chr4 : 92,438,845 92,439,507
662 bp 90 TFs 2 linked genes
This 662 bp open chromatin element is linked to GRID2 and LNCPRESS2 and is bound by 90 transcription factors.
Linked Genes
2 genes
Link type
Gene Expression Dist. to TSS Distance Link type
GRID2 135.2 kb Distal Multiome
LNCPRESS2 162.1 kb Distal Multiome+HiCAR
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:92,433,845 – 92,444,507
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
90 transcription factors
Source
Cell type
AFF4 1 dataset
ChIP WTC11 ENCFF556XTF 445 bp overlap
ASCL1 2 datasets
Motif DE_12h DE_12h-ASCL1_MA1631.2 9 bp overlap
Motif ES_0h ES_0h-ASCL1_MA1631.2 9 bp overlap
ASH2L 2 datasets
ChIP H1 ENCFF399KAM 500 bp overlap
ChIP WA01 ENCSR850KIP.ASH2L.WA01 296 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 106 bp overlap
Atoh1 1 dataset
Motif DE_12h DE_12h-Atoh1_MA1467.3 7 bp overlap
CDX2 2 datasets
Motif DE_12h DE_12h-CDX2_MA0465.3 8 bp overlap
Motif ES_0h ES_0h-CDX2_MA0465.3 8 bp overlap
CEBPD 1 dataset
Motif DE_12h DE_12h-CEBPD_MA0836.3 8 bp overlap
CTCF 3 datasets
Motif DE_12h DE_12h-CTCF_MA0139.2 15 bp overlap
Motif DE_12h DE_12h-CTCF_MA1930.2 33 bp overlap
Motif ES_0h ES_0h-CTCF_MA0139.2 15 bp overlap
CTCFL 2 datasets
Motif DE_12h DE_12h-CTCFL_MA1102.3 8 bp overlap
Motif ES_0h ES_0h-CTCFL_MA1102.3 8 bp overlap
E2F8 1 dataset
Motif DE_12h DE_12h-E2F8_MA0865.3 9 bp overlap
Ebf4 2 datasets
Motif DE_12h DE_12h-Ebf4_MA2122.1 11 bp overlap
Motif ES_0h ES_0h-Ebf4_MA2122.1 11 bp overlap
FERD3L 1 dataset
Motif DE_12h DE_12h-FERD3L_MA1485.1 14 bp overlap
FIGLA 4 datasets
Motif DE_12h DE_12h-FIGLA_MA0820.2 6 bp overlap
Motif DE_12h DE_12h-FIGLA_MA0820.2 6 bp overlap
Motif ES_0h ES_0h-FIGLA_MA0820.2 6 bp overlap
Motif ES_0h ES_0h-FIGLA_MA0820.2 6 bp overlap
FOXD2 2 datasets
Motif DE_12h DE_12h-FOXD2_MA0847.4 11 bp overlap
Motif ES_0h ES_0h-FOXD2_MA0847.4 11 bp overlap
FOXF2 2 datasets
Motif DE_12h DE_12h-FOXF2_MA0030.2 9 bp overlap
Motif ES_0h ES_0h-FOXF2_MA0030.2 9 bp overlap
FOXH1 2 datasets
Motif DE_12h DE_12h-FOXH1_MA0479.2 8 bp overlap
Motif ES_0h ES_0h-FOXH1_MA0479.2 8 bp overlap
FOXK1 3 datasets
Motif DE_12h DE_12h-FOXK1_MA0852.3 7 bp overlap
Motif ES_0h ES_0h-FOXK1_MA0852.3 7 bp overlap
ChIP WTC11 ENCFF875IGU 390 bp overlap
FOXK2 2 datasets
Motif DE_12h DE_12h-FOXK2_MA1103.3 7 bp overlap
Motif ES_0h ES_0h-FOXK2_MA1103.3 7 bp overlap
FOXL1 2 datasets
Motif DE_12h DE_12h-FOXL1_MA0033.2 7 bp overlap
Motif ES_0h ES_0h-FOXL1_MA0033.2 7 bp overlap
FOXO4 2 datasets
Motif DE_12h DE_12h-FOXO4_MA0848.1 7 bp overlap
Motif ES_0h ES_0h-FOXO4_MA0848.1 7 bp overlap
FOXO6 2 datasets
Motif DE_12h DE_12h-FOXO6_MA0849.1 7 bp overlap
Motif ES_0h ES_0h-FOXO6_MA0849.1 7 bp overlap
FOXP2 2 datasets
Motif DE_12h DE_12h-FOXP2_MA0593.2 9 bp overlap
Motif ES_0h ES_0h-FOXP2_MA0593.2 9 bp overlap
FOXP3 2 datasets
Motif DE_12h DE_12h-FOXP3_MA0850.1 7 bp overlap
Motif ES_0h ES_0h-FOXP3_MA0850.1 7 bp overlap
Foxf1 2 datasets
Motif DE_12h DE_12h-Foxf1_MA1606.2 7 bp overlap
Motif ES_0h ES_0h-Foxf1_MA1606.2 7 bp overlap
Foxo1 2 datasets
Motif DE_12h DE_12h-Foxo1_MA0480.3 7 bp overlap
Motif ES_0h ES_0h-Foxo1_MA0480.3 7 bp overlap
Foxo3 2 datasets
Motif DE_12h DE_12h-Foxo3_MA0157.4 7 bp overlap
Motif ES_0h ES_0h-Foxo3_MA0157.4 7 bp overlap
Foxq1 2 datasets
Motif DE_12h DE_12h-Foxq1_MA0040.2 10 bp overlap
Motif ES_0h ES_0h-Foxq1_MA0040.2 10 bp overlap
GATA2 1 dataset
Motif DE_12h DE_12h-GATA2_MA0036.4 7 bp overlap
GATA4 1 dataset
Motif DE_12h DE_12h-GATA4_MA0482.3 8 bp overlap
GATA5 1 dataset
Motif DE_12h DE_12h-GATA5_MA0766.3 8 bp overlap
GATA6 6 datasets
Motif DE_12h DE_12h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 414 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 541 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 527 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 256 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 285 bp overlap
GLI3 1 dataset
Motif DE_12h DE_12h-GLI3_MA1491.3 15 bp overlap
GRHL1 1 dataset
Motif DE_12h DE_12h-GRHL1_MA0647.2 10 bp overlap
GRHL2 1 dataset
Motif DE_12h DE_12h-GRHL2_MA1105.3 8 bp overlap
Gata3 1 dataset
Motif DE_12h DE_12h-Gata3_MA0037.5 8 bp overlap
Gli1 1 dataset
Motif DE_12h DE_12h-Gli1_MA1990.2 10 bp overlap
Gli2 1 dataset
Motif DE_12h DE_12h-Gli2_MA0734.4 9 bp overlap
HAND2 1 dataset
Motif DE_12h DE_12h-HAND2_MA1638.2 6 bp overlap
JUN 7 datasets
ChIP DE_D1 S08-DE-d1-JUN-exp1 560 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 569 bp overlap
ChIP DE_D2 S02-DE-d2-JUN-exp1 568 bp overlap
ChIP DE_D2 S54-DE-d2-JUN-exp2 371 bp overlap
ChIP ESC S24-ESC-d0-JUN-exp1 395 bp overlap
ChIP ESC S34-ESC-d0-JUN-exp2 504 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 233 bp overlap
KLF1 2 datasets
Motif DE_12h DE_12h-KLF1_MA0493.3 8 bp overlap
Motif ES_0h ES_0h-KLF1_MA0493.3 8 bp overlap
KLF13 5 datasets
Motif DE_12h DE_12h-KLF13_MA0657.2 17 bp overlap
Motif DE_12h DE_12h-KLF13_MA0657.2 17 bp overlap
Motif DE_24h DE_24h-KLF13_MA0657.2 17 bp overlap
Motif ES_0h ES_0h-KLF13_MA0657.2 17 bp overlap
Motif ES_0h ES_0h-KLF13_MA0657.2 17 bp overlap
KLF16 2 datasets
Motif DE_12h DE_12h-KLF16_MA0741.1 11 bp overlap
Motif ES_0h ES_0h-KLF16_MA0741.1 11 bp overlap
KLF2 2 datasets
Motif DE_12h DE_12h-KLF2_MA1515.2 8 bp overlap
Motif ES_0h ES_0h-KLF2_MA1515.2 8 bp overlap
KLF4 2 datasets
Motif DE_12h DE_12h-KLF4_MA0039.5 8 bp overlap
Motif ES_0h ES_0h-KLF4_MA0039.5 8 bp overlap
KLF5 2 datasets
Motif DE_12h DE_12h-KLF5_MA0599.1 10 bp overlap
Motif ES_0h ES_0h-KLF5_MA0599.1 10 bp overlap
MAFK 2 datasets
Motif DE_12h DE_12h-MAFK_MA0496.4 10 bp overlap
Motif ES_0h ES_0h-MAFK_MA0496.4 10 bp overlap
NANOG 5 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 318 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 264 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 230 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 215 bp overlap
Nfat5 2 datasets
Motif DE_12h DE_12h-Nfat5_MA0606.3 8 bp overlap
Motif ES_0h ES_0h-Nfat5_MA0606.3 8 bp overlap
Nfatc2 2 datasets
Motif DE_12h DE_12h-Nfatc2_MA0152.3 8 bp overlap
Motif ES_0h ES_0h-Nfatc2_MA0152.3 8 bp overlap
Nr1h3::Rxra 3 datasets
Motif DE_12h DE_12h-Nr1h3Rxra_MA0494.2 16 bp overlap
Motif DE_12h DE_12h-Nr1h3Rxra_MA0494.2 16 bp overlap
Motif ES_0h ES_0h-Nr1h3Rxra_MA0494.2 16 bp overlap
POLR2A 2 datasets
ChIP GM23338 ENCFF450WCS 378 bp overlap
ChIP GM23338 ENCFF450WCS 420 bp overlap
POU5F1 3 datasets
ChIP DE_D1 DED1-OCT4_Batch_II 472 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 290 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 154 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 345 bp overlap
PPARG 2 datasets
Motif DE_12h DE_12h-PPARG_MA0066.2 19 bp overlap
Motif ES_0h ES_0h-PPARG_MA0066.2 19 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 270 bp overlap
SMAD2_3 4 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 455 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 511 bp overlap
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 447 bp overlap
ChIP ESC S33-ESC-d0-SMAD2_3-exp2 359 bp overlap
SNAI1 2 datasets
Motif DE_12h DE_12h-SNAI1_MA1558.2 7 bp overlap
Motif ES_0h ES_0h-SNAI1_MA1558.2 7 bp overlap
SNAI3 2 datasets
Motif DE_12h DE_12h-SNAI3_MA1559.2 9 bp overlap
Motif ES_0h ES_0h-SNAI3_MA1559.2 9 bp overlap
SOX2 1 dataset
ChIP hiPSC GSE56567.SOX2.hiPSC 151 bp overlap
SP8 5 datasets
Motif DE_12h DE_12h-SP8_MA0747.2 11 bp overlap
Motif DE_12h DE_12h-SP8_MA0747.2 11 bp overlap
Motif DE_24h DE_24h-SP8_MA0747.2 11 bp overlap
Motif ES_0h ES_0h-SP8_MA0747.2 11 bp overlap
Motif ES_0h ES_0h-SP8_MA0747.2 11 bp overlap
SPIB 1 dataset
Motif DE_12h DE_12h-SPIB_MA0081.3 13 bp overlap
SREBF1 2 datasets
Motif DE_12h DE_12h-SREBF1_MA0595.1 10 bp overlap
Motif ES_0h ES_0h-SREBF1_MA0595.1 10 bp overlap
TCF12 2 datasets
Motif DE_12h DE_12h-TCF12_MA1648.2 7 bp overlap
Motif ES_0h ES_0h-TCF12_MA1648.2 7 bp overlap
TCF3 2 datasets
Motif DE_12h DE_12h-TCF3_MA0522.4 7 bp overlap
Motif ES_0h ES_0h-TCF3_MA0522.4 7 bp overlap
TCF4 2 datasets
Motif DE_12h DE_12h-TCF4_MA0830.3 8 bp overlap
Motif ES_0h ES_0h-TCF4_MA0830.3 8 bp overlap
TFAP2C 2 datasets
ChIP UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_1d_TFAP2C_Induc 163 bp overlap
ChIP UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc GSE101074.TFAP2C.UCLA1-hESCs_TFAP2C--_2d_TFAP2C_Induc 86 bp overlap
TRPS1 1 dataset
Motif DE_12h DE_12h-TRPS1_MA1970.2 8 bp overlap
TWIST1 1 dataset
Motif DE_12h DE_12h-TWIST1_MA1123.3 8 bp overlap
USF2 1 dataset
ChIP WTC11 ENCFF139JAW 417 bp overlap
Yy1 1 dataset
Motif DE_12h DE_12h-Yy1_MA0095.4 8 bp overlap
ZBTB7B 1 dataset
Motif DE_12h DE_12h-ZBTB7B_MA0694.2 10 bp overlap
ZBTB7C 1 dataset
Motif DE_12h DE_12h-ZBTB7C_MA0695.2 8 bp overlap
ZEB1 4 datasets
Motif DE_12h DE_12h-ZEB1_MA0103.4 6 bp overlap
Motif DE_12h DE_12h-ZEB1_MA0103.4 6 bp overlap
Motif ES_0h ES_0h-ZEB1_MA0103.4 6 bp overlap
Motif ES_0h ES_0h-ZEB1_MA0103.4 6 bp overlap
ZFP42 1 dataset
Motif DE_12h DE_12h-ZFP42_MA1651.2 13 bp overlap
ZIC1 3 datasets
Motif DE_12h DE_12h-ZIC1_MA0696.1 14 bp overlap
Motif DE_24h DE_24h-ZIC1_MA0696.1 14 bp overlap
Motif ES_0h ES_0h-ZIC1_MA0696.1 14 bp overlap
ZIC4 3 datasets
Motif DE_12h DE_12h-ZIC4_MA0751.2 14 bp overlap
Motif DE_24h DE_24h-ZIC4_MA0751.2 14 bp overlap
Motif ES_0h ES_0h-ZIC4_MA0751.2 14 bp overlap
ZIC5 3 datasets
Motif DE_12h DE_12h-ZIC5_MA1584.2 15 bp overlap
Motif DE_24h DE_24h-ZIC5_MA1584.2 15 bp overlap
Motif ES_0h ES_0h-ZIC5_MA1584.2 15 bp overlap
ZNF157 2 datasets
Motif DE_12h DE_12h-ZNF157_MA2331.1 21 bp overlap
Motif ES_0h ES_0h-ZNF157_MA2331.1 21 bp overlap
ZNF16 2 datasets
Motif DE_12h DE_12h-ZNF16_MA1654.2 21 bp overlap
Motif ES_0h ES_0h-ZNF16_MA1654.2 21 bp overlap
ZNF184 1 dataset
ChIP WTC11 ENCFF352POG 425 bp overlap
ZNF341 2 datasets
Motif DE_12h DE_12h-ZNF341_MA1655.2 8 bp overlap
Motif ES_0h ES_0h-ZNF341_MA1655.2 8 bp overlap
ZNF416 1 dataset
Motif DE_12h DE_12h-ZNF416_MA1979.2 10 bp overlap
ZNF582 1 dataset
Motif DE_12h DE_12h-ZNF582_MA1983.2 19 bp overlap
ZNF682 2 datasets
Motif DE_12h DE_12h-ZNF682_MA1599.2 11 bp overlap
Motif ES_0h ES_0h-ZNF682_MA1599.2 11 bp overlap
ZNF707 1 dataset
Motif DE_12h DE_12h-ZNF707_MA1715.1 15 bp overlap
ZNF784 2 datasets
Motif DE_12h DE_12h-ZNF784_MA1717.2 8 bp overlap
Motif ES_0h ES_0h-ZNF784_MA1717.2 8 bp overlap