chr4 : 113,752,535 113,753,179
644 bp 88 TFs 1 linked gene
This 644 bp open chromatin element is linked to CAMK2D and is bound by 88 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
CAMK2D 7.9 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:113,747,535 – 113,758,179
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
88 transcription factors
Source
Cell type
BCL6B 3 datasets
Motif DE_48h DE_48h-BCL6B_MA0731.1 17 bp overlap
Motif DE_60h DE_60h-BCL6B_MA0731.1 17 bp overlap
Motif DE_72h DE_72h-BCL6B_MA0731.1 17 bp overlap
CHD4 1 dataset
ChIP pre-B-cell GSE107886.CHD4.pre-B-cell 221 bp overlap
CRX 3 datasets
ChIP retina_Hu20 GSE137311.CRX.retina_Hu20 283 bp overlap
ChIP retina_Hu3 GSE137311.CRX.retina_Hu3 343 bp overlap
ChIP retina_Hu6 GSE137311.CRX.retina_Hu6 391 bp overlap
CTCF 3 datasets
ChIP retina_Hu22 GSE137311.CTCF.retina_Hu22 318 bp overlap
ChIP retina_Hu25 GSE137311.CTCF.retina_Hu25 412 bp overlap
ChIP retina_Hu6 GSE137311.CTCF.retina_Hu6 317 bp overlap
ELF1 3 datasets
Motif DE_36h DE_36h-ELF1_MA0473.4 9 bp overlap
Motif DE_60h DE_60h-ELF1_MA0473.4 9 bp overlap
Motif DE_72h DE_72h-ELF1_MA0473.4 9 bp overlap
ELF3 3 datasets
Motif DE_36h DE_36h-ELF3_MA0640.3 9 bp overlap
Motif DE_60h DE_60h-ELF3_MA0640.3 9 bp overlap
Motif DE_72h DE_72h-ELF3_MA0640.3 9 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 191 bp overlap
ERF::FOXO1 3 datasets
Motif DE_36h DE_36h-ERFFOXO1_MA1936.2 12 bp overlap
Motif DE_60h DE_60h-ERFFOXO1_MA1936.2 12 bp overlap
Motif DE_72h DE_72h-ERFFOXO1_MA1936.2 12 bp overlap
ESR1 1 dataset
ChIP breast-cancer_3487 GSE126004.ESR1.breast-cancer_3487 163 bp overlap
ETV5::FOXI1 3 datasets
Motif DE_36h DE_36h-ETV5FOXI1_MA1946.2 12 bp overlap
Motif DE_60h DE_60h-ETV5FOXI1_MA1946.2 12 bp overlap
Motif DE_72h DE_72h-ETV5FOXI1_MA1946.2 12 bp overlap
ETV5::FOXO1 3 datasets
Motif DE_36h DE_36h-ETV5FOXO1_MA1947.2 10 bp overlap
Motif DE_60h DE_60h-ETV5FOXO1_MA1947.2 10 bp overlap
Motif DE_72h DE_72h-ETV5FOXO1_MA1947.2 10 bp overlap
Erg 3 datasets
Motif DE_36h DE_36h-Erg_MA0474.4 10 bp overlap
Motif DE_60h DE_60h-Erg_MA0474.4 10 bp overlap
Motif DE_72h DE_72h-Erg_MA0474.4 10 bp overlap
FLI1::FOXI1 3 datasets
Motif DE_36h DE_36h-FLI1FOXI1_MA1950.2 11 bp overlap
Motif DE_60h DE_60h-FLI1FOXI1_MA1950.2 11 bp overlap
Motif DE_72h DE_72h-FLI1FOXI1_MA1950.2 11 bp overlap
FOS 2 datasets
ChIP HCT-116_A7A GSE152144.FOS.HCT-116_A7A 399 bp overlap
ChIP HCT-116_A7A GSE152144.FOS.HCT-116_A7A 171 bp overlap
FOXA1 7 datasets
ChIP T-47D_shRNF2 GSE137579.FOXA1.T-47D_shRNF2 201 bp overlap
ChIP T-47D_shRNF2_24h GSE137579.FOXA1.T-47D_shRNF2_24h 155 bp overlap
ChIP ZR751 GSE72249.FOXA1.ZR751 227 bp overlap
ChIP ZR751_DEX GSE72249.FOXA1.ZR751_DEX 224 bp overlap
ChIP ZR751_E2_TAM ERP000380.FOXA1.ZR751_E2_TAM 163 bp overlap
ChIP breast-cancer_3487 GSE126004.FOXA1.breast-cancer_3487 201 bp overlap
ChIP breast-cancer_Veh-2856 GSE128018.FOXA1.breast-cancer_Veh-2856 346 bp overlap
FOXA2 9 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.FOXA2.BJ1-hTERT_FOXA2_GATA4_Coexp 248 bp overlap
ChIP Caco-2 GSE66218.FOXA2.Caco-2 155 bp overlap
ChIP DE DE-FOXA2-1 539 bp overlap
ChIP DE DE-FOXA2-2 533 bp overlap
Motif DE_36h DE_36h-FOXA2_MA0047.4 8 bp overlap
Motif DE_48h DE_48h-FOXA2_MA0047.4 8 bp overlap
Motif DE_60h DE_60h-FOXA2_MA0047.4 8 bp overlap
Motif DE_72h DE_72h-FOXA2_MA0047.4 8 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 270 bp overlap
FOXA3 4 datasets
Motif DE_36h DE_36h-FOXA3_MA1683.2 7 bp overlap
Motif DE_48h DE_48h-FOXA3_MA1683.2 7 bp overlap
Motif DE_60h DE_60h-FOXA3_MA1683.2 7 bp overlap
Motif DE_72h DE_72h-FOXA3_MA1683.2 7 bp overlap
FOXD2 2 datasets
Motif DE_48h DE_48h-FOXD2_MA0847.4 11 bp overlap
Motif DE_60h DE_60h-FOXD2_MA0847.4 11 bp overlap
FOXE1 4 datasets
Motif DE_36h DE_36h-FOXE1_MA1487.3 12 bp overlap
Motif DE_48h DE_48h-FOXE1_MA1487.3 12 bp overlap
Motif DE_60h DE_60h-FOXE1_MA1487.3 12 bp overlap
Motif DE_72h DE_72h-FOXE1_MA1487.3 12 bp overlap
FOXF2 4 datasets
Motif DE_36h DE_36h-FOXF2_MA0030.2 9 bp overlap
Motif DE_48h DE_48h-FOXF2_MA0030.2 9 bp overlap
Motif DE_60h DE_60h-FOXF2_MA0030.2 9 bp overlap
Motif DE_72h DE_72h-FOXF2_MA0030.2 9 bp overlap
FOXH1 3 datasets
Motif DE_36h DE_36h-FOXH1_MA0479.2 8 bp overlap
Motif DE_60h DE_60h-FOXH1_MA0479.2 8 bp overlap
Motif DE_72h DE_72h-FOXH1_MA0479.2 8 bp overlap
FOXI1 4 datasets
Motif DE_36h DE_36h-FOXI1_MA0042.2 7 bp overlap
Motif DE_48h DE_48h-FOXI1_MA0042.2 7 bp overlap
Motif DE_60h DE_60h-FOXI1_MA0042.2 7 bp overlap
Motif DE_72h DE_72h-FOXI1_MA0042.2 7 bp overlap
FOXO1::ELF1 3 datasets
Motif DE_36h DE_36h-FOXO1ELF1_MA1953.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1ELF1_MA1953.2 13 bp overlap
Motif DE_72h DE_72h-FOXO1ELF1_MA1953.2 13 bp overlap
FOXO1::ELK1 3 datasets
Motif DE_36h DE_36h-FOXO1ELK1_MA1954.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1ELK1_MA1954.2 13 bp overlap
Motif DE_72h DE_72h-FOXO1ELK1_MA1954.2 13 bp overlap
FOXO1::ELK3 3 datasets
Motif DE_36h DE_36h-FOXO1ELK3_MA1955.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1ELK3_MA1955.2 13 bp overlap
Motif DE_72h DE_72h-FOXO1ELK3_MA1955.2 13 bp overlap
FOXO1::FLI1 3 datasets
Motif DE_36h DE_36h-FOXO1FLI1_MA1956.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1FLI1_MA1956.2 13 bp overlap
Motif DE_72h DE_72h-FOXO1FLI1_MA1956.2 13 bp overlap
FOXP1 4 datasets
Motif DE_36h DE_36h-FOXP1_MA0481.4 7 bp overlap
Motif DE_48h DE_48h-FOXP1_MA0481.4 7 bp overlap
Motif DE_60h DE_60h-FOXP1_MA0481.4 7 bp overlap
Motif DE_72h DE_72h-FOXP1_MA0481.4 7 bp overlap
FOXP4 4 datasets
Motif DE_36h DE_36h-FOXP4_MA2117.1 7 bp overlap
Motif DE_48h DE_48h-FOXP4_MA2117.1 7 bp overlap
Motif DE_60h DE_60h-FOXP4_MA2117.1 7 bp overlap
Motif DE_72h DE_72h-FOXP4_MA2117.1 7 bp overlap
Foxj3 4 datasets
Motif DE_36h DE_36h-Foxj3_MA0851.2 9 bp overlap
Motif DE_48h DE_48h-Foxj3_MA0851.2 9 bp overlap
Motif DE_60h DE_60h-Foxj3_MA0851.2 9 bp overlap
Motif DE_72h DE_72h-Foxj3_MA0851.2 9 bp overlap
GABPA 3 datasets
Motif DE_36h DE_36h-GABPA_MA0062.4 10 bp overlap
Motif DE_60h DE_60h-GABPA_MA0062.4 10 bp overlap
Motif DE_72h DE_72h-GABPA_MA0062.4 10 bp overlap
GATA2 4 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_72h DE_72h-GATA2_MA0036.4 7 bp overlap
GATA3 1 dataset
ChIP MCF-7_E2 GSE60270.GATA3.MCF-7_E2 160 bp overlap
GATA4 11 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.GATA4.BJ1-hTERT_FOXA2_GATA4_Coexp 185 bp overlap
ChIP DE DE-GATA4-1 504 bp overlap
ChIP DE DE-GATA4-2 535 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_72h DE_72h-GATA4_MA0482.3 8 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 223 bp overlap
ChIP foregut GSE117136.GATA4.foregut 457 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 428 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 247 bp overlap
GATA6 19 datasets
ChIP DE DE-GATA6-1 454 bp overlap
ChIP DE DE-GATA6-2 475 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_72h DE_72h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 410 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 398 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 462 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 510 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 442 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 455 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 419 bp overlap
ChIP foregut GSE117136.GATA6.foregut 435 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 367 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 357 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 270 bp overlap
GRHL1 1 dataset
Motif DE_36h DE_36h-GRHL1_MA0647.2 10 bp overlap
GRHL2 3 datasets
ChIP HBE GSE46194.GRHL2.HBE 110 bp overlap
ChIP OVCA429 GSE71018.GRHL2.OVCA429 108 bp overlap
ChIP T-47D GSE99680.GRHL2.T-47D 144 bp overlap
Gata3 6 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_72h DE_72h-Gata3_MA0037.5 8 bp overlap
HOXB13 1 dataset
ChIP prostate_2483_T GSE130408.HOXB13.prostate_2483_T 118 bp overlap
Hmx1 4 datasets
Motif DE_36h DE_36h-Hmx1_MA0896.2 9 bp overlap
Motif DE_48h DE_48h-Hmx1_MA0896.2 9 bp overlap
Motif DE_60h DE_60h-Hmx1_MA0896.2 9 bp overlap
Motif DE_72h DE_72h-Hmx1_MA0896.2 9 bp overlap
Hmx3 4 datasets
Motif DE_36h DE_36h-Hmx3_MA0898.2 9 bp overlap
Motif DE_48h DE_48h-Hmx3_MA0898.2 9 bp overlap
Motif DE_60h DE_60h-Hmx3_MA0898.2 9 bp overlap
Motif DE_72h DE_72h-Hmx3_MA0898.2 9 bp overlap
KMT2A 1 dataset
ChIP L826 GSE83671.KMT2A.L826 196 bp overlap
MAFF 3 datasets
Motif DE_36h DE_36h-MAFF_MA0495.4 11 bp overlap
Motif DE_36h DE_36h-MAFF_MA0495.4 11 bp overlap
Motif DE_72h DE_72h-MAFF_MA0495.4 11 bp overlap
MEF2D 2 datasets
ChIP retina_Hu13 GSE137311.MEF2D.retina_Hu13 316 bp overlap
ChIP retina_Hu25 GSE137311.MEF2D.retina_Hu25 332 bp overlap
MEIS1 1 dataset
Motif DE_36h DE_36h-MEIS1_MA0498.3 5 bp overlap
MZF1 1 dataset
Motif DE_60h DE_60h-MZF1_MA0056.3 8 bp overlap
NEUROD1 1 dataset
ChIP D283-Med GSE92582.NEUROD1.D283-Med 193 bp overlap
NFATC3 2 datasets
Motif DE_36h DE_36h-NFATC3_MA0625.3 6 bp overlap
Motif DE_72h DE_72h-NFATC3_MA0625.3 6 bp overlap
NRL 1 dataset
ChIP retina_Hu13 GSE137311.NRL.retina_Hu13 285 bp overlap
Nanog 4 datasets
Motif DE_36h DE_36h-Nanog_MA2339.1 7 bp overlap
Motif DE_48h DE_48h-Nanog_MA2339.1 7 bp overlap
Motif DE_60h DE_60h-Nanog_MA2339.1 7 bp overlap
Motif DE_72h DE_72h-Nanog_MA2339.1 7 bp overlap
Nfatc1 2 datasets
Motif DE_36h DE_36h-Nfatc1_MA0624.3 6 bp overlap
Motif DE_72h DE_72h-Nfatc1_MA0624.3 6 bp overlap
Nfatc2 1 dataset
Motif DE_36h DE_36h-Nfatc2_MA0152.3 8 bp overlap
OTX2 1 dataset
ChIP retina_Hu7 GSE137311.OTX2.retina_Hu7 418 bp overlap
PDX1 2 datasets
ChIP hiPSC GSE125768.PDX1.hiPSC 266 bp overlap
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 150 bp overlap
PRDM9 1 dataset
Motif DE_36h DE_36h-PRDM9_MA1723.2 20 bp overlap
RB1 1 dataset
ChIP MCF-7_siH1.2 GSE98728.RB1.MCF-7_siH1.2 175 bp overlap
RBPJ 2 datasets
Motif DE_48h DE_48h-RBPJ_MA1116.2 6 bp overlap
Motif DE_60h DE_60h-RBPJ_MA1116.2 6 bp overlap
RORB 1 dataset
ChIP retina_Hu22 GSE137311.RORB.retina_Hu22 350 bp overlap
SCRT1 3 datasets
Motif DE_36h DE_36h-SCRT1_MA0743.3 10 bp overlap
Motif DE_60h DE_60h-SCRT1_MA0743.3 10 bp overlap
Motif DE_72h DE_72h-SCRT1_MA0743.3 10 bp overlap
SCRT2 3 datasets
Motif DE_36h DE_36h-SCRT2_MA0744.3 10 bp overlap
Motif DE_60h DE_60h-SCRT2_MA0744.3 10 bp overlap
Motif DE_72h DE_72h-SCRT2_MA0744.3 10 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 500 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 301 bp overlap
SMAD2_3 2 datasets
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 471 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 369 bp overlap
SOX10 3 datasets
Motif DE_48h DE_48h-SOX10_MA0442.3 6 bp overlap
Motif DE_60h DE_60h-SOX10_MA0442.3 6 bp overlap
Motif DE_72h DE_72h-SOX10_MA0442.3 6 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 275 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 299 bp overlap
SUPT5H 1 dataset
ChIP HCT-116_pThr806 GSE138548.SUPT5H.HCT-116_pThr806 170 bp overlap
SUPT6H 1 dataset
ChIP HCT-116_Inhibitor GSE130509.SUPT6H.HCT-116_Inhibitor 415 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 179 bp overlap
TCF7L2 1 dataset
ChIP hepatocellular-carcinoma-cell_420 GSE138781.TCF7L2.hepatocellular-carcinoma-cell_420 129 bp overlap
TFCP2 1 dataset
Motif DE_36h DE_36h-TFCP2_MA1968.2 9 bp overlap
THRA 2 datasets
Motif DE_60h DE_60h-THRA_MA1969.2 18 bp overlap
Motif DE_72h DE_72h-THRA_MA1969.2 18 bp overlap
THRB 2 datasets
Motif DE_60h DE_60h-THRB_MA1576.2 18 bp overlap
Motif DE_72h DE_72h-THRB_MA1576.2 18 bp overlap
TRPS1 6 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_72h DE_72h-TRPS1_MA1970.2 8 bp overlap
ZIC1 3 datasets
Motif DE_36h DE_36h-ZIC1_MA0696.1 14 bp overlap
Motif DE_60h DE_60h-ZIC1_MA0696.1 14 bp overlap
Motif DE_72h DE_72h-ZIC1_MA0696.1 14 bp overlap
ZIC5 3 datasets
Motif DE_36h DE_36h-ZIC5_MA1584.2 15 bp overlap
Motif DE_60h DE_60h-ZIC5_MA1584.2 15 bp overlap
Motif DE_72h DE_72h-ZIC5_MA1584.2 15 bp overlap
ZNF143 1 dataset
Motif DE_60h DE_60h-ZNF143_MA0088.2 16 bp overlap
ZNF184 2 datasets
Motif DE_36h DE_36h-ZNF184_MA2120.1 13 bp overlap
Motif DE_72h DE_72h-ZNF184_MA2120.1 13 bp overlap
ZNF257 2 datasets
Motif DE_36h DE_36h-ZNF257_MA1710.2 10 bp overlap
Motif DE_72h DE_72h-ZNF257_MA1710.2 10 bp overlap
ZNF558 4 datasets
Motif DE_36h DE_36h-ZNF558_MA2335.1 29 bp overlap
Motif DE_48h DE_48h-ZNF558_MA2335.1 29 bp overlap
Motif DE_60h DE_60h-ZNF558_MA2335.1 29 bp overlap
Motif DE_72h DE_72h-ZNF558_MA2335.1 29 bp overlap
ZNF675 2 datasets
Motif DE_48h DE_48h-ZNF675_MA1714.2 19 bp overlap
Motif DE_60h DE_60h-ZNF675_MA1714.2 19 bp overlap
ZNF677 3 datasets
Motif DE_48h DE_48h-ZNF677_MA2101.1 12 bp overlap
Motif DE_60h DE_60h-ZNF677_MA2101.1 12 bp overlap
Motif DE_72h DE_72h-ZNF677_MA2101.1 12 bp overlap
ZNF76 1 dataset
Motif DE_60h DE_60h-ZNF76_MA1716.2 17 bp overlap
ZNF85 3 datasets
Motif DE_48h DE_48h-ZNF85_MA1720.2 12 bp overlap
Motif DE_60h DE_60h-ZNF85_MA1720.2 12 bp overlap
Motif DE_72h DE_72h-ZNF85_MA1720.2 12 bp overlap
ZSCAN21 3 datasets
Motif DE_48h DE_48h-ZSCAN21_MA2336.1 7 bp overlap
Motif DE_60h DE_60h-ZSCAN21_MA2336.1 7 bp overlap
Motif DE_72h DE_72h-ZSCAN21_MA2336.1 7 bp overlap
ZSCAN4 4 datasets
Motif DE_36h DE_36h-ZSCAN4_MA1155.1 15 bp overlap
Motif DE_48h DE_48h-ZSCAN4_MA1155.1 15 bp overlap
Motif DE_60h DE_60h-ZSCAN4_MA1155.1 15 bp overlap
Motif DE_72h DE_72h-ZSCAN4_MA1155.1 15 bp overlap