chr9 : 36,415,916 36,416,528
612 bp 55 TFs 4 linked genes
This 612 bp open chromatin element is linked to 4 target genes and is bound by 55 transcription factors.
Linked Genes
4 genes
Gene Expression Dist. to TSS Distance Link type
RNF38 71.3 kb Distal Multiome
MELK 156.7 kb Distal Multiome
GNE 157.8 kb Distal Multiome
CLTA 225.3 kb Distal Multiome
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr9:36,410,916 – 36,421,528
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
55 transcription factors
Source
Cell type
AFF4 1 dataset
ChIP HeLa_EGF GSE40632.AFF4.HeLa_EGF 196 bp overlap
ARNTL 1 dataset
ChIP U2OS GSE44236.ARNTL.U2OS 220 bp overlap
ASH2L 1 dataset
ChIP WA01 ENCSR850KIP.ASH2L.WA01 221 bp overlap
ATF2 2 datasets
ChIP WA01 ENCSR000BQU.ATF2.WA01 157 bp overlap
ChIP WA01 ENCSR000BQU.ATF2.WA01 70 bp overlap
BRF1 1 dataset
ChIP H9_Activin GSE94418.BRF1.H9_Activin 148 bp overlap
CREB1 2 datasets
ChIP GM23338 ENCFF432ZEW 214 bp overlap
ChIP GM23338 ENCSR214ZAV.CREB1.GM23338 151 bp overlap
CTCF 136 datasets
ChIP 22Rv1 ENCSR857PBV.CTCF.22Rv1 211 bp overlap
ChIP 22Rv1_hydroxy_10nM_4h ENCSR847XGE.CTCF.22Rv1_hydroxy_10nM_4h 222 bp overlap
ChIP 786-O_NORMOXIA GSE78113.CTCF.786-O_NORMOXIA 230 bp overlap
ChIP A-549 ENCSR000AUE.CTCF.A-549 254 bp overlap
ChIP A-549 ENCSR000AUF.CTCF.A-549 270 bp overlap
ChIP A-549 ENCSR000DPF.CTCF.A-549 152 bp overlap
ChIP A-549 ENCSR000BHV.CTCF.A-549 105 bp overlap
ChIP A-673 ENCSR611JJS.CTCF.A-673 196 bp overlap
ChIP A549 ENCFF034FVO 224 bp overlap
ChIP A549 ENCFF669BWC 328 bp overlap
ChIP A673 ENCFF123WOM 215 bp overlap
ChIP BE2C ENCFF757SRF 176 bp overlap
ChIP D54 ENCSR000DKN.CTCF.D54 118 bp overlap
ChIP D721Med ENCFF513FYD 136 bp overlap
ChIP DND-41 ENCFF913MRA 186 bp overlap
ChIP DND41 ENCSR000AQU.CTCF.DND41 212 bp overlap
ChIP DOHH2 ENCFF637WNW 497 bp overlap
ChIP DOHH2 ENCFF637WNW 145 bp overlap
ChIP DOHH2 ENCSR084RDK.CTCF.DOHH2 260 bp overlap
ChIP GM04604 GSE148179.CTCF.GM04604 163 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 170 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 172 bp overlap
ChIP GM12871 ENCSR000DRL.CTCF.GM12871 106 bp overlap
ChIP GM23338 ENCFF531QOI 201 bp overlap
ChIP GM23338 ENCFF772DML 144 bp overlap
ChIP GP5D GSE51234.CTCF.GP5D 508 bp overlap
ChIP GP5D_SIRAD21 GSE51234.CTCF.GP5D_SIRAD21 301 bp overlap
ChIP H1 ENCFF230QSV 140 bp overlap
ChIP H1 ENCFF414GZI 159 bp overlap
ChIP H1 ENCFF764RHO 188 bp overlap
ChIP H9 ENCFF152GTF 196 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 230 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 160 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_2 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_2 215 bp overlap
ChIP HAP1_clone25 GSE152721.CTCF.HAP1_clone25 197 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 208 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 223 bp overlap
ChIP HCT-116 ENCSR000BSE.CTCF.HCT-116 171 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.CTCF.HCT-116_RAD21-mAC 347 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.CTCF.HCT-116_RAD21-mAC_500uM_auxin 162 bp overlap
ChIP HCT116 ENCFF003KHP 206 bp overlap
ChIP HCT116 ENCFF373YMA 267 bp overlap
ChIP HFFc6 ENCFF005CJI 307 bp overlap
ChIP HT-1080_CBL0137-6h GSE122462.CTCF.HT-1080_CBL0137-6h 281 bp overlap
ChIP HUES-64 GSE97394.CTCF.HUES-64 204 bp overlap
ChIP HUES-64_DNMT-KO GSE97394.CTCF.HUES-64_DNMT-KO 242 bp overlap
ChIP HeLa GSE126990.CTCF.HeLa 218 bp overlap
ChIP HeLa-Kyoto GSE102884.CTCF.HeLa-Kyoto 217 bp overlap
ChIP HeLa-Kyoto GSE138405.CTCF.HeLa-Kyoto 218 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 242 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 254 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 202 bp overlap
ChIP HeLa-S3 ENCFF565UFR 161 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 227 bp overlap
ChIP HeLa_dC9Sun-D3A_CDCC85C GSE107607.CTCF.HeLa_dC9Sun-D3A_CDCC85C 157 bp overlap
ChIP HeLa_dC9Sun-D3A_MIR152 GSE107607.CTCF.HeLa_dC9Sun-D3A_MIR152 204 bp overlap
ChIP HepG2 ENCFF348BUL 131 bp overlap
ChIP HepG2 ENCFF757EKU 242 bp overlap
ChIP ID00014 GSE76922.CTCF.ID00014 212 bp overlap
ChIP K-562 GSE70764.CTCF.K-562 188 bp overlap
ChIP K-562 ENCSR000EGM.CTCF.K-562 147 bp overlap
ChIP K-562 ENCSR000DMA.CTCF.K-562 159 bp overlap
ChIP K-562_MonoC GSE137216.CTCF.K-562_MonoC 189 bp overlap
ChIP K-562_RF2-insertion GSE140868.CTCF.K-562_RF2-insertion 81 bp overlap
ChIP K-562_RabbitC GSE137216.CTCF.K-562_RabbitC 186 bp overlap
ChIP K562 ENCFF082GOI 185 bp overlap
ChIP K562 ENCFF400DFR 189 bp overlap
ChIP KARPAS422 ENCSR113REG.CTCF.KARPAS422 365 bp overlap
ChIP Kelly_sensitive GSE103030.CTCF.Kelly_sensitive 182 bp overlap
ChIP LNCAP ENCFF223HIG 445 bp overlap
ChIP Loucy ENCFF359TVQ 228 bp overlap
ChIP Loucy ENCSR464DKE.CTCF.Loucy 304 bp overlap
ChIP MCF-7 ENCFF198DQX 130 bp overlap
ChIP MCF-7 ENCFF414SZG 147 bp overlap
ChIP MCF-7 ENCFF494VXA 130 bp overlap
ChIP MCF-7 ENCFF844STM 147 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 183 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 122 bp overlap
ChIP MCF-7 GSE137216.CTCF.MCF-7 202 bp overlap
ChIP MCF-7 GSE70764.CTCF.MCF-7 137 bp overlap
ChIP MCF-7_HYPOXIA GSE78113.CTCF.MCF-7_HYPOXIA 245 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 240 bp overlap
ChIP OCI-LY1 ENCFF455ESK 244 bp overlap
ChIP OCI-LY3 ENCFF939BYJ 282 bp overlap
ChIP OCI-LY7 ENCFF086AXQ 294 bp overlap
ChIP OCI-Ly1 ENCSR072EUE.CTCF.OCI-Ly1 219 bp overlap
ChIP OCI-Ly3 ENCSR756ZKG.CTCF.OCI-Ly3 220 bp overlap
ChIP OCI-Ly7 ENCSR027HML.CTCF.OCI-Ly7 250 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 171 bp overlap
ChIP PC-3 ENCFF487TUI 326 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 216 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 163 bp overlap
ChIP RWPE-2 ENCSR856JJB.CTCF.RWPE-2 231 bp overlap
ChIP SH-SY5Y GSE101295.CTCF.SH-SY5Y 115 bp overlap
ChIP SK-N-AS GSE101295.CTCF.SK-N-AS 136 bp overlap
ChIP SK-N-SH ENCSR000EIC.CTCF.SK-N-SH 167 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 298 bp overlap
ChIP SU-DHL-6 ENCSR125DKL.CTCF.SU-DHL-6 72 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-000m-Flavo-000m 193 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-020m-Flavo-000m 196 bp overlap
ChIP THP-1_Pam3csk-045m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-045m-Flavo-000m 166 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.CTCF.THP-1_Pam3csk-150m-Flavo-030m 162 bp overlap
ChIP THP-1_Pam3csk-180m-Flavo-060m GSE103477.CTCF.THP-1_Pam3csk-180m-Flavo-060m 213 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 200 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 90 bp overlap
ChIP WA01_MESEN_STEM GSE52457.CTCF.WA01_MESEN_STEM 144 bp overlap
ChIP WERI-Rb-1 ENCFF349QKF 149 bp overlap
ChIP WERI-Rb-1 ENCSR000DXW.CTCF.WERI-Rb-1 66 bp overlap
ChIP cardiac-muscle ENCSR713SXF.CTCF.cardiac-muscle 313 bp overlap
ChIP endodermal cell ENCFF471YCZ 148 bp overlap
ChIP endothelial cell of umbilical vein ENCFF455OQM 121 bp overlap
ChIP erythroblast_adult_erythroblasts GSE102184.CTCF.erythroblast_adult_erythroblasts 203 bp overlap
ChIP foreskin keratinocyte ENCFF980OWR 224 bp overlap
ChIP hESC_D0 GSE116862.CTCF.hESC_D0 211 bp overlap
ChIP hESC_NAIVE GSE69646.CTCF.hESC_NAIVE 275 bp overlap
ChIP hESC_PRIMED GSE69646.CTCF.hESC_PRIMED 244 bp overlap
ChIP hiPSC GSE115407.CTCF.hiPSC 233 bp overlap
ChIP hiPSC_Angelman GSE117283.CTCF.hiPSC_Angelman 165 bp overlap
ChIP hiPSC_CVI GSE106870.CTCF.hiPSC_CVI 155 bp overlap
ChIP hiPSC_HUES9 GSE106870.CTCF.hiPSC_HUES9 200 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 190 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 253 bp overlap
ChIP hiPSC_IIA11 GSE106870.CTCF.hiPSC_IIA11 217 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 253 bp overlap
ChIP hiPSC_Prader-Willi GSE117283.CTCF.hiPSC_Prader-Willi 180 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 371 bp overlap
ChIP keratinocyte ENCFF046PBT 144 bp overlap
ChIP keratinocyte GSE154221.CTCF.keratinocyte 433 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 119 bp overlap
ChIP keratinocyte ENCSR000ALJ.CTCF.keratinocyte 164 bp overlap
ChIP leukemia GSE142161.CTCF.leukemia 260 bp overlap
ChIP neural progenitor cell ENCFF420RBO 253 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 233 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 195 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d0 119 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 253 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF274GAT 153 bp overlap
E2F6 1 dataset
ChIP WA01 ENCSR000BSI.E2F6.WA01 147 bp overlap
EBF1 3 datasets
ChIP GM12878 ENCFF813OXE 265 bp overlap
ChIP MUTUL GSE75503.EBF1.MUTUL 554 bp overlap
ChIP NALM-6 GSE126300.EBF1.NALM-6 419 bp overlap
ETS1 1 dataset
ChIP cardiomyocyte_D0 GSE129986.ETS1.cardiomyocyte_D0 157 bp overlap
GABPA 1 dataset
ChIP WA01 ENCSR000BIW.GABPA.WA01 152 bp overlap
GATA2 1 dataset
ChIP hiPSC_SLC6ebs GSE107639.GATA2.hiPSC_SLC6ebs 111 bp overlap
HDAC2 1 dataset
ChIP WA01 ENCSR000BNR.HDAC2.WA01 115 bp overlap
HIC2 2 datasets
Motif DE_12h DE_12h-HIC2_MA0738.2 6 bp overlap
Motif ES_0h ES_0h-HIC2_MA0738.2 6 bp overlap
IKZF1 1 dataset
ChIP pre-B-cell GSE107886.IKZF1.pre-B-cell 232 bp overlap
INO80 1 dataset
ChIP Huh-7 GSE97411.INO80.Huh-7 437 bp overlap
JUN 1 dataset
ChIP ESC S34-ESC-d0-JUN-exp2 192 bp overlap
KDM1A 1 dataset
ChIP H1 ENCFF696SGD 505 bp overlap
KLF4 1 dataset
ChIP WIBR3 GSE130417.KLF4.WIBR3 223 bp overlap
KMT2A 1 dataset
ChIP RCH-ACV GSE83671.KMT2A.RCH-ACV 203 bp overlap
MCRS1 1 dataset
ChIP Huh-7 GSE97411.MCRS1.Huh-7 262 bp overlap
MED1 1 dataset
ChIP MCF-7_SHRARS GSE60270.MED1.MCF-7_SHRARS 102 bp overlap
MEIS1 2 datasets
Motif DE_12h DE_12h-MEIS1_MA0498.3 5 bp overlap
Motif ES_0h ES_0h-MEIS1_MA0498.3 5 bp overlap
NANOG 3 datasets
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 421 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 319 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 188 bp overlap
NR2F6 2 datasets
ChIP K-562 ENCSR707QWA.NR2F6.K-562 161 bp overlap
ChIP K562 ENCFF674RQA 271 bp overlap
OVOL1 2 datasets
Motif DE_12h DE_12h-OVOL1_MA1544.2 10 bp overlap
Motif ES_0h ES_0h-OVOL1_MA1544.2 10 bp overlap
OVOL2 2 datasets
Motif DE_12h DE_12h-OVOL2_MA1545.2 7 bp overlap
Motif ES_0h ES_0h-OVOL2_MA1545.2 7 bp overlap
PCBP1 1 dataset
ChIP Hep-G2 GSE120104.PCBP1.Hep-G2 171 bp overlap
POLR2A 2 datasets
ChIP H1 ENCFF566JSR 473 bp overlap
ChIP H1 ENCFF833NJP 372 bp overlap
POU5F1 3 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 376 bp overlap
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 328 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 612 bp overlap
PPARG 1 dataset
ChIP HUVEC-C_DMSO_HYPO GSE50144.PPARG.HUVEC-C_DMSO_HYPO 147 bp overlap
PRDM14 1 dataset
ChIP hESC GSE138674.PRDM14.hESC 218 bp overlap
RAD21 8 datasets
ChIP H1 ENCFF698EWO 186 bp overlap
ChIP H1 ENCFF967OJF 182 bp overlap
ChIP HCT-116 GSE131606.RAD21.HCT-116 194 bp overlap
ChIP HepG2 ENCFF360ZSW 159 bp overlap
ChIP WA09_heat-shock GSE105028.RAD21.WA09_heat-shock 178 bp overlap
ChIP hiPSC_IV8 GSE106870.RAD21.hiPSC_IV8 162 bp overlap
ChIP hiPSC_IV8 GSE106870.RAD21.hiPSC_IV8 153 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 161 bp overlap
RUNX1 1 dataset
ChIP 697 GSE138031.RUNX1.697 277 bp overlap
RUVBL2 1 dataset
ChIP Huh-7 GSE97411.RUVBL2.Huh-7 283 bp overlap
SIN3A 1 dataset
ChIP hiPSC_IID12 GSE106870.SIN3A.hiPSC_IID12 167 bp overlap
SMC1A 2 datasets
ChIP A-549 GSE76893.SMC1A.A-549 134 bp overlap
ChIP Hep-G2 GSE76893.SMC1A.Hep-G2 144 bp overlap
SNAI2 1 dataset
ChIP SMS-CTR_D48 GSE137168.SNAI2.SMS-CTR_D48 138 bp overlap
SOX2 1 dataset
ChIP hiPSC GSE56567.SOX2.hiPSC 273 bp overlap
SP4 1 dataset
ChIP WA01 ENCSR000BQV.SP4.WA01 96 bp overlap
T 1 dataset
ChIP H9_MESODERM GSE60606.T.H9_MESODERM 299 bp overlap
TCF3 1 dataset
ChIP RCH-ACV GSE85988.TCF3.RCH-ACV 329 bp overlap
TEAD4 1 dataset
ChIP SNU-216 GSE44416.TEAD4.SNU-216 174 bp overlap
TFAP2A 2 datasets
ChIP WA09 GSE105081.TFAP2A.WA09 169 bp overlap
ChIP WA09 GSE105081.TFAP2A.WA09 195 bp overlap
TFAP2C 2 datasets
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 612 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 612 bp overlap
THAP1 2 datasets
Motif DE_12h DE_12h-THAP1_MA0597.3 8 bp overlap
Motif ES_0h ES_0h-THAP1_MA0597.3 8 bp overlap
YY1 4 datasets
ChIP Huh-7 GSE97411.YY1.Huh-7 372 bp overlap
ChIP NT2-D1 ENCSR000EXG.YY1.NT2-D1 133 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 237 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 225 bp overlap
ZNF143 1 dataset
Motif DE_12h DE_12h-ZNF143_MA0088.2 16 bp overlap
ZNF148 1 dataset
ChIP K-562 GSE121133.ZNF148.K-562 197 bp overlap
ZNF528 2 datasets
ChIP HEK293 GSE76494.ZNF528.HEK293 133 bp overlap
ChIP HEK293T GSE78099.ZNF528.HEK293T 76 bp overlap
ZNF574 1 dataset
ChIP HepG2 ENCFF206MMY 338 bp overlap
ZNF76 2 datasets
Motif DE_12h DE_12h-ZNF76_MA1716.2 17 bp overlap
Motif ES_0h ES_0h-ZNF76_MA1716.2 17 bp overlap
ZNF90 1 dataset
ChIP HEK293T GSE78099.ZNF90.HEK293T 255 bp overlap
ZSCAN16 2 datasets
Motif DE_12h DE_12h-ZSCAN16_MA2100.1 18 bp overlap
Motif ES_0h ES_0h-ZSCAN16_MA2100.1 18 bp overlap