chr18 : 36,189,006 – 36,189,191
185 bp 53 TFs 2 linked genes
This 185 bp open chromatin element is linked to MOCOS and COSMOC and is bound by 53 transcription factors.
Linked Genes
2 genes
Gene↕ Expression Dist. to TSS↑ Distance↕ Link type↕
MOCOS 1.5 kb Proximal Proximity
COSMOC 1.5 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr18:36,184,006 – 36,194,191
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
53 transcription factors
Source
Cell type
ARNT 2 datasets
ChIP K-562 ENCSR613NUC.ARNT.K-562 185 bp overlap
ChIP K562 ENCFF451RAF 184 bp overlap
ATF4 1 dataset
ChIP K-562 ENCSR145TSJ.ATF4.K-562 129 bp overlap
BCOR 1 dataset
ChIP WA01_RNF2-R GSE104690.BCOR.WA01_RNF2-R 166 bp overlap
BRD2 2 datasets
ChIP SK-MEL-147 GSE94488.BRD2.SK-MEL-147 75 bp overlap
ChIP SUM149PT_R_DMSO GSE131097.BRD2.SUM149PT_R_DMSO 133 bp overlap
BRD9 1 dataset
ChIP Mel270_DMSO GSE124720.BRD9.Mel270_DMSO 147 bp overlap
CEBPA 1 dataset
ChIP Hep-G2 ERP000209.CEBPA.Hep-G2 124 bp overlap
CEBPB 6 datasets
ChIP H1 ENCFF871PTR 185 bp overlap
ChIP HeLa-S3 ENCFF722WEG 185 bp overlap
ChIP K-562 ENCSR000EHE.CEBPB.K-562 175 bp overlap
ChIP K-562 ENCSR000BRQ.CEBPB.K-562 133 bp overlap
ChIP K562 ENCFF189VBN 124 bp overlap
ChIP WA01 ENCSR000EBV.CEBPB.WA01 160 bp overlap
CHD1 1 dataset
ChIP HeLa-S3 ENCSR392EAA.CHD1.HeLa-S3 180 bp overlap
CTCF 114 datasets
ChIP 786-O_NORMOXIA GSE78113.CTCF.786-O_NORMOXIA 152 bp overlap
ChIP A-549 ENCSR000AUF.CTCF.A-549 107 bp overlap
ChIP DOHH2 ENCFF637WNW 185 bp overlap
ChIP DOHH2 ENCSR084RDK.CTCF.DOHH2 185 bp overlap
ChIP GM06990 ENCFF471OQT 159 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 130 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 152 bp overlap
ChIP GM12864 ENCFF357DQE 185 bp overlap
ChIP GM12864 ENCSR000DRB.CTCF.GM12864 152 bp overlap
ChIP GM12869 ENCSR000DRJ.CTCF.GM12869 106 bp overlap
ChIP GM12870 ENCSR000DRK.CTCF.GM12870 103 bp overlap
ChIP GM12871 ENCSR000DRL.CTCF.GM12871 98 bp overlap
ChIP GM12872 ENCFF697BYI 185 bp overlap
ChIP GM12873 ENCSR000DRP.CTCF.GM12873 115 bp overlap
ChIP GM12878 ENCFF511URZ 143 bp overlap
ChIP GM12878 ENCSR000DKV.CTCF.GM12878 128 bp overlap
ChIP GM23338 ENCFF531QOI 185 bp overlap
ChIP GM23338 ENCFF772DML 169 bp overlap
ChIP GP5D GSE51234.CTCF.GP5D 185 bp overlap
ChIP GP5D_SIRAD21 GSE51234.CTCF.GP5D_SIRAD21 171 bp overlap
ChIP H1 ENCFF764RHO 185 bp overlap
ChIP H9 ENCFF152GTF 185 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 176 bp overlap
ChIP HAP1_clone21 GSE152721.CTCF.HAP1_clone21 171 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 170 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_2 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_2 177 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 171 bp overlap
ChIP HAP1_clone25 GSE152721.CTCF.HAP1_clone25 185 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 185 bp overlap
ChIP HEK293 ENCFF498RMM 185 bp overlap
ChIP HEK293 ENCSR617IFZ.CTCF.HEK293 158 bp overlap
ChIP HeLa GSE126990.CTCF.HeLa 185 bp overlap
ChIP HeLa-Kyoto GSE138405.CTCF.HeLa-Kyoto 185 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 171 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 182 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 185 bp overlap
ChIP HeLa-S3 ENCFF565UFR 160 bp overlap
ChIP HeLa-S3 ENCSR000AOA.CTCF.HeLa-S3 185 bp overlap
ChIP HeLa-S3 ENCSR000DLO.CTCF.HeLa-S3 121 bp overlap
ChIP HeLa-S3_biotin GSE108869.CTCF.HeLa-S3_biotin 165 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 185 bp overlap
ChIP HeLa-S3_unsynchro GSE108173.CTCF.HeLa-S3_unsynchro 162 bp overlap
ChIP HeLa_dC9Sun-D3AMut_MIR152 GSE107607.CTCF.HeLa_dC9Sun-D3AMut_MIR152 125 bp overlap
ChIP HeLa_dC9Sun-D3A_CDCC85C GSE107607.CTCF.HeLa_dC9Sun-D3A_CDCC85C 130 bp overlap
ChIP HeLa_dC9Sun-D3A_MIR152 GSE107607.CTCF.HeLa_dC9Sun-D3A_MIR152 160 bp overlap
ChIP HeLa_dC9Sun-D3A_SHB GSE107607.CTCF.HeLa_dC9Sun-D3A_SHB 164 bp overlap
ChIP Hep-G2 ERP000209.CTCF.Hep-G2 166 bp overlap
ChIP Hep-G2 ENCSR000DLS.CTCF.Hep-G2 146 bp overlap
ChIP Hep-G2 ENCSR000AMA.CTCF.Hep-G2 145 bp overlap
ChIP Hep-G2 ENCSR000DUG.CTCF.Hep-G2 124 bp overlap
ChIP Hep-G2 ENCSR000BIE.CTCF.Hep-G2 130 bp overlap
ChIP HepG2 ENCFF194VBQ 185 bp overlap
ChIP HepG2 ENCFF348BUL 163 bp overlap
ChIP HepG2 ENCFF668CTD 137 bp overlap
ChIP HepG2 ENCFF757EKU 185 bp overlap
ChIP IMR-90 GSE43070.CTCF.IMR-90 126 bp overlap
ChIP K-562 ENCSR000AKO.CTCF.K-562 185 bp overlap
ChIP K-562 GSE70764.CTCF.K-562 185 bp overlap
ChIP K-562 GSE70482.CTCF.K-562 175 bp overlap
ChIP K-562 ENCSR000EGM.CTCF.K-562 160 bp overlap
ChIP K-562 ENCSR000BPJ.CTCF.K-562 150 bp overlap
ChIP K-562 ENCSR000DMA.CTCF.K-562 136 bp overlap
ChIP K-562_7140 GSE131349.CTCF.K-562_7140 111 bp overlap
ChIP K-562_8004 GSE131349.CTCF.K-562_8004 96 bp overlap
ChIP K-562_MonoC GSE137216.CTCF.K-562_MonoC 185 bp overlap
ChIP K-562_MonoN GSE137216.CTCF.K-562_MonoN 128 bp overlap
ChIP K-562_RF2-insertion GSE140868.CTCF.K-562_RF2-insertion 156 bp overlap
ChIP K-562_RabbitC GSE137216.CTCF.K-562_RabbitC 185 bp overlap
ChIP K-562_WT GSE140868.CTCF.K-562_WT 88 bp overlap
ChIP K-562_dCas9-KRAB GSE132212.CTCF.K-562_dCas9-KRAB 177 bp overlap
ChIP K-562_dCas9-LSD1 GSE132212.CTCF.K-562_dCas9-LSD1 185 bp overlap
ChIP K-562_sgGal4 GSE132212.CTCF.K-562_sgGal4 165 bp overlap
ChIP K562 ENCFF082GOI 171 bp overlap
ChIP K562 ENCFF111MGE 180 bp overlap
ChIP K562 ENCFF400DFR 167 bp overlap
ChIP K562 ENCFF430KTH 185 bp overlap
ChIP KARPAS422 ENCSR113REG.CTCF.KARPAS422 176 bp overlap
ChIP LNCaP clone FGC ENCFF957KCI 160 bp overlap
ChIP MCF-7 ENCFF198DQX 179 bp overlap
ChIP MCF-7 ENCFF494VXA 179 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 149 bp overlap
ChIP NB4 ENCSR000DWN.CTCF.NB4 141 bp overlap
ChIP OCI-LY1 ENCFF455ESK 185 bp overlap
ChIP OCI-LY3 ENCFF939BYJ 185 bp overlap
ChIP OCI-Ly1 ENCSR072EUE.CTCF.OCI-Ly1 185 bp overlap
ChIP OCI-Ly3 ENCSR756ZKG.CTCF.OCI-Ly3 185 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 185 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 152 bp overlap
ChIP Panc1 ENCFF056JQX 185 bp overlap
ChIP Panc1 ENCFF056JQX 185 bp overlap
ChIP SK-MEL-147 GSE94488.CTCF.SK-MEL-147 104 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 185 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-000m-Flavo-240m 154 bp overlap
ChIP THP-1_Pam3csk-030m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-030m-Flavo-000m 155 bp overlap
ChIP THP-1_Pam3csk-045m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-045m-Flavo-000m 125 bp overlap
ChIP THP-1_Pam3csk-060m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-060m-Flavo-000m 144 bp overlap
ChIP THP-1_Pam3csk-120m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-120m-Flavo-000m 119 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.CTCF.THP-1_Pam3csk-150m-Flavo-030m 134 bp overlap
ChIP THP-1_Pam3csk-180m-Flavo-060m GSE103477.CTCF.THP-1_Pam3csk-180m-Flavo-060m 125 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 123 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 138 bp overlap
ChIP ZR751 ERP000783.CTCF.ZR751 120 bp overlap
ChIP epithelial_retinal_pigment ENCSR000DVI.CTCF.epithelial_retinal_pigment 133 bp overlap
ChIP erythroblast_adult_erythroblasts GSE102184.CTCF.erythroblast_adult_erythroblasts 142 bp overlap
ChIP fetal_erythroblasts GSE102184.CTCF.fetal_erythroblasts 116 bp overlap
ChIP hepatocyte ERP000395.CTCF.hepatocyte 153 bp overlap
ChIP hiPSC GSE115407.CTCF.hiPSC 185 bp overlap
ChIP hiPSC_Angelman GSE117283.CTCF.hiPSC_Angelman 163 bp overlap
ChIP hiPSC_CVI GSE106870.CTCF.hiPSC_CVI 157 bp overlap
ChIP hiPSC_Prader-Willi GSE117283.CTCF.hiPSC_Prader-Willi 133 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 100 bp overlap
ChIP leukemia_CEBPA-24h GSE142161.CTCF.leukemia_CEBPA-24h 178 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 185 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 125 bp overlap
CTCFL 1 dataset
ChIP OVCAR-8 GSE70764.CTCFL.OVCAR-8 181 bp overlap
Cebpa 3 datasets
ChIP BLaER1 ENCFF262VBH 56 bp overlap
ChIP BLaER1 ENCFF274GAT 122 bp overlap
ChIP BLaER1 ENCFF364PUR 185 bp overlap
EGR1 1 dataset
ChIP Hep-G2 ENCSR026GSW.EGR1.Hep-G2 93 bp overlap
ELF3 2 datasets
ChIP PDAC GSE64557.ELF3.PDAC 124 bp overlap
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 185 bp overlap
ELF4 1 dataset
ChIP HepG2 ENCFF752OAT 139 bp overlap
ETS1 1 dataset
ChIP HEY-A8 GSE101832.ETS1.HEY-A8 166 bp overlap
EZH2 3 datasets
ChIP SU-DHL-6 GSE134136.EZH2.SU-DHL-6 179 bp overlap
ChIP WA01 ENCSR000ASY.EZH2.WA01 82 bp overlap
ChIP astrocyte ENCSR000ARR.EZH2.astrocyte 185 bp overlap
FOS 1 dataset
ChIP CFPAC-1 GSE119930.FOS.CFPAC-1 89 bp overlap
GATA6 3 datasets
ChIP DE DE-GATA6-2 164 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 185 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 185 bp overlap
HDGF 1 dataset
ChIP K-562 ENCSR197ALX.HDGF.K-562 185 bp overlap
HMGXB4 2 datasets
ChIP Hep-G2 ENCSR174GOO.HMGXB4.Hep-G2 135 bp overlap
ChIP Hep-G2 ENCSR599XKG.HMGXB4.Hep-G2 156 bp overlap
KLF1 1 dataset
ChIP HEK293 ENCSR859BMR.KLF1.HEK293 167 bp overlap
MAZ 1 dataset
ChIP HepG2 ENCFF068NYH 185 bp overlap
MTA2 1 dataset
ChIP K-562 ENCSR411UYA.MTA2.K-562 150 bp overlap
POLR2A 2 datasets
ChIP HeLa-S3 ENCFF455YYQ 141 bp overlap
ChIP HeLa-S3 ENCFF773DNG 185 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 185 bp overlap
PRDM1 1 dataset
ChIP HEK293 ENCFF302TBP 185 bp overlap
RAD21 10 datasets
ChIP GP5D GSE51234.RAD21.GP5D 185 bp overlap
ChIP H1 ENCFF698EWO 185 bp overlap
ChIP Hep-G2 ERP000209.RAD21.Hep-G2 145 bp overlap
ChIP Hep-G2 ENCSR000EEG.RAD21.Hep-G2 142 bp overlap
ChIP Hep-G2 ENCSR054FKH.RAD21.Hep-G2 141 bp overlap
ChIP HepG2 ENCFF360ZSW 176 bp overlap
ChIP HepG2 ENCFF906QIS 185 bp overlap
ChIP K-562_RF2-insertion GSE140868.RAD21.K-562_RF2-insertion 122 bp overlap
ChIP K562 ENCFF634XYR 185 bp overlap
ChIP WA09 GSE105028.RAD21.WA09 146 bp overlap
RNF2 1 dataset
ChIP H1 ENCFF239FFS 185 bp overlap
SIN3A 1 dataset
ChIP K-562 ENCSR920BLG.SIN3A.K-562 169 bp overlap
SMARCA4 3 datasets
ChIP K-562 ENCSR587OQL.SMARCA4.K-562 185 bp overlap
ChIP K-562 ENCSR643VTW.SMARCA4.K-562 185 bp overlap
ChIP endometrial-epithelial-cells_KO GSE106660.SMARCA4.endometrial-epithelial-cells_KO 122 bp overlap
SMARCE1 1 dataset
ChIP K-562 ENCSR157TCS.SMARCE1.K-562 185 bp overlap
SMC1A 1 dataset
ChIP Hep-G2 GSE76893.SMC1A.Hep-G2 185 bp overlap
STAG1 3 datasets
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 185 bp overlap
ChIP Hep-G2 ERP000209.STAG1.Hep-G2 185 bp overlap
ChIP HepG2 ENCFF843EBZ 185 bp overlap
STAT3 1 dataset
ChIP HCC70_EtOH GSE85579.STAT3.HCC70_EtOH 114 bp overlap
TARDBP 1 dataset
ChIP GM12878 ENCSR412QBS.TARDBP.GM12878 93 bp overlap
TBP 1 dataset
ChIP HeLa-S3 ENCSR000EDD.TBP.HeLa-S3 117 bp overlap
TBX2 1 dataset
ChIP HepG2 ENCFF811TLA 185 bp overlap
TCF7L2 1 dataset
ChIP HCT-116 ENCSR000EUV.TCF7L2.HCT-116 185 bp overlap
TFAP4 3 datasets
ChIP Hep-G2 ENCSR103SZL.TFAP4.Hep-G2 184 bp overlap
ChIP HepG2 ENCFF932XOY 115 bp overlap
ChIP K562 ENCFF727PXG 185 bp overlap
THAP7 1 dataset
ChIP HepG2 ENCFF034KPY 124 bp overlap
USF2 1 dataset
ChIP HeLa-S3 ENCSR000ECW.USF2.HeLa-S3 78 bp overlap
ZBTB2 1 dataset
ChIP Hep-G2 GSE97661.ZBTB2.Hep-G2 185 bp overlap
ZBTB7B 1 dataset
ChIP Hep-G2 ENCSR322ULL.ZBTB7B.Hep-G2 140 bp overlap
ZFHX2 1 dataset
ChIP HEK293 ENCFF167TUA 185 bp overlap
ZNF184 1 dataset
ChIP K-562 ENCSR621ATC.ZNF184.K-562 185 bp overlap
ZNF253 1 dataset
ChIP HepG2 ENCFF422LRI 56 bp overlap
ZNF319 2 datasets
ChIP K562 ENCFF561ZSB 88 bp overlap
ChIP K562 ENCFF561ZSB 185 bp overlap
ZNF350 1 dataset
ChIP HepG2 ENCFF595LWL 126 bp overlap
ZNF366 1 dataset
ChIP HEK293 ENCSR106EBH.ZNF366.HEK293 94 bp overlap
ZNF462 1 dataset
ChIP GM23338 ENCFF896CCA 185 bp overlap
ZNF747 1 dataset
ChIP HepG2 ENCFF528MQU 185 bp overlap
ZNF879 1 dataset
ChIP HepG2 ENCFF479BKR 126 bp overlap
ZSCAN31 1 dataset
ChIP HepG2 ENCFF066FRL 115 bp overlap
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