chr4 : 140,212,328 140,214,033
1,705 bp 74 TFs 3 linked genes
This 1.7 kb open chromatin element is linked to MAML3, SCOC, and CLGN and is bound by 74 transcription factors.
Linked Genes
3 genes
Link type
Gene Expression Dist. to TSS Distance Link type
MAML3 58.8 kb Distal Multiome+HiCAR
SCOC 160.7 kb Distal Multiome
CLGN 214.7 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:140,207,328 – 140,219,033
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
74 transcription factors
Source
Cell type
ARID1A 1 dataset
ChIP endometrial-epithelial-cells GSE106660.ARID1A.endometrial-epithelial-cells 236 bp overlap
ATF3 1 dataset
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 141 bp overlap
BRD2 1 dataset
ChIP SUM159PT_MGSK2801 GSE116879.BRD2.SUM159PT_MGSK2801 258 bp overlap
BRD3 3 datasets
ChIP H-1_DE GSE126661.BRD3.H-1_DE 299 bp overlap
ChIP H-1_DE GSE126661.BRD3.H-1_DE 297 bp overlap
ChIP H-1_DE GSE126661.BRD3.H-1_DE 575 bp overlap
BRD4 3 datasets
ChIP NMC24335 GSE96775.BRD4.NMC24335 382 bp overlap
ChIP SUM159PT_30nMbortezomib_8h GSE87418.BRD4.SUM159PT_30nMbortezomib_8h 224 bp overlap
ChIP SUM159PT_DMSO_72h GSE87418.BRD4.SUM159PT_DMSO_72h 289 bp overlap
CDX2 2 datasets
ChIP Caco-2_DIFF GSE23436.CDX2.Caco-2_DIFF 154 bp overlap
ChIP adult-duodenal-cell GSE115314.CDX2.adult-duodenal-cell 550 bp overlap
CEBPA 1 dataset
ChIP Kasumi-1_CEBPA-ER_E2 GSE102697.CEBPA.Kasumi-1_CEBPA-ER_E2 136 bp overlap
CTCF 9 datasets
ChIP Peyer's patch ENCFF701KWW 226 bp overlap
ChIP Peyer's patch ENCFF746TCR 253 bp overlap
ChIP Peyer's patch ENCFF828IDE 122 bp overlap
ChIP Peyers-patch ENCSR419ANE.CTCF.Peyers-patch 409 bp overlap
ChIP Peyers-patch ENCSR375VXU.CTCF.Peyers-patch 273 bp overlap
ChIP Peyers-patch ENCSR799WDT.CTCF.Peyers-patch 302 bp overlap
ChIP colon_transverse ENCSR102CSD.CTCF.colon_transverse 524 bp overlap
ChIP colon_transverse ENCSR833FWC.CTCF.colon_transverse 283 bp overlap
ChIP transverse colon ENCFF749DPF 373 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF274GAT 420 bp overlap
DAXX 1 dataset
ChIP PC-3 GSE68647.DAXX.PC-3 163 bp overlap
EGR1 1 dataset
ChIP HL-60_PMA GSE106359.EGR1.HL-60_PMA 173 bp overlap
ESR1 4 datasets
ChIP U2OS_100nM-E2 GSE151039.ESR1.U2OS_100nM-E2 476 bp overlap
ChIP U2OS_10nM-E2 GSE151039.ESR1.U2OS_10nM-E2 406 bp overlap
ChIP U2OS_10nM-E2-B GSE151039.ESR1.U2OS_10nM-E2-B 465 bp overlap
ChIP U2OS_E2 GSE26110.ESR1.U2OS_E2 279 bp overlap
EZH2 1 dataset
ChIP DND41 ENCSR000ASW.EZH2.DND41 545 bp overlap
FOS 9 datasets
ChIP CFPAC-1 GSE119930.FOS.CFPAC-1 509 bp overlap
ChIP IMR-90 ENCFF179EDA 258 bp overlap
ChIP IMR-90 ENCSR124AIG.FOS.IMR-90 280 bp overlap
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 179 bp overlap
ChIP MCF-10A ENCSR000DON.FOS.MCF-10A 221 bp overlap
ChIP MCF-7 ENCFF282FWZ 264 bp overlap
ChIP MCF-7 ENCSR569XNP.FOS.MCF-7 302 bp overlap
ChIP endothelial cell of umbilical vein ENCFF415XBG 318 bp overlap
ChIP endothelial_umbilical-vein ENCSR000EVU.FOS.endothelial_umbilical-vein 259 bp overlap
FOSL1 3 datasets
ChIP HCT-116 ENCSR000BTE.FOSL1.HCT-116 196 bp overlap
ChIP HCT116 ENCFF540ZXN 332 bp overlap
ChIP MDA-MB-231 GSE95303.FOSL1.MDA-MB-231 181 bp overlap
FOSL2 4 datasets
ChIP A-549 ENCSR448TVS.FOSL2.A-549 216 bp overlap
ChIP A549 ENCFF195CES 306 bp overlap
ChIP LPS141 GSE111253.FOSL2.LPS141 195 bp overlap
ChIP SK-N-SH ENCSR000BVB.FOSL2.SK-N-SH 141 bp overlap
FOXA1 1 dataset
ChIP CFPAC-1 GSE119930.FOXA1.CFPAC-1 337 bp overlap
FOXA2 6 datasets
ChIP CFPAC-1 GSE119930.FOXA2.CFPAC-1 299 bp overlap
ChIP CFPAC-1_EV GSE119930.FOXA2.CFPAC-1_EV 487 bp overlap
ChIP CFPAC-1_HOXB8-OE GSE119930.FOXA2.CFPAC-1_HOXB8-OE 584 bp overlap
ChIP DE DE-FOXA2-1 1429 bp overlap
ChIP DE DE-FOXA2-2 482 bp overlap
ChIP DE DE-FOXA2-2 856 bp overlap
GATA4 4 datasets
ChIP DE DE-GATA4-1 1536 bp overlap
ChIP DE DE-GATA4-2 1705 bp overlap
ChIP foregut GSE117136.GATA4.foregut 876 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 316 bp overlap
GATA6 14 datasets
ChIP DE DE-GATA6-1 1592 bp overlap
ChIP DE DE-GATA6-2 1705 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 944 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 938 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 849 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 844 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 1180 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 402 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 962 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 259 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 707 bp overlap
ChIP foregut GSE117136.GATA6.foregut 772 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 356 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 264 bp overlap
HMBOX1 2 datasets
ChIP K-562 ENCSR757IIU.HMBOX1.K-562 475 bp overlap
ChIP K562 ENCFF317JJX 449 bp overlap
HNF4A 10 datasets
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 132 bp overlap
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 172 bp overlap
ChIP GP5D GSE51234.HNF4A.GP5D 879 bp overlap
ChIP IM95 GSE114018.HNF4A.IM95 664 bp overlap
ChIP IM95 GSE114018.HNF4A.IM95 309 bp overlap
ChIP IM95 GSE114018.HNF4A.IM95 368 bp overlap
ChIP KATO-III GSE114018.HNF4A.KATO-III 94 bp overlap
ChIP KATO-III GSE114018.HNF4A.KATO-III 166 bp overlap
ChIP LoVo_PHASES GSE51290.HNF4A.LoVo_PHASES 400 bp overlap
ChIP LoVo_PHASES GSE51290.HNF4A.LoVo_PHASES 736 bp overlap
IKZF2 1 dataset
Motif DE_60h DE_60h-IKZF2_MA2326.1 6 bp overlap
IRF4 1 dataset
ChIP NCI-H929 GSE56857.IRF4.NCI-H929 85 bp overlap
JUN 8 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 540 bp overlap
ChIP DE_D1 S08-DE-d1-JUN-exp1 356 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 469 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 311 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 266 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 443 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 374 bp overlap
ChIP MCF10A-Er-Src_EtOH GSE115597.JUN.MCF10A-Er-Src_EtOH 163 bp overlap
JUNB 3 datasets
ChIP CD4 GSE116695.JUNB.CD4 263 bp overlap
ChIP CFPAC-1 GSE119930.JUNB.CFPAC-1 200 bp overlap
ChIP MCF10A-Er-Src_EtOH GSE115597.JUNB.MCF10A-Er-Src_EtOH 123 bp overlap
JUND 2 datasets
ChIP HCT116 ENCFF748ZQX 272 bp overlap
ChIP HeLa-S3 ENCSR000EDH.JUND.HeLa-S3 147 bp overlap
KLF17 1 dataset
ChIP HEK293 ENCSR065WUF.KLF17.HEK293 210 bp overlap
KLF4 1 dataset
ChIP keratinocyte_diff GSE57702.KLF4.keratinocyte_diff 182 bp overlap
KLF5 2 datasets
ChIP ESO-26 GSE132680.KLF5.ESO-26 255 bp overlap
ChIP GP5D GSE51234.KLF5.GP5D 335 bp overlap
KLF9 1 dataset
ChIP GBM1A GSE62211.KLF9.GBM1A 99 bp overlap
LDB1 1 dataset
ChIP Kasumi-1_ctrl GSE121280.LDB1.Kasumi-1_ctrl 227 bp overlap
MAX 2 datasets
ChIP HeLa-S3 ENCSR000EZF.MAX.HeLa-S3 58 bp overlap
ChIP P493-6 GSE36354.MAX.P493-6 224 bp overlap
MED1 4 datasets
ChIP HCT-116 GSE121798.MED1.HCT-116 653 bp overlap
ChIP HCT-116_shLuc GSE121798.MED1.HCT-116_shLuc 577 bp overlap
ChIP HCT-116_shLuc_thaps GSE121798.MED1.HCT-116_shLuc_thaps 587 bp overlap
ChIP myoblast GSE60026.MED1.myoblast 214 bp overlap
MEIS1 1 dataset
ChIP CHRF28811 ERR063469.MEIS1.CHRF28811 103 bp overlap
MYC 1 dataset
ChIP GP5D GSE51234.MYC.GP5D 541 bp overlap
MYCN 1 dataset
ChIP SHEP-21N GSE80151.MYCN.SHEP-21N 88 bp overlap
NANOG 5 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 242 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 480 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 248 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 199 bp overlap
NFKB1 1 dataset
ChIP MCF10A-Er-Src_TAM GSE115597.NFKB1.MCF10A-Er-Src_TAM 130 bp overlap
NR1H2 3 datasets
ChIP HT29_GW3965_48H GSE77039.NR1H2.HT29_GW3965_48H 86 bp overlap
ChIP HT29_GW3965_48H GSE77039.NR1H2.HT29_GW3965_48H 230 bp overlap
ChIP HT29_GW3965_48H GSE77039.NR1H2.HT29_GW3965_48H 280 bp overlap
NUTM1 3 datasets
ChIP NMC24335 GSE96775.NUTM1.NMC24335 446 bp overlap
ChIP NMC24335 GSE96775.NUTM1.NMC24335 264 bp overlap
ChIP embryonic-kidney_induced GSE133122.NUTM1.embryonic-kidney_induced 1029 bp overlap
Nanog 1 dataset
Motif DE_60h DE_60h-Nanog_MA2339.1 7 bp overlap
PGR 1 dataset
ChIP AB32 GSE31129.PGR.AB32 233 bp overlap
POLR2A 5 datasets
ChIP Peyer's patch ENCFF767HVN 417 bp overlap
ChIP transverse colon ENCFF098HBD 465 bp overlap
ChIP transverse colon ENCFF193UMS 255 bp overlap
ChIP transverse colon ENCFF607LKE 204 bp overlap
ChIP transverse colon ENCFF610RWV 296 bp overlap
POU5F1 3 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 255 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 266 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 229 bp overlap
PPARG 5 datasets
ChIP HT29_ROSIG_2H GSE77039.PPARG.HT29_ROSIG_2H 157 bp overlap
ChIP HT29_ROSIG_2H GSE77039.PPARG.HT29_ROSIG_2H 301 bp overlap
ChIP HT29_ROSIG_2H GSE77039.PPARG.HT29_ROSIG_2H 217 bp overlap
ChIP HT29_ROSIG_48H GSE77039.PPARG.HT29_ROSIG_48H 444 bp overlap
ChIP HT29_ROSIG_48H GSE77039.PPARG.HT29_ROSIG_48H 888 bp overlap
RAD21 1 dataset
ChIP GP5D GSE51234.RAD21.GP5D 355 bp overlap
REST 1 dataset
ChIP hippocampus GSE144226.REST.hippocampus 199 bp overlap
SMAD2 3 datasets
ChIP endoderm GSE29422.SMAD2.endoderm 217 bp overlap
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 691 bp overlap
ChIP hESC_activinA_15h GSE99202.SMAD2.hESC_activinA_15h 488 bp overlap
SMAD2-3 3 datasets
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 1270 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 821 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 357 bp overlap
SMAD2_3 6 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 813 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 769 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 843 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 793 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 918 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 313 bp overlap
SMAD3 4 datasets
ChIP NCI-H441 GSE51509.SMAD3.NCI-H441 152 bp overlap
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 197 bp overlap
ChIP NCI-H441_SIITTF1 GSE51509.SMAD3.NCI-H441_SIITTF1 501 bp overlap
ChIP PC-3_hypoxia GSE106305.SMAD3.PC-3_hypoxia 161 bp overlap
SMAD4 2 datasets
ChIP HGrC1_C134W-TGF GSE138496.SMAD4.HGrC1_C134W-TGF 151 bp overlap
ChIP endoderm GSE29422.SMAD4.endoderm 143 bp overlap
SMARCA4 12 datasets
ChIP A-549_AG15677 GSE132290.SMARCA4.A-549_AG15677 109 bp overlap
ChIP A-549_AG15678 GSE132290.SMARCA4.A-549_AG15678 96 bp overlap
ChIP A-549_AG15721 GSE132290.SMARCA4.A-549_AG15721 91 bp overlap
ChIP A-549_AG15722 GSE132290.SMARCA4.A-549_AG15722 80 bp overlap
ChIP A-549_AG15723 GSE132290.SMARCA4.A-549_AG15723 91 bp overlap
ChIP A-549_AG15724 GSE132290.SMARCA4.A-549_AG15724 66 bp overlap
ChIP A-549_AG15725 GSE132290.SMARCA4.A-549_AG15725 120 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 255 bp overlap
ChIP TTC-1240 GSE124903.SMARCA4.TTC-1240 202 bp overlap
ChIP TTC-1240_R377H GSE124903.SMARCA4.TTC-1240_R377H 275 bp overlap
ChIP TTC-1240_SMARCB1-FL GSE124903.SMARCA4.TTC-1240_SMARCB1-FL 186 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 154 bp overlap
SMARCB1 1 dataset
ChIP proliferating-human-fibroblast_ASO_Ctrl GSE131711.SMARCB1.proliferating-human-fibroblast_ASO_Ctrl 804 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S10-DE-d1-BAF155-exp1 324 bp overlap
ChIP TTC-1240_R377H GSE124903.SMARCC1.TTC-1240_R377H 233 bp overlap
SNAI2 1 dataset
ChIP PC-9_2DF GSE131687.SNAI2.PC-9_2DF 659 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 405 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 768 bp overlap
SPI1 1 dataset
ChIP Kasumi-1_SIRUNX1ETO GSE60130.SPI1.Kasumi-1_SIRUNX1ETO 177 bp overlap
STAT3 2 datasets
ChIP MCF-10A ENCSR000DOQ.STAT3.MCF-10A 142 bp overlap
ChIP MCF10A-Er-Src_TAM GSE115597.STAT3.MCF10A-Er-Src_TAM 144 bp overlap
TAL1 1 dataset
ChIP CHRF28811 ERP008568.TAL1.CHRF28811 97 bp overlap
TCF12 1 dataset
ChIP SK-N-SH ENCFF147AHB 344 bp overlap
TEAD4 1 dataset
ChIP PC-9_1DF_DMSO GSE131687.TEAD4.PC-9_1DF_DMSO 255 bp overlap
TP53 1 dataset
ChIP MDA-MB-231 GSE95303.TP53.MDA-MB-231 169 bp overlap
YY1AP1 1 dataset
ChIP PC-9_1DF GSE131687.YY1AP1.PC-9_1DF 276 bp overlap