chr1 : 102,993,871 102,995,366
1,495 bp 101 TFs 0 linked genes
This 1.5 kb open chromatin element has no linked target genes and is bound by 101 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr1:102,988,871 – 103,000,366
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
101 transcription factors
Source
Cell type
ATF2 4 datasets
Motif DE_36h DE_36h-ATF2_MA1632.2 10 bp overlap
Motif DE_48h DE_48h-ATF2_MA1632.2 10 bp overlap
Motif DE_60h DE_60h-ATF2_MA1632.2 10 bp overlap
Motif DE_72h DE_72h-ATF2_MA1632.2 10 bp overlap
ATF7 1 dataset
ChIP Hep-G2 ENCSR545FXC.ATF7.Hep-G2 241 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF262VBH 251 bp overlap
Crx 2 datasets
Motif DE_48h DE_48h-Crx_MA0467.3 6 bp overlap
Motif DE_60h DE_60h-Crx_MA0467.3 6 bp overlap
DPF2 2 datasets
ChIP BIN-67_ctrl GSE117734.DPF2.BIN-67_ctrl 87 bp overlap
ChIP BIN-67_ctrl GSE117734.DPF2.BIN-67_ctrl 248 bp overlap
DUX4 5 datasets
Motif DE_36h DE_36h-DUX4_MA0468.1 11 bp overlap
Motif DE_48h DE_48h-DUX4_MA0468.1 11 bp overlap
Motif DE_60h DE_60h-DUX4_MA0468.1 11 bp overlap
Motif DE_60h DE_60h-DUX4_MA0468.1 11 bp overlap
Motif DE_72h DE_72h-DUX4_MA0468.1 11 bp overlap
Dux 5 datasets
Motif DE_36h DE_36h-Dux_MA0611.3 11 bp overlap
Motif DE_48h DE_48h-Dux_MA0611.3 11 bp overlap
Motif DE_60h DE_60h-Dux_MA0611.3 11 bp overlap
Motif DE_60h DE_60h-Dux_MA0611.3 11 bp overlap
Motif DE_72h DE_72h-Dux_MA0611.3 11 bp overlap
E2F6 2 datasets
Motif DE_48h DE_48h-E2F6_MA0471.3 8 bp overlap
Motif DE_60h DE_60h-E2F6_MA0471.3 8 bp overlap
EGR4 1 dataset
Motif DE_60h DE_60h-EGR4_MA0733.2 11 bp overlap
EOMES 2 datasets
ChIP hESC GSE26097.EOMES.hESC 359 bp overlap
ChIP hESC GSE26097.EOMES.hESC 229 bp overlap
FIGLA 2 datasets
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
FOS 4 datasets
Motif DE_36h DE_36h-FOS_MA1951.2 13 bp overlap
Motif DE_48h DE_48h-FOS_MA1951.2 13 bp overlap
Motif DE_60h DE_60h-FOS_MA1951.2 13 bp overlap
Motif DE_72h DE_72h-FOS_MA1951.2 13 bp overlap
FOSB::JUN 4 datasets
Motif DE_36h DE_36h-FOSBJUN_MA1127.1 11 bp overlap
Motif DE_48h DE_48h-FOSBJUN_MA1127.1 11 bp overlap
Motif DE_60h DE_60h-FOSBJUN_MA1127.1 11 bp overlap
Motif DE_72h DE_72h-FOSBJUN_MA1127.1 11 bp overlap
FOXA1 2 datasets
ChIP 22Rv1_CST_Crispr-70 GSE123618.FOXA1.22Rv1_CST_Crispr-70 189 bp overlap
ChIP LNCaP_ETOH24H GSE58428.FOXA1.LNCaP_ETOH24H 248 bp overlap
FOXA2 4 datasets
ChIP DE DE-FOXA2-1 747 bp overlap
ChIP DE DE-FOXA2-1 548 bp overlap
ChIP DE DE-FOXA2-2 650 bp overlap
ChIP DE DE-FOXA2-2 492 bp overlap
FOXD2 2 datasets
Motif DE_60h DE_60h-FOXD2_MA0847.4 11 bp overlap
Motif DE_72h DE_72h-FOXD2_MA0847.4 11 bp overlap
FOXE1 5 datasets
Motif DE_36h DE_36h-FOXE1_MA1487.3 12 bp overlap
Motif DE_48h DE_48h-FOXE1_MA1487.3 12 bp overlap
Motif DE_60h DE_60h-FOXE1_MA1487.3 12 bp overlap
Motif DE_60h DE_60h-FOXE1_MA1487.3 12 bp overlap
Motif DE_72h DE_72h-FOXE1_MA1487.3 12 bp overlap
FOXG1 2 datasets
Motif DE_60h DE_60h-FOXG1_MA0613.1 8 bp overlap
Motif DE_72h DE_72h-FOXG1_MA0613.1 8 bp overlap
FOXN3 2 datasets
Motif DE_60h DE_60h-FOXN3_MA1489.1 8 bp overlap
Motif DE_72h DE_72h-FOXN3_MA1489.1 8 bp overlap
GATA2 5 datasets
ChIP CD34_ACY957 GSE60792.GATA2.CD34_ACY957 141 bp overlap
ChIP LNCaP_CSFCS GSE69043.GATA2.LNCaP_CSFCS 130 bp overlap
ChIP SKH1 GSE87283.GATA2.SKH1 376 bp overlap
ChIP SKH1_RUNX1-EVI1_KD GSE87283.GATA2.SKH1_RUNX1-EVI1_KD 237 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 250 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 1467 bp overlap
ChIP DE DE-GATA4-2 771 bp overlap
ChIP DE DE-GATA4-2 631 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 192 bp overlap
ChIP foregut GSE117136.GATA4.foregut 259 bp overlap
GATA6 14 datasets
ChIP DE DE-GATA6-1 783 bp overlap
ChIP DE DE-GATA6-1 539 bp overlap
ChIP DE DE-GATA6-2 766 bp overlap
ChIP DE DE-GATA6-2 601 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 267 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 340 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 510 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 478 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 506 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 333 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 503 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 265 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 332 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 277 bp overlap
GLI3 1 dataset
Motif DE_60h DE_60h-GLI3_MA1491.3 15 bp overlap
GLIS1 5 datasets
Motif DE_36h DE_36h-GLIS1_MA0735.2 15 bp overlap
Motif DE_48h DE_48h-GLIS1_MA0735.2 15 bp overlap
Motif DE_60h DE_60h-GLIS1_MA0735.2 15 bp overlap
Motif DE_60h DE_60h-GLIS1_MA0735.2 15 bp overlap
Motif DE_72h DE_72h-GLIS1_MA0735.2 15 bp overlap
GLIS2 1 dataset
Motif DE_60h DE_60h-GLIS2_MA0736.1 14 bp overlap
GLIS3 5 datasets
Motif DE_36h DE_36h-GLIS3_MA0737.1 14 bp overlap
Motif DE_48h DE_48h-GLIS3_MA0737.1 14 bp overlap
Motif DE_60h DE_60h-GLIS3_MA0737.1 14 bp overlap
Motif DE_60h DE_60h-GLIS3_MA0737.1 14 bp overlap
Motif DE_72h DE_72h-GLIS3_MA0737.1 14 bp overlap
GSC 2 datasets
Motif DE_48h DE_48h-GSC_MA0648.2 6 bp overlap
Motif DE_60h DE_60h-GSC_MA0648.2 6 bp overlap
GSC2 2 datasets
Motif DE_48h DE_48h-GSC2_MA0891.2 6 bp overlap
Motif DE_60h DE_60h-GSC2_MA0891.2 6 bp overlap
Gfi1B 1 dataset
Motif DE_60h DE_60h-Gfi1B_MA0483.2 10 bp overlap
Gli1 1 dataset
Motif DE_60h DE_60h-Gli1_MA1990.2 10 bp overlap
Gli2 1 dataset
Motif DE_60h DE_60h-Gli2_MA0734.4 9 bp overlap
Hic1 1 dataset
Motif DE_60h DE_60h-Hic1_MA0739.2 8 bp overlap
ISL2 3 datasets
Motif DE_36h DE_36h-ISL2_MA0914.2 6 bp overlap
Motif DE_48h DE_48h-ISL2_MA0914.2 6 bp overlap
Motif DE_60h DE_60h-ISL2_MA0914.2 6 bp overlap
JUN 6 datasets
Motif DE_36h DE_36h-JUN_MA0488.2 10 bp overlap
Motif DE_48h DE_48h-JUN_MA0488.2 10 bp overlap
Motif DE_60h DE_60h-JUN_MA0488.2 10 bp overlap
Motif DE_72h DE_72h-JUN_MA0488.2 10 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 414 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 441 bp overlap
JUND 4 datasets
Motif DE_36h DE_36h-JUND_MA0492.2 11 bp overlap
Motif DE_48h DE_48h-JUND_MA0492.2 11 bp overlap
Motif DE_60h DE_60h-JUND_MA0492.2 11 bp overlap
Motif DE_72h DE_72h-JUND_MA0492.2 11 bp overlap
MAX::MYC 6 datasets
Motif DE_36h DE_36h-MAXMYC_MA0059.2 10 bp overlap
Motif DE_48h DE_48h-MAXMYC_MA0059.2 10 bp overlap
Motif DE_60h DE_60h-MAXMYC_MA0059.2 10 bp overlap
Motif DE_60h DE_60h-MAXMYC_MA0059.2 10 bp overlap
Motif DE_72h DE_72h-MAXMYC_MA0059.2 10 bp overlap
Motif DE_72h DE_72h-MAXMYC_MA0059.2 10 bp overlap
MAZ 2 datasets
Motif DE_60h DE_60h-MAZ_MA1522.2 8 bp overlap
Motif DE_72h DE_72h-MAZ_MA1522.2 8 bp overlap
MGA::EVX1 2 datasets
Motif DE_48h DE_48h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_60h DE_60h-MGAEVX1_MA1960.2 11 bp overlap
MSANTD3 5 datasets
Motif DE_36h DE_36h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_48h DE_48h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_60h DE_60h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_60h DE_60h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_72h DE_72h-MSANTD3_MA1523.2 7 bp overlap
MXI1 6 datasets
Motif DE_36h DE_36h-MXI1_MA1108.3 6 bp overlap
Motif DE_48h DE_48h-MXI1_MA1108.3 6 bp overlap
Motif DE_60h DE_60h-MXI1_MA1108.3 6 bp overlap
Motif DE_60h DE_60h-MXI1_MA1108.3 6 bp overlap
Motif DE_72h DE_72h-MXI1_MA1108.3 6 bp overlap
Motif DE_72h DE_72h-MXI1_MA1108.3 6 bp overlap
MYOD1 2 datasets
Motif DE_48h DE_48h-MYOD1_MA0499.3 9 bp overlap
Motif DE_60h DE_60h-MYOD1_MA0499.3 9 bp overlap
Mecom 1 dataset
Motif DE_60h DE_60h-Mecom_MA0029.2 11 bp overlap
NFATC3 1 dataset
Motif DE_60h DE_60h-NFATC3_MA0625.3 6 bp overlap
NFKB2 2 datasets
Motif DE_48h DE_48h-NFKB2_MA0778.2 11 bp overlap
Motif DE_60h DE_60h-NFKB2_MA0778.2 11 bp overlap
Nfatc1 1 dataset
Motif DE_60h DE_60h-Nfatc1_MA0624.3 6 bp overlap
Nfatc2 1 dataset
Motif DE_60h DE_60h-Nfatc2_MA0152.3 8 bp overlap
Nkx3-2 3 datasets
Motif DE_36h DE_36h-Nkx3-2_MA0122.4 10 bp overlap
Motif DE_48h DE_48h-Nkx3-2_MA0122.4 10 bp overlap
Motif DE_60h DE_60h-Nkx3-2_MA0122.4 10 bp overlap
Nr2F6 4 datasets
Motif DE_36h DE_36h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_48h DE_48h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_60h DE_60h-Nr2F6_MA0728.1 15 bp overlap
Motif DE_72h DE_72h-Nr2F6_MA0728.1 15 bp overlap
Nrf1 2 datasets
Motif DE_60h DE_60h-Nrf1_MA0506.3 12 bp overlap
Motif DE_72h DE_72h-Nrf1_MA0506.3 12 bp overlap
OTX1 2 datasets
Motif DE_48h DE_48h-OTX1_MA0711.2 6 bp overlap
Motif DE_60h DE_60h-OTX1_MA0711.2 6 bp overlap
PATZ1 2 datasets
Motif DE_60h DE_60h-PATZ1_MA1961.2 11 bp overlap
Motif DE_72h DE_72h-PATZ1_MA1961.2 11 bp overlap
PBX1 2 datasets
Motif DE_60h DE_60h-PBX1_MA0070.2 9 bp overlap
Motif DE_72h DE_72h-PBX1_MA0070.2 9 bp overlap
PITX1 2 datasets
Motif DE_48h DE_48h-PITX1_MA0682.3 6 bp overlap
Motif DE_60h DE_60h-PITX1_MA0682.3 6 bp overlap
PITX3 2 datasets
Motif DE_48h DE_48h-PITX3_MA0714.2 6 bp overlap
Motif DE_60h DE_60h-PITX3_MA0714.2 6 bp overlap
POU2F1::SOX2 2 datasets
Motif DE_48h DE_48h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif DE_60h DE_60h-POU2F1SOX2_MA1962.1 17 bp overlap
PPARA::RXRA 2 datasets
Motif DE_48h DE_48h-PPARARXRA_MA1148.2 17 bp overlap
Motif DE_60h DE_60h-PPARARXRA_MA1148.2 17 bp overlap
PRDM9 4 datasets
Motif DE_60h DE_60h-PRDM9_MA1723.2 20 bp overlap
Motif DE_60h DE_60h-PRDM9_MA1723.2 20 bp overlap
Motif DE_72h DE_72h-PRDM9_MA1723.2 20 bp overlap
Motif DE_72h DE_72h-PRDM9_MA1723.2 20 bp overlap
Pou5f1::Sox2 2 datasets
Motif DE_48h DE_48h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_60h DE_60h-Pou5f1Sox2_MA0142.1 15 bp overlap
Ptf1A 2 datasets
Motif DE_48h DE_48h-Ptf1A_MA1620.2 8 bp overlap
Motif DE_60h DE_60h-Ptf1A_MA1620.2 8 bp overlap
RARA 5 datasets
Motif DE_36h DE_36h-RARA_MA0729.1 18 bp overlap
Motif DE_48h DE_48h-RARA_MA0729.1 18 bp overlap
Motif DE_60h DE_60h-RARA_MA0729.1 18 bp overlap
Motif DE_60h DE_60h-RARA_MA0729.1 18 bp overlap
Motif DE_72h DE_72h-RARA_MA0729.1 18 bp overlap
RBPJ 2 datasets
Motif DE_48h DE_48h-RBPJ_MA1116.2 6 bp overlap
Motif DE_60h DE_60h-RBPJ_MA1116.2 6 bp overlap
REL 1 dataset
Motif DE_60h DE_60h-REL_MA0101.1 10 bp overlap
RELA 1 dataset
Motif DE_60h DE_60h-RELA_MA0107.1 10 bp overlap
RHOXF1 2 datasets
Motif DE_48h DE_48h-RHOXF1_MA0719.2 6 bp overlap
Motif DE_60h DE_60h-RHOXF1_MA0719.2 6 bp overlap
Rarb 6 datasets
Motif DE_36h DE_36h-Rarb_MA0857.1 16 bp overlap
Motif DE_48h DE_48h-Rarb_MA0857.1 16 bp overlap
Motif DE_60h DE_60h-Rarb_MA0857.1 16 bp overlap
Motif DE_60h DE_60h-Rarb_MA0857.1 16 bp overlap
Motif DE_72h DE_72h-Rarb_MA0857.1 16 bp overlap
Motif DE_72h DE_72h-Rarb_MA0857.1 16 bp overlap
Rarg 4 datasets
Motif DE_36h DE_36h-Rarg_MA0859.2 15 bp overlap
Motif DE_48h DE_48h-Rarg_MA0859.2 15 bp overlap
Motif DE_60h DE_60h-Rarg_MA0859.2 15 bp overlap
Motif DE_72h DE_72h-Rarg_MA0859.2 15 bp overlap
SNAI2 2 datasets
Motif DE_48h DE_48h-SNAI2_MA0745.3 8 bp overlap
Motif DE_60h DE_60h-SNAI2_MA0745.3 8 bp overlap
SNAI3 2 datasets
Motif DE_48h DE_48h-SNAI3_MA1559.2 9 bp overlap
Motif DE_60h DE_60h-SNAI3_MA1559.2 9 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 319 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 384 bp overlap
SP5 8 datasets
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_36h DE_36h-SP5_MA1965.2 6 bp overlap
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
Motif DE_48h DE_48h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
Motif DE_72h DE_72h-SP5_MA1965.2 6 bp overlap
Motif DE_72h DE_72h-SP5_MA1965.2 6 bp overlap
SPI1 1 dataset
ChIP KG-1 GSE128834.SPI1.KG-1 117 bp overlap
SPIB 2 datasets
Motif DE_48h DE_48h-SPIB_MA0081.3 13 bp overlap
Motif DE_60h DE_60h-SPIB_MA0081.3 13 bp overlap
SPIC 1 dataset
Motif DE_48h DE_48h-SPIC_MA0687.2 13 bp overlap
SRY 2 datasets
Motif DE_60h DE_60h-SRY_MA0084.2 7 bp overlap
Motif DE_72h DE_72h-SRY_MA0084.2 7 bp overlap
Spi1 2 datasets
Motif DE_48h DE_48h-Spi1_MA0080.7 13 bp overlap
Motif DE_60h DE_60h-Spi1_MA0080.7 13 bp overlap
Stat6 1 dataset
Motif DE_60h DE_60h-Stat6_MA0520.2 10 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 270 bp overlap
TAL1::TCF3 3 datasets
Motif DE_36h DE_36h-TAL1TCF3_MA0091.2 10 bp overlap
Motif DE_48h DE_48h-TAL1TCF3_MA0091.2 10 bp overlap
Motif DE_60h DE_60h-TAL1TCF3_MA0091.2 10 bp overlap
TBX18 2 datasets
Motif DE_48h DE_48h-TBX18_MA1565.2 9 bp overlap
Motif DE_60h DE_60h-TBX18_MA1565.2 9 bp overlap
TFAP2A 1 dataset
Motif DE_60h DE_60h-TFAP2A_MA0003.5 9 bp overlap
TFAP2B 1 dataset
Motif DE_60h DE_60h-TFAP2B_MA0812.2 9 bp overlap
TFAP2C 1 dataset
Motif DE_60h DE_60h-TFAP2C_MA0814.3 9 bp overlap
TFAP2E 1 dataset
Motif DE_60h DE_60h-TFAP2E_MA1569.2 9 bp overlap
Tbx6 2 datasets
Motif DE_48h DE_48h-Tbx6_MA1567.3 9 bp overlap
Motif DE_60h DE_60h-Tbx6_MA1567.3 9 bp overlap
VEZF1 1 dataset
Motif DE_60h DE_60h-VEZF1_MA1578.2 6 bp overlap
ZEB1 2 datasets
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
ZIC5 2 datasets
Motif DE_60h DE_60h-ZIC5_MA1584.2 15 bp overlap
Motif DE_72h DE_72h-ZIC5_MA1584.2 15 bp overlap
ZKSCAN5 4 datasets
Motif DE_36h DE_36h-ZKSCAN5_MA1652.2 9 bp overlap
Motif DE_48h DE_48h-ZKSCAN5_MA1652.2 9 bp overlap
Motif DE_60h DE_60h-ZKSCAN5_MA1652.2 9 bp overlap
Motif DE_72h DE_72h-ZKSCAN5_MA1652.2 9 bp overlap
ZNF148 2 datasets
Motif DE_60h DE_60h-ZNF148_MA1653.2 10 bp overlap
Motif DE_72h DE_72h-ZNF148_MA1653.2 10 bp overlap
ZNF184 1 dataset
Motif DE_48h DE_48h-ZNF184_MA2120.1 13 bp overlap
ZNF35 1 dataset
Motif DE_60h DE_60h-ZNF35_MA2333.1 7 bp overlap
ZNF418 2 datasets
Motif DE_60h DE_60h-ZNF418_MA1980.1 15 bp overlap
Motif DE_72h DE_72h-ZNF418_MA1980.1 15 bp overlap
ZNF549 2 datasets
Motif DE_60h DE_60h-ZNF549_MA1728.2 8 bp overlap
Motif DE_72h DE_72h-ZNF549_MA1728.2 8 bp overlap
ZNF667 3 datasets
Motif DE_36h DE_36h-ZNF667_MA1984.2 11 bp overlap
Motif DE_48h DE_48h-ZNF667_MA1984.2 11 bp overlap
Motif DE_60h DE_60h-ZNF667_MA1984.2 11 bp overlap
ZNF701 4 datasets
Motif DE_48h DE_48h-ZNF701_MA1987.2 17 bp overlap
Motif DE_60h DE_60h-ZNF701_MA1987.2 17 bp overlap
Motif DE_60h DE_60h-ZNF701_MA1987.2 17 bp overlap
Motif DE_72h DE_72h-ZNF701_MA1987.2 17 bp overlap
ZNF740 1 dataset
Motif DE_60h DE_60h-ZNF740_MA0753.3 10 bp overlap
ZNF93 2 datasets
Motif DE_60h DE_60h-ZNF93_MA1721.2 14 bp overlap
Motif DE_72h DE_72h-ZNF93_MA1721.2 14 bp overlap