chr4 : 48,071,082 48,071,611
529 bp 60 TFs 1 linked gene
This 529 bp open chromatin element is linked to TXK and is bound by 60 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
TXK 8.6 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:48,066,082 – 48,076,611
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
60 transcription factors
Source
Cell type
ATF3 1 dataset
ChIP HCT-116_CAMP GSE74355.ATF3.HCT-116_CAMP 205 bp overlap
BCOR 1 dataset
ChIP B-cell_GERMINAL_CENTER GSE43350.BCOR.B-cell_GERMINAL_CENTER 170 bp overlap
BRD2 2 datasets
ChIP HCC1806_BAZ2BsiRNA GSE116879.BRD2.HCC1806_BAZ2BsiRNA 214 bp overlap
ChIP HCC1806_NTsiRNA GSE116879.BRD2.HCC1806_NTsiRNA 258 bp overlap
BRD4 6 datasets
ChIP HCC1806_100nMtrametinib_24h GSE87418.BRD4.HCC1806_100nMtrametinib_24h 457 bp overlap
ChIP HCC1806_300nMJQ1_24h GSE87418.BRD4.HCC1806_300nMJQ1_24h 303 bp overlap
ChIP HCC1806_DMSO_24h GSE87418.BRD4.HCC1806_DMSO_24h 220 bp overlap
ChIP HT29 GSE73319.BRD4.HT29 327 bp overlap
ChIP PC-3_GDC-R GSE137207.BRD4.PC-3_GDC-R 246 bp overlap
ChIP PC-3_GDC-R_SAHA GSE137207.BRD4.PC-3_GDC-R_SAHA 284 bp overlap
CBFB 1 dataset
ChIP GM12878 ENCSR860UHK.CBFB.GM12878 189 bp overlap
CDX2 3 datasets
ChIP Caco-2_DIFF GSE23436.CDX2.Caco-2_DIFF 329 bp overlap
ChIP Caco-2_PROLIF GSE23436.CDX2.Caco-2_PROLIF 268 bp overlap
ChIP adult-duodenal-cell GSE115314.CDX2.adult-duodenal-cell 219 bp overlap
CEBPB 1 dataset
ChIP HeLa-S3 ENCFF722WEG 265 bp overlap
CHD4 2 datasets
ChIP 501-mel GSE134848.CHD4.501-mel 148 bp overlap
ChIP HaCaT GSE139685.CHD4.HaCaT 256 bp overlap
CREB1 2 datasets
ChIP GM12878 ENCFF870CVH 307 bp overlap
ChIP GM12878 ENCSR000BUF.CREB1.GM12878 238 bp overlap
CREBBP 1 dataset
ChIP PC-3 GSE147455.CREBBP.PC-3 127 bp overlap
CREM 2 datasets
ChIP GM12878 ENCFF391UGE 254 bp overlap
ChIP GM12878 ENCSR839XZU.CREM.GM12878 176 bp overlap
CTCF 81 datasets
ChIP 22Rv1_hydroxy_10nM_4h ENCSR847XGE.CTCF.22Rv1_hydroxy_10nM_4h 184 bp overlap
ChIP 786-O_NORMOXIA GSE78113.CTCF.786-O_NORMOXIA 200 bp overlap
ChIP A-549 ENCSR000AUF.CTCF.A-549 166 bp overlap
ChIP A-673 ENCSR611JJS.CTCF.A-673 278 bp overlap
ChIP Caco-2 ENCFF934QYS 137 bp overlap
ChIP Caco-2 ENCSR000DQN.CTCF.Caco-2 113 bp overlap
ChIP FT282 GSE131931.CTCF.FT282 87 bp overlap
ChIP FT282_EV GSE131931.CTCF.FT282_EV 205 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 194 bp overlap
ChIP GM12878 ENCFF511URZ 165 bp overlap
ChIP GM23338 ENCFF531QOI 279 bp overlap
ChIP GM23338 ENCFF772DML 153 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 185 bp overlap
ChIP HAP1_2kb GSE152721.CTCF.HAP1_2kb 290 bp overlap
ChIP HAP1_clone21 GSE152721.CTCF.HAP1_clone21 191 bp overlap
ChIP HAP1_clone21_CTCF-CRISPR GSE152721.CTCF.HAP1_clone21_CTCF-CRISPR 162 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 172 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_2 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_2 212 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 213 bp overlap
ChIP HAP1_clone25 GSE152721.CTCF.HAP1_clone25 170 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 182 bp overlap
ChIP HeLa GSE126990.CTCF.HeLa 178 bp overlap
ChIP HeLa-Kyoto GSE138405.CTCF.HeLa-Kyoto 178 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 175 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 155 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 189 bp overlap
ChIP HeLa-S3 ENCFF565UFR 146 bp overlap
ChIP HeLa-S3 ENCSR000DLO.CTCF.HeLa-S3 117 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 147 bp overlap
ChIP HeLa-S3_unsynchro GSE108173.CTCF.HeLa-S3_unsynchro 171 bp overlap
ChIP Hep-G2 ENCSR000AMA.CTCF.Hep-G2 149 bp overlap
ChIP K-562 GSE70764.CTCF.K-562 160 bp overlap
ChIP K-562 ENCSR000EGM.CTCF.K-562 92 bp overlap
ChIP K-562_8005 GSE131349.CTCF.K-562_8005 60 bp overlap
ChIP K-562_MonoC GSE137216.CTCF.K-562_MonoC 129 bp overlap
ChIP Loucy ENCFF359TVQ 265 bp overlap
ChIP Loucy ENCSR464DKE.CTCF.Loucy 212 bp overlap
ChIP MCF-7 ENCFF198DQX 173 bp overlap
ChIP MCF-7 ENCFF494VXA 173 bp overlap
ChIP MCF-7 GSE137216.CTCF.MCF-7 187 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 150 bp overlap
ChIP MCF-7 ENCSR000DMS.CTCF.MCF-7 139 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 125 bp overlap
ChIP MCF-7L_t0 GSE108787.CTCF.MCF-7L_t0 175 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 113 bp overlap
ChIP OCI-LY3 ENCFF939BYJ 150 bp overlap
ChIP OCI-LY7 ENCFF086AXQ 302 bp overlap
ChIP OCI-Ly1 ENCSR072EUE.CTCF.OCI-Ly1 152 bp overlap
ChIP OCI-Ly3 ENCSR756ZKG.CTCF.OCI-Ly3 237 bp overlap
ChIP OCI-Ly7 ENCSR027HML.CTCF.OCI-Ly7 187 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 164 bp overlap
ChIP PC-3 ENCFF487TUI 277 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 228 bp overlap
ChIP RWPE-2 ENCSR856JJB.CTCF.RWPE-2 161 bp overlap
ChIP RWPE2 ENCFF911IEE 406 bp overlap
ChIP SK-N-AS GSE101295.CTCF.SK-N-AS 106 bp overlap
ChIP SK-N-SH ENCFF731NJX 221 bp overlap
ChIP SK-N-SH ENCSR000DXQ.CTCF.SK-N-SH 93 bp overlap
ChIP THP-1_Pam3csk-045m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-045m-Flavo-000m 111 bp overlap
ChIP THP-1_Pam3csk-060m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-060m-Flavo-000m 129 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.CTCF.THP-1_Pam3csk-150m-Flavo-030m 126 bp overlap
ChIP brain microvascular endothelial cell ENCFF526VHE 202 bp overlap
ChIP choroid plexus epithelial cell ENCFF407YNR 152 bp overlap
ChIP choroid-plexus_epithelial ENCSR000DTL.CTCF.choroid-plexus_epithelial 90 bp overlap
ChIP endothelial_brain-microvascular ENCSR000DTA.CTCF.endothelial_brain-microvascular 144 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 355 bp overlap
ChIP epithelial_kidney ENCSR000DVH.CTCF.epithelial_kidney 121 bp overlap
ChIP epithelial_retinal_pigment ENCSR000DVI.CTCF.epithelial_retinal_pigment 110 bp overlap
ChIP erythroblast_adult_erythroblasts GSE102184.CTCF.erythroblast_adult_erythroblasts 166 bp overlap
ChIP fetal_erythroblasts GSE102184.CTCF.fetal_erythroblasts 131 bp overlap
ChIP fibroblast_PULMONARY_ARTERY ENCSR000DUX.CTCF.fibroblast_PULMONARY_ARTERY 135 bp overlap
ChIP foreskin keratinocyte ENCFF980OWR 247 bp overlap
ChIP hepatocyte ENCSR252QYR.CTCF.hepatocyte 105 bp overlap
ChIP keratinocyte ENCFF046PBT 160 bp overlap
ChIP keratinocyte ENCFF291YDC 160 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 143 bp overlap
ChIP keratinocyte ENCSR000ALJ.CTCF.keratinocyte 109 bp overlap
ChIP leukemia_CEBPA-24h GSE142161.CTCF.leukemia_CEBPA-24h 115 bp overlap
ChIP primary-glioblastoma_G583 GSE121601.CTCF.primary-glioblastoma_G583 171 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 275 bp overlap
ChIP retinal pigment epithelial cell ENCFF810AAG 180 bp overlap
CTCFL 1 dataset
ChIP FT282 GSE131931.CTCFL.FT282 408 bp overlap
DPF2 1 dataset
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.DPF2.BIN-67_lentivirus-SMARCA4-T910M 190 bp overlap
ELF3 3 datasets
ChIP PDAC GSE64557.ELF3.PDAC 319 bp overlap
ChIP PDAC_KOKLF5 GSE64557.ELF3.PDAC_KOKLF5 201 bp overlap
ChIP PDAC_SHCTR GSE64557.ELF3.PDAC_SHCTR 193 bp overlap
EP300 1 dataset
ChIP HeLa-S3 ENCSR000ECV.EP300.HeLa-S3 133 bp overlap
ESR1 7 datasets
ChIP MCF-7 GSE119057.ESR1.MCF-7 136 bp overlap
ChIP MCF-7_ER_t10 GSE119057.ESR1.MCF-7_ER_t10 156 bp overlap
ChIP MCF-7_ER_t20 GSE119057.ESR1.MCF-7_ER_t20 154 bp overlap
ChIP MCF-7_ER_t30 GSE119057.ESR1.MCF-7_ER_t30 142 bp overlap
ChIP MCF-7_ER_t50 GSE119057.ESR1.MCF-7_ER_t50 155 bp overlap
ChIP MCF-7_ER_t60 GSE119057.ESR1.MCF-7_ER_t60 144 bp overlap
ChIP MCF-7_ER_t70 GSE119057.ESR1.MCF-7_ER_t70 144 bp overlap
FOS 2 datasets
ChIP CFPAC-1 GSE119930.FOS.CFPAC-1 424 bp overlap
ChIP HeLa-S3 ENCSR000EZE.FOS.HeLa-S3 157 bp overlap
FOXA1 3 datasets
ChIP CFPAC-1 GSE119930.FOXA1.CFPAC-1 348 bp overlap
ChIP PDAC GSE64557.FOXA1.PDAC 397 bp overlap
ChIP PDAC_SHCTR GSE64557.FOXA1.PDAC_SHCTR 506 bp overlap
FOXA2 6 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.FOXA2.BJ1-hTERT_FOXA2_GATA4_Coexp 212 bp overlap
ChIP CFPAC-1_EV GSE119930.FOXA2.CFPAC-1_EV 353 bp overlap
ChIP CFPAC-1_HOXB8-OE GSE119930.FOXA2.CFPAC-1_HOXB8-OE 305 bp overlap
ChIP Caco-2 GSE66218.FOXA2.Caco-2 182 bp overlap
ChIP DE DE-FOXA2-1 295 bp overlap
ChIP DE DE-FOXA2-2 325 bp overlap
GATA4 6 datasets
ChIP A-549 GSE85002.GATA4.A-549 191 bp overlap
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.GATA4.BJ1-hTERT_FOXA2_GATA4_Coexp 396 bp overlap
ChIP DE DE-GATA4-1 406 bp overlap
ChIP DE DE-GATA4-2 482 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 518 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 459 bp overlap
GATA6 11 datasets
ChIP AGS GSE51705.GATA6.AGS 245 bp overlap
ChIP AGS GSE51936.GATA6.AGS 80 bp overlap
ChIP Caco-2_PROLIF GSE23436.GATA6.Caco-2_PROLIF 272 bp overlap
ChIP DE DE-GATA6-1 438 bp overlap
ChIP DE DE-GATA6-2 515 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 349 bp overlap
ChIP ESO-26 GSE132680.GATA6.ESO-26 528 bp overlap
ChIP OACP4-C GSE132680.GATA6.OACP4-C 529 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 447 bp overlap
ChIP foregut GSE117136.GATA6.foregut 294 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 403 bp overlap
HNF1B 1 dataset
ChIP PDAC GSE64557.HNF1B.PDAC 310 bp overlap
HNF4A 2 datasets
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 203 bp overlap
ChIP Caco-2_PROLIF GSE23436.HNF4A.Caco-2_PROLIF 131 bp overlap
HOXB8 1 dataset
ChIP CFPAC-1_HOXB8-OE GSE119930.HOXB8.CFPAC-1_HOXB8-OE 464 bp overlap
IRF1 1 dataset
ChIP PDAC GSE64557.IRF1.PDAC 529 bp overlap
ISL1 1 dataset
ChIP SK-N-SH ENCFF285GEQ 395 bp overlap
JUN 3 datasets
ChIP DE_D2 S02-DE-d2-JUN-exp1 332 bp overlap
ChIP HAEC_TNFa_4h GSE89970.JUN.HAEC_TNFa_4h 366 bp overlap
ChIP HeLa-S3 ENCSR000EDG.JUN.HeLa-S3 177 bp overlap
JUNB 1 dataset
ChIP GM12878 ENCSR897MMC.JUNB.GM12878 399 bp overlap
JUND 1 dataset
ChIP HeLa-S3 ENCSR000EDH.JUND.HeLa-S3 242 bp overlap
KLF4 1 dataset
ChIP PDAC GSE64557.KLF4.PDAC 336 bp overlap
KLF6 1 dataset
ChIP PDAC GSE64557.KLF6.PDAC 339 bp overlap
KMT2D 1 dataset
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.KMT2D.BIN-67_lentivirus-SMARCA4 334 bp overlap
LHX2 1 dataset
ChIP retina_pigment GSE60024.LHX2.retina_pigment 184 bp overlap
MAML1 1 dataset
ChIP SCC_4h GSE156486.MAML1.SCC_4h 428 bp overlap
NR1H2 1 dataset
ChIP HT29_GW3965_48H GSE77039.NR1H2.HT29_GW3965_48H 196 bp overlap
NR3C1 1 dataset
ChIP HeLa-B2_TA GSE24518.NR3C1.HeLa-B2_TA 138 bp overlap
POLR2A 1 dataset
ChIP stomach ENCFF820WZN 345 bp overlap
RAD21 4 datasets
ChIP GM12878 ENCSR000BMY.RAD21.GM12878 104 bp overlap
ChIP HAP1 GSE152721.RAD21.HAP1 226 bp overlap
ChIP K-562_RF2-insertion GSE140868.RAD21.K-562_RF2-insertion 74 bp overlap
ChIP MCF-7 ENCSR703TNG.RAD21.MCF-7 121 bp overlap
RBPJ 1 dataset
ChIP SCC_4h GSE156486.RBPJ.SCC_4h 332 bp overlap
RELA 15 datasets
ChIP Detroit-562_LPS GSE91018.RELA.Detroit-562_LPS 435 bp overlap
ChIP Detroit-562_TNFa GSE91018.RELA.Detroit-562_TNFa 424 bp overlap
ChIP aortic-endothelial-cell_IL1B_D10 GSE139377.RELA.aortic-endothelial-cell_IL1B_D10 392 bp overlap
ChIP aortic-endothelial-cell_IL1B_D11 GSE139377.RELA.aortic-endothelial-cell_IL1B_D11 304 bp overlap
ChIP aortic-endothelial-cell_IL1B_D14 GSE139377.RELA.aortic-endothelial-cell_IL1B_D14 231 bp overlap
ChIP aortic-endothelial-cell_IL1B_D21 GSE139377.RELA.aortic-endothelial-cell_IL1B_D21 430 bp overlap
ChIP aortic-endothelial-cell_IL1B_D26 GSE139377.RELA.aortic-endothelial-cell_IL1B_D26 216 bp overlap
ChIP aortic-endothelial-cell_IL1B_D3 GSE139377.RELA.aortic-endothelial-cell_IL1B_D3 245 bp overlap
ChIP aortic-endothelial-cell_IL1B_D31 GSE139377.RELA.aortic-endothelial-cell_IL1B_D31 385 bp overlap
ChIP aortic-endothelial-cell_IL1B_D35 GSE139377.RELA.aortic-endothelial-cell_IL1B_D35 173 bp overlap
ChIP aortic-endothelial-cell_IL1B_D39 GSE139377.RELA.aortic-endothelial-cell_IL1B_D39 363 bp overlap
ChIP aortic-endothelial-cell_IL1B_D4 GSE139377.RELA.aortic-endothelial-cell_IL1B_D4 358 bp overlap
ChIP aortic-endothelial-cell_IL1B_D47 GSE139377.RELA.aortic-endothelial-cell_IL1B_D47 335 bp overlap
ChIP aortic-endothelial-cell_IL1B_D7 GSE139377.RELA.aortic-endothelial-cell_IL1B_D7 150 bp overlap
ChIP aortic-endothelial-cell_IL1B_D9 GSE139377.RELA.aortic-endothelial-cell_IL1B_D9 357 bp overlap
SMAD3 2 datasets
ChIP HCC1954 GSE104760.SMAD3.HCC1954 506 bp overlap
ChIP HCC1954_TGFb GSE104760.SMAD3.HCC1954_TGFb 517 bp overlap
SMAD4 1 dataset
ChIP Caco-2 GSE112946.SMAD4.Caco-2 334 bp overlap
SMARCA4 4 datasets
ChIP A-549_AG15677 GSE132290.SMARCA4.A-549_AG15677 73 bp overlap
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4 413 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SMARCA4.BIN-67_lentivirus-SMARCA4-T910M 480 bp overlap
ChIP HeLa-S3 ENCSR000EZC.SMARCA4.HeLa-S3 306 bp overlap
SMARCC1 3 datasets
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4 305 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SMARCC1.BIN-67_lentivirus-SMARCA4-T910M 332 bp overlap
ChIP TTC-549_Dox GSE71504.SMARCC1.TTC-549_Dox 228 bp overlap
SMC1A 1 dataset
ChIP MCF-7_EtOH GSE115602.SMC1A.MCF-7_EtOH 100 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 311 bp overlap
SOX2 2 datasets
ChIP HCC2814 GSE137459.SOX2.HCC2814 282 bp overlap
ChIP KNS-62 GSE137459.SOX2.KNS-62 296 bp overlap
SOX4 1 dataset
ChIP HCC1954 GSE104760.SOX4.HCC1954 257 bp overlap
SS18 2 datasets
ChIP BIN-67_lentivirus-SMARCA4 GSE117734.SS18.BIN-67_lentivirus-SMARCA4 449 bp overlap
ChIP BIN-67_lentivirus-SMARCA4-T910M GSE117734.SS18.BIN-67_lentivirus-SMARCA4-T910M 329 bp overlap
STAG1 1 dataset
ChIP MCF-7 ERP000209.STAG1.MCF-7 120 bp overlap
TCF21 1 dataset
ChIP HCASMC GSE124011.TCF21.HCASMC 141 bp overlap
TEAD1 1 dataset
ChIP MSTO GSE68170.TEAD1.MSTO 272 bp overlap
TFAP2B 1 dataset
ChIP SK-N-SH ENCFF869XXQ 255 bp overlap
TWIST1 2 datasets
ChIP SHEP-21N GSE80151.TWIST1.SHEP-21N 85 bp overlap
ChIP SHEP-21N_24h GSE80151.TWIST1.SHEP-21N_24h 85 bp overlap
ZBTB2 1 dataset
ChIP GM12878 GSE97661.ZBTB2.GM12878 110 bp overlap
ZEB1 1 dataset
ChIP PDAC GSE64557.ZEB1.PDAC 324 bp overlap
ZFP64 1 dataset
ChIP HEK293 GSE76494.ZFP64.HEK293 213 bp overlap
ZNF750 1 dataset
ChIP keratinocyte_diff GSE57702.ZNF750.keratinocyte_diff 152 bp overlap