataset
Motif DE_60h DE_60h-DUXA_MA0884.2 13 bp overlap
EMX1 1 dataset
Motif DE_60h DE_60h-EMX1_MA0612.3 6 bp overlap
EMX2 1 dataset
Motif DE_60h DE_60h-EMX2_MA0886.2 6 bp overlap
EN1 1 dataset
Motif DE_60h DE_60h-EN1_MA0027.3 6 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 294 bp overlap
ESR1 32 datasets
Motif DE_60h DE_60h-ESR1_MA0112.4 15 bp overlap
ChIP MCF-7 GSE48930.ESR1.MCF-7 275 bp overlap
ChIP MCF-7 GSE45822.ESR1.MCF-7 350 bp overlap
ChIP MCF-7 GSE76893.ESR1.MCF-7 259 bp overlap
ChIP MCF-7 GSE95302.ESR1.MCF-7 241 bp overlap
ChIP MCF-7 GSE103023.ESR1.MCF-7 151 bp overlap
ChIP MCF-7_E2 GSE60270.ESR1.MCF-7_E2 338 bp overlap
ChIP MCF-7_E2 GSE72249.ESR1.MCF-7_E2 237 bp overlap
ChIP MCF-7_E2 GSE59530.ESR1.MCF-7_E2 188 bp overlap
ChIP MCF-7_E2 ERP000209.ESR1.MCF-7_E2 143 bp overlap
ChIP MCF-7_E2-10min-ERalpha GSE94023.ESR1.MCF-7_E2-10min-ERalpha 151 bp overlap
ChIP MCF-7_E2-20min-ERalpha GSE94023.ESR1.MCF-7_E2-20min-ERalpha 258 bp overlap
ChIP MCF-7_E2-40min-ERalpha GSE94023.ESR1.MCF-7_E2-40min-ERalpha 249 bp overlap
ChIP MCF-7_E2-5min-ERalpha GSE94023.ESR1.MCF-7_E2-5min-ERalpha 194 bp overlap
ChIP MCF-7_E2-80min-ERalpha GSE94023.ESR1.MCF-7_E2-80min-ERalpha 208 bp overlap
ChIP MCF-7_SHCRT_E2 ERP000380.ESR1.MCF-7_SHCRT_E2 110 bp overlap
ChIP MCF-7_SHFOXA1_E2_TNF GSE59530.ESR1.MCF-7_SHFOXA1_E2_TNF 132 bp overlap
ChIP MCF-7_SICTR_E2 GSE40129.ESR1.MCF-7_SICTR_E2 113 bp overlap
ChIP MCF-7_estradiol-DHT_4h GSE99626.ESR1.MCF-7_estradiol-DHT_4h 180 bp overlap
ChIP MCF-7_estradiol-Dex_75min GSE99626.ESR1.MCF-7_estradiol-Dex_75min 151 bp overlap
ChIP MCF-7_estradiol_4h GSE99626.ESR1.MCF-7_estradiol_4h 219 bp overlap
ChIP MCF-7_jc5849 GSE126004.ESR1.MCF-7_jc5849 329 bp overlap
ChIP MCF-7_jc5850 GSE126004.ESR1.MCF-7_jc5850 280 bp overlap
ChIP MCF-7_jc5853 GSE126004.ESR1.MCF-7_jc5853 221 bp overlap
ChIP MCF-7_jc5856 GSE126004.ESR1.MCF-7_jc5856 306 bp overlap
ChIP MCF-7_parental_4-hydroxytamoxifen GSE123284.ESR1.MCF-7_parental_4-hydroxytamoxifen 200 bp overlap
ChIP MCF-7_shTEAD4 GSE125594.ESR1.MCF-7_shTEAD4 279 bp overlap
ChIP MDA-MB-134-VI_E2 GSE109103.ESR1.MDA-MB-134-VI_E2 228 bp overlap
ChIP SUM44PE_ESR1_wildtype GSE100074.ESR1.SUM44PE_ESR1_wildtype 418 bp overlap
ChIP T-47D_CR3flp GSE99479.ESR1.T-47D_CR3flp 275 bp overlap
ChIP T-47D_siCont-IL6 GSE126004.ESR1.T-47D_siCont-IL6 262 bp overlap
ChIP ZR751 ERP000783.ESR1.ZR751 201 bp overlap
ESR1_Y537C 1 dataset
ChIP SUM44PE__ESR1_mutant_LTED GSE100074.ESR1_Y537C.SUM44PE__ESR1_mutant_LTED 467 bp overlap
ESR1_Y537S 2 datasets
ChIP MCF-7_E2 GSE94493.ESR1_Y537S.MCF-7_E2 288 bp overlap
ChIP MCF-7_dox GSE94493.ESR1_Y537S.MCF-7_dox 250 bp overlap
ESR2 1 dataset
Motif DE_60h DE_60h-ESR2_MA0258.2 15 bp overlap
ESX1 1 dataset
Motif DE_60h DE_60h-ESX1_MA0644.3 7 bp overlap
EVX1 1 dataset
Motif DE_60h DE_60h-EVX1_MA0887.2 6 bp overlap
EVX2 1 dataset
Motif DE_60h DE_60h-EVX2_MA0888.2 6 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 467 bp overlap
ChIP DE DE-FOXA2-2 419 bp overlap
GATA1 4 datasets
ChIP K-562 GSE107726.GATA1.K-562 245 bp overlap
ChIP K-562_dCas9-LSD1 GSE132212.GATA1.K-562_dCas9-LSD1 179 bp overlap
ChIP K-562_enCRISPRi-LK GSE132212.GATA1.K-562_enCRISPRi-LK 220 bp overlap
ChIP erythroid-progenitor GSE124163.GATA1.erythroid-progenitor 126 bp overlap
GATA2 1 dataset
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
GATA4 4 datasets
ChIP DE DE-GATA4-1 526 bp overlap
ChIP DE DE-GATA4-2 534 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 334 bp overlap
GATA6 7 datasets
ChIP DE DE-GATA6-1 442 bp overlap
ChIP DE DE-GATA6-2 512 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 341 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 316 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 382 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 343 bp overlap
GSX1 1 dataset
Motif DE_60h DE_60h-GSX1_MA0892.2 6 bp overlap
GSX2 1 dataset
Motif DE_60h DE_60h-GSX2_MA0893.3 7 bp overlap
Gata3 1 dataset
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
HAND2 1 dataset
Motif DE_60h DE_60h-HAND2_MA1638.2 6 bp overlap
HOXA1 1 dataset
Motif DE_60h DE_60h-HOXA1_MA1495.2 6 bp overlap
HOXA2 1 dataset
Motif DE_60h DE_60h-HOXA2_MA0900.3 6 bp overlap
HOXA3 1 dataset
Motif DE_60h DE_60h-HOXA3_MA2119.1 7 bp overlap
HOXA5 1 dataset
Motif DE_60h DE_60h-HOXA5_MA0158.2 8 bp overlap
HOXA6 1 dataset
Motif DE_60h DE_60h-HOXA6_MA1497.2 7 bp overlap
HOXB1 1 dataset
Motif DE_60h DE_60h-HOXB1_MA2093.1 7 bp overlap
HOXB2 1 dataset
Motif DE_60h DE_60h-HOXB2_MA0902.3 6 bp overlap
HOXB3 1 dataset
Motif DE_60h DE_60h-HOXB3_MA0903.2 6 bp overlap
HOXB5 1 dataset
Motif DE_60h DE_60h-HOXB5_MA0904.3 6 bp overlap
HOXB6 1 dataset
Motif DE_60h DE_60h-HOXB6_MA1500.2 7 bp overlap
HOXB7 1 dataset
Motif DE_60h DE_60h-HOXB7_MA1501.2 7 bp overlap
HOXB8 1 dataset
Motif DE_60h DE_60h-HOXB8_MA1502.2 7 bp overlap
HOXC8 1 dataset
Motif DE_60h DE_60h-HOXC8_MA1505.2 6 bp overlap
HOXD3 1 dataset
Motif DE_60h DE_60h-HOXD3_MA0912.2 8 bp overlap
HOXD8 1 dataset
Motif DE_60h DE_60h-HOXD8_MA0910.3 7 bp overlap
Hmga1 1 dataset
Motif DE_60h DE_60h-Hmga1_MA2124.1 8 bp overlap
ISX 1 dataset
Motif DE_60h DE_60h-ISX_MA0654.2 6 bp overlap
LHX5 1 dataset
Motif DE_60h DE_60h-LHX5_MA1519.2 7 bp overlap
LHX6 2 datasets
Motif DE_60h DE_60h-LHX6_MA0658.2 8 bp overlap
Motif DE_60h DE_60h-LHX6_MA0658.2 8 bp overlap
LMX1A 1 dataset
Motif DE_60h DE_60h-LMX1A_MA0702.3 7 bp overlap
LMX1B 1 dataset
Motif DE_60h DE_60h-LMX1B_MA0703.3 8 bp overlap
Lhx4 1 dataset
Motif DE_60h DE_60h-Lhx4_MA0704.2 6 bp overlap
Lhx8 1 dataset
Motif DE_60h DE_60h-Lhx8_MA0705.2 6 bp overlap
MEF2A 1 dataset
Motif DE_60h DE_60h-MEF2A_MA0052.5 10 bp overlap
MEF2C 1 dataset
Motif DE_60h DE_60h-MEF2C_MA0497.2 11 bp overlap
MEF2D 3 datasets
ChIP retina_Hu13 GSE137311.MEF2D.retina_Hu13 386 bp overlap
ChIP retina_Hu21 GSE137311.MEF2D.retina_Hu21 397 bp overlap
ChIP retina_Hu25 GSE137311.MEF2D.retina_Hu25 328 bp overlap
MEOX1 1 dataset
Motif DE_60h DE_60h-MEOX1_MA0661.2 7 bp overlap
MEOX2 1 dataset
Motif DE_60h DE_60h-MEOX2_MA0706.2 7 bp overlap
MIXL1 1 dataset
Motif DE_60h DE_60h-MIXL1_MA0662.2 6 bp overlap
MNX1 1 dataset
Motif DE_60h DE_60h-MNX1_MA0707.3 6 bp overlap
MYB 1 dataset
Motif DE_60h DE_60h-MYB_MA0100.4 6 bp overlap
NEUROD1 4 datasets
ChIP D283-Med GSE92582.NEUROD1.D283-Med 351 bp overlap
ChIP D283-Med_shNEUROD1-1579 GSE92582.NEUROD1.D283-Med_shNEUROD1-1579 228 bp overlap
ChIP D341-Med_shNEUROD1-1154 GSE92582.NEUROD1.D341-Med_shNEUROD1-1154 273 bp overlap
ChIP D341-Med_shNEUROD1-1579 GSE92582.NEUROD1.D341-Med_shNEUROD1-1579 352 bp overlap
NKX6-1 1 dataset
Motif DE_60h DE_60h-NKX6-1_MA0674.2 7 bp overlap
NKX6-2 1 dataset
Motif DE_60h DE_60h-NKX6-2_MA0675.2 6 bp overlap
NOTO 1 dataset
Motif DE_60h DE_60h-NOTO_MA0710.2 7 bp overlap
NR2C1 1 dataset
Motif DE_60h DE_60h-NR2C1_MA1535.2 6 bp overlap
NR2C2 1 dataset
Motif DE_60h DE_60h-NR2C2_MA1536.2 6 bp overlap
NRIP1 1 dataset
ChIP MCF-7_E2 ERP005838.NRIP1.MCF-7_E2 166 bp overlap
NRL 1 dataset
ChIP retina_Hu13 GSE137311.NRL.retina_Hu13 288 bp overlap
Nr1H2 1 dataset
Motif DE_60h DE_60h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 1 dataset
Motif DE_60h DE_60h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 1 dataset
Motif DE_60h DE_60h-Nr1h3_MA2337.1 6 bp overlap
OTX2 1 dataset
ChIP retina_Hu7 GSE137311.OTX2.retina_Hu7 500 bp overlap
PDX1 1 dataset
Motif DE_60h DE_60h-PDX1_MA0132.3 6 bp overlap
PHF19 1 dataset
ChIP DU145_SH4 GSE135623.PHF19.DU145_SH4 239 bp overlap
PHOX2A 2 datasets
Motif DE_60h DE_60h-PHOX2A_MA0713.1 11 bp overlap
Motif DE_60h DE_60h-PHOX2A_MA0713.1 11 bp overlap
PHOX2B 1 dataset
Motif DE_60h DE_60h-PHOX2B_MA0681.3 12 bp overlap
POU4F2 1 dataset
Motif DE_60h DE_60h-POU4F2_MA0683.2 15 bp overlap
POU6F1 1 dataset
Motif DE_60h DE_60h-POU6F1_MA0628.2 6 bp overlap
PRRX1 1 dataset
Motif DE_60h DE_60h-PRRX1_MA0716.2 6 bp overlap
Ptf1A 1 dataset
Motif DE_60h DE_60h-Ptf1A_MA1618.2 9 bp overlap
RAX2 1 dataset
Motif DE_60h DE_60h-RAX2_MA0717.2 6 bp overlap
RORA 1 dataset
Motif DE_60h DE_60h-RORA_MA0072.2 11 bp overlap
RORB 1 dataset
ChIP retina_Hu22 GSE137311.RORB.retina_Hu22 346 bp overlap
SHOX 1 dataset
Motif DE_60h DE_60h-SHOX_MA0630.2 6 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 355 bp overlap
SMARCC1 1 dataset
ChIP G-401_Dox GSE71504.SMARCC1.G-401_Dox 203 bp overlap
STAT1 1 dataset
Motif DE_60h DE_60h-STAT1_MA0137.4 9 bp overlap
STAT3 1 dataset
ChIP MCF-7_jc5834 GSE126004.STAT3.MCF-7_jc5834 294 bp overlap
Shox2 1 dataset
Motif DE_60h DE_60h-Shox2_MA0720.2 6 bp overlap
TLX2 1 dataset
Motif DE_60h DE_60h-TLX2_MA1577.2 6 bp overlap
TRPS1 1 dataset
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
TWIST1 1 dataset
Motif DE_60h DE_60h-TWIST1_MA1123.3 8 bp overlap
UNCX 1 dataset
Motif DE_60h DE_60h-UNCX_MA0721.2 6 bp overlap
VAX1 1 dataset
Motif DE_60h DE_60h-VAX1_MA0722.2 7 bp overlap
VAX2 1 dataset
Motif DE_60h DE_60h-VAX2_MA0723.3 6 bp overlap
VSX1 1 dataset
Motif DE_60h DE_60h-VSX1_MA0725.2 7 bp overlap
VSX2 1 dataset
Motif DE_60h DE_60h-VSX2_MA0726.2 7 bp overlap
ZBTB18 1 dataset
Motif DE_60h DE_60h-ZBTB18_MA0698.2 11 bp overlap
ZBTB26 1 dataset
Motif DE_60h DE_60h-ZBTB26_MA1579.2 8 bp overlap
ZNF134 2 datasets
ChIP K-562 ENCSR553NTC.ZNF134.K-562 279 bp overlap
ChIP K562 ENCFF502NWS 345 bp overlap
ZNF157 1 dataset
Motif DE_60h DE_60h-ZNF157_MA2331.1 21 bp overlap
ZNF449 1 dataset
Motif DE_60h DE_60h-ZNF449_MA1656.2 10 bp overlap
ZNF85 1 dataset
Motif DE_60h DE_60h-ZNF85_MA1720.2 12 bp overlap
mix-a 1 dataset
Motif DE_60h DE_60h-mix-a_MA0621.2 7 bp overlap