chr4 : 140,515,302 140,516,703
1,401 bp 90 TFs 4 linked genes
This 1.4 kb open chromatin element is linked to 4 target genes and is bound by 90 transcription factors.
Linked Genes
4 genes
Distance
Link type
Gene Expression Dist. to TSS Distance Link type
ELMOD2 7.5 kb Proximal Proximity
CLGN 88.7 kb Distal Multiome
SCOC 142.7 kb Distal Multiome
TBC1D9 240.0 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:140,510,302 – 140,521,703
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
90 transcription factors
Source
Cell type
BCL6B 1 dataset
Motif DE_60h DE_60h-BCL6B_MA0731.1 17 bp overlap
CRX 2 datasets
ChIP retina_Hu3 GSE137311.CRX.retina_Hu3 378 bp overlap
ChIP retina_Hu6 GSE137311.CRX.retina_Hu6 330 bp overlap
CTCF 2 datasets
ChIP OCI-LY7 ENCFF086AXQ 100 bp overlap
ChIP OCI-Ly3 ENCSR756ZKG.CTCF.OCI-Ly3 179 bp overlap
DPRX 2 datasets
Motif DE_48h DE_48h-DPRX_MA1480.2 9 bp overlap
Motif DE_60h DE_60h-DPRX_MA1480.2 9 bp overlap
ELF1 1 dataset
Motif DE_60h DE_60h-ELF1_MA0473.4 9 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 223 bp overlap
EWSR1-FLI1 2 datasets
Motif DE_60h DE_60h-EWSR1-FLI1_MA0149.1 18 bp overlap
Motif DE_72h DE_72h-EWSR1-FLI1_MA0149.1 18 bp overlap
EZH2 1 dataset
ChIP neural progenitor cell ENCFF018MKA 468 bp overlap
FIGLA 1 dataset
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
FOXA2 5 datasets
ChIP DE DE-FOXA2-1 476 bp overlap
ChIP DE DE-FOXA2-1 569 bp overlap
ChIP DE DE-FOXA2-2 470 bp overlap
ChIP DE DE-FOXA2-2 426 bp overlap
Motif DE_60h DE_60h-FOXA2_MA0047.4 8 bp overlap
FOXA3 1 dataset
Motif DE_60h DE_60h-FOXA3_MA1683.2 7 bp overlap
FOXD3 1 dataset
Motif DE_60h DE_60h-FOXD3_MA0041.3 14 bp overlap
FOXE1 2 datasets
Motif DE_48h DE_48h-FOXE1_MA1487.3 12 bp overlap
Motif DE_60h DE_60h-FOXE1_MA1487.3 12 bp overlap
FOXF2 1 dataset
Motif DE_60h DE_60h-FOXF2_MA0030.2 9 bp overlap
FOXI1 1 dataset
Motif DE_60h DE_60h-FOXI1_MA0042.2 7 bp overlap
FOXJ2::ELF1 2 datasets
Motif DE_60h DE_60h-FOXJ2ELF1_MA1952.2 11 bp overlap
Motif DE_72h DE_72h-FOXJ2ELF1_MA1952.2 11 bp overlap
FOXN3 1 dataset
Motif DE_60h DE_60h-FOXN3_MA1489.1 8 bp overlap
FOXO1 1 dataset
ChIP primary-chondrocyte GSE144026.FOXO1.primary-chondrocyte 336 bp overlap
FOXO1::FLI1 2 datasets
Motif DE_60h DE_60h-FOXO1FLI1_MA1956.2 13 bp overlap
Motif DE_72h DE_72h-FOXO1FLI1_MA1956.2 13 bp overlap
FOXP1 1 dataset
Motif DE_60h DE_60h-FOXP1_MA0481.4 7 bp overlap
FOXP4 1 dataset
Motif DE_60h DE_60h-FOXP4_MA2117.1 7 bp overlap
Foxj3 1 dataset
Motif DE_60h DE_60h-Foxj3_MA0851.2 9 bp overlap
GATA1 2 datasets
Motif DE_48h DE_48h-GATA1_MA0035.5 7 bp overlap
Motif DE_60h DE_60h-GATA1_MA0035.5 7 bp overlap
GATA2 2 datasets
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
GATA4 6 datasets
ChIP DE DE-GATA4-1 501 bp overlap
ChIP DE DE-GATA4-1 533 bp overlap
ChIP DE DE-GATA4-2 1387 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 327 bp overlap
GATA5 2 datasets
Motif DE_60h DE_60h-GATA5_MA0766.3 8 bp overlap
Motif DE_72h DE_72h-GATA5_MA0766.3 8 bp overlap
GATA6 21 datasets
ChIP DE DE-GATA6-1 516 bp overlap
ChIP DE DE-GATA6-1 584 bp overlap
ChIP DE DE-GATA6-2 672 bp overlap
ChIP DE DE-GATA6-2 560 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 547 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 658 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 582 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 588 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 518 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 482 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 600 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 568 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 697 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 623 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 567 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 606 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 518 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 558 bp overlap
ChIP foregut GSE117136.GATA6.foregut 261 bp overlap
Gata3 2 datasets
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HOXB4 2 datasets
Motif DE_60h DE_60h-HOXB4_MA1499.2 6 bp overlap
Motif DE_72h DE_72h-HOXB4_MA1499.2 6 bp overlap
HOXC4 2 datasets
Motif DE_60h DE_60h-HOXC4_MA1504.2 6 bp overlap
Motif DE_72h DE_72h-HOXC4_MA1504.2 6 bp overlap
HOXD4 2 datasets
Motif DE_60h DE_60h-HOXD4_MA1507.2 6 bp overlap
Motif DE_72h DE_72h-HOXD4_MA1507.2 6 bp overlap
Ikzf3 1 dataset
Motif DE_60h DE_60h-Ikzf3_MA1992.2 9 bp overlap
KLF17 2 datasets
Motif DE_48h DE_48h-KLF17_MA1514.2 14 bp overlap
Motif DE_60h DE_60h-KLF17_MA1514.2 14 bp overlap
LIN54 2 datasets
Motif DE_48h DE_48h-LIN54_MA0619.2 7 bp overlap
Motif DE_60h DE_60h-LIN54_MA0619.2 7 bp overlap
MEIS1 1 dataset
Motif DE_60h DE_60h-MEIS1_MA0498.3 5 bp overlap
Mecom 2 datasets
Motif DE_60h DE_60h-Mecom_MA0029.2 11 bp overlap
Motif DE_72h DE_72h-Mecom_MA0029.2 11 bp overlap
NANOG 1 dataset
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 611 bp overlap
NEUROD1 1 dataset
ChIP D341-Med_shGFP GSE92582.NEUROD1.D341-Med_shGFP 272 bp overlap
NKX6-1 2 datasets
Motif DE_60h DE_60h-NKX6-1_MA0674.2 7 bp overlap
Motif DE_72h DE_72h-NKX6-1_MA0674.2 7 bp overlap
Nfat5 2 datasets
Motif DE_48h DE_48h-Nfat5_MA0606.3 8 bp overlap
Motif DE_60h DE_60h-Nfat5_MA0606.3 8 bp overlap
OTX2 2 datasets
ChIP retina_Hu7 GSE137311.OTX2.retina_Hu7 364 bp overlap
ChIP retina_pigment GSE60024.OTX2.retina_pigment 183 bp overlap
POU1F1 2 datasets
Motif DE_48h DE_48h-POU1F1_MA0784.3 14 bp overlap
Motif DE_60h DE_60h-POU1F1_MA0784.3 14 bp overlap
POU2F1 2 datasets
Motif DE_48h DE_48h-POU2F1_MA0785.2 9 bp overlap
Motif DE_60h DE_60h-POU2F1_MA0785.2 9 bp overlap
POU2F1::SOX2 3 datasets
Motif DE_48h DE_48h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif DE_48h DE_48h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif DE_60h DE_60h-POU2F1SOX2_MA1962.1 17 bp overlap
POU2F2 2 datasets
Motif DE_48h DE_48h-POU2F2_MA0507.3 13 bp overlap
Motif DE_60h DE_60h-POU2F2_MA0507.3 13 bp overlap
POU3F1 2 datasets
Motif DE_48h DE_48h-POU3F1_MA0786.2 10 bp overlap
Motif DE_60h DE_60h-POU3F1_MA0786.2 10 bp overlap
POU3F2 2 datasets
Motif DE_48h DE_48h-POU3F2_MA0787.1 12 bp overlap
Motif DE_60h DE_60h-POU3F2_MA0787.1 12 bp overlap
POU3F3 2 datasets
Motif DE_48h DE_48h-POU3F3_MA0788.1 13 bp overlap
Motif DE_60h DE_60h-POU3F3_MA0788.1 13 bp overlap
POU5F1 1 dataset
ChIP BJ_INDUCED GSE36570.POU5F1.BJ_INDUCED 124 bp overlap
POU5F1B 2 datasets
Motif DE_48h DE_48h-POU5F1B_MA0792.1 9 bp overlap
Motif DE_60h DE_60h-POU5F1B_MA0792.1 9 bp overlap
PROX1 2 datasets
Motif DE_60h DE_60h-PROX1_MA0794.1 12 bp overlap
Motif DE_72h DE_72h-PROX1_MA0794.1 12 bp overlap
Pou5f1::Sox2 3 datasets
Motif DE_48h DE_48h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_48h DE_48h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_60h DE_60h-Pou5f1Sox2_MA0142.1 15 bp overlap
Prdm15 1 dataset
Motif DE_48h DE_48h-Prdm15_MA1616.2 11 bp overlap
Ptf1A 1 dataset
Motif DE_48h DE_48h-Ptf1A_MA1620.2 8 bp overlap
RBBP5 1 dataset
ChIP WA01 ENCSR000AQC.RBBP5.WA01 142 bp overlap
SIX2 3 datasets
Motif DE_48h DE_48h-SIX2_MA1119.2 11 bp overlap
Motif DE_60h DE_60h-SIX2_MA1119.2 11 bp overlap
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 152 bp overlap
SMAD2-3 5 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 1401 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 671 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 668 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 378 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 591 bp overlap
SMAD2_3 8 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 307 bp overlap
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 593 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 418 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 763 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 555 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 534 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 807 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 1339 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 328 bp overlap
ChIP DE_D1 S15-DE-d1-BAF155-exp1 359 bp overlap
SOX10 1 dataset
Motif DE_60h DE_60h-SOX10_MA0442.3 6 bp overlap
SOX13 2 datasets
Motif DE_60h DE_60h-SOX13_MA1120.2 7 bp overlap
Motif DE_72h DE_72h-SOX13_MA1120.2 7 bp overlap
SOX17 2 datasets
ChIP DE_D2 DED2-SOX17_Batch_II 296 bp overlap
ChIP DE_D2 DED2-SOX17_Batch_II 318 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 1092 bp overlap
SOX2 2 datasets
Motif DE_60h DE_60h-SOX2_MA0143.5 7 bp overlap
Motif DE_72h DE_72h-SOX2_MA0143.5 7 bp overlap
SOX21 1 dataset
ChIP neuroepithelilal-cells GSE110505.SOX21.neuroepithelilal-cells 363 bp overlap
SOX4 2 datasets
Motif DE_60h DE_60h-SOX4_MA0867.3 8 bp overlap
Motif DE_72h DE_72h-SOX4_MA0867.3 8 bp overlap
SOX8 2 datasets
Motif DE_60h DE_60h-SOX8_MA0868.3 7 bp overlap
Motif DE_72h DE_72h-SOX8_MA0868.3 7 bp overlap
SOX9 2 datasets
Motif DE_60h DE_60h-SOX9_MA0077.2 8 bp overlap
Motif DE_72h DE_72h-SOX9_MA0077.2 8 bp overlap
SP5 1 dataset
Motif DE_60h DE_60h-SP5_MA1965.2 6 bp overlap
SRY 2 datasets
Motif DE_60h DE_60h-SRY_MA0084.2 7 bp overlap
Motif DE_72h DE_72h-SRY_MA0084.2 7 bp overlap
Sox17 2 datasets
Motif DE_60h DE_60h-Sox17_MA0078.3 10 bp overlap
Motif DE_72h DE_72h-Sox17_MA0078.3 10 bp overlap
Sox3 2 datasets
Motif DE_60h DE_60h-Sox3_MA0514.3 7 bp overlap
Motif DE_72h DE_72h-Sox3_MA0514.3 7 bp overlap
Sox5 2 datasets
Motif DE_60h DE_60h-Sox5_MA0087.3 8 bp overlap
Motif DE_72h DE_72h-Sox5_MA0087.3 8 bp overlap
Sox6 2 datasets
Motif DE_60h DE_60h-Sox6_MA0515.1 10 bp overlap
Motif DE_72h DE_72h-Sox6_MA0515.1 10 bp overlap
Sox7 2 datasets
Motif DE_60h DE_60h-Sox7_MA2095.1 10 bp overlap
Motif DE_72h DE_72h-Sox7_MA2095.1 10 bp overlap
Stat2 2 datasets
Motif DE_48h DE_48h-Stat2_MA1623.2 10 bp overlap
Motif DE_60h DE_60h-Stat2_MA1623.2 10 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 179 bp overlap
TRPS1 2 datasets
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
ZBTB18 1 dataset
Motif DE_48h DE_48h-ZBTB18_MA0698.2 11 bp overlap
ZBTB26 2 datasets
Motif DE_60h DE_60h-ZBTB26_MA1579.2 8 bp overlap
Motif DE_72h DE_72h-ZBTB26_MA1579.2 8 bp overlap
ZEB1 1 dataset
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
ZNF157 2 datasets
ChIP HEK293 ENCFF799MOR 117 bp overlap
ChIP HEK293 ENCSR564YYW.ZNF157.HEK293 241 bp overlap
ZNF384 2 datasets
Motif DE_48h DE_48h-ZNF384_MA1125.2 8 bp overlap
Motif DE_60h DE_60h-ZNF384_MA1125.2 8 bp overlap
ZNF558 2 datasets
Motif DE_60h DE_60h-ZNF558_MA2335.1 29 bp overlap
Motif DE_72h DE_72h-ZNF558_MA2335.1 29 bp overlap
ZNF766 2 datasets
Motif DE_60h DE_60h-ZNF766_MA2098.1 9 bp overlap
Motif DE_72h DE_72h-ZNF766_MA2098.1 9 bp overlap
ZNF85 2 datasets
Motif DE_48h DE_48h-ZNF85_MA1720.2 12 bp overlap
Motif DE_60h DE_60h-ZNF85_MA1720.2 12 bp overlap
Zfp335 1 dataset
Motif DE_60h DE_60h-Zfp335_MA2002.2 7 bp overlap