chr10 : 9,339,575 – 9,340,197
622 bp 100 TFs 0 linked genes
This 622 bp open chromatin element has no linked target genes and is bound by 100 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr10:9,334,575 – 9,345,197
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
100 transcription factors
Source
Cell type
Arid3a 2 datasets
Motif DE_36h DE_36h-Arid3a_MA0151.1 6 bp overlap
Motif DE_60h DE_60h-Arid3a_MA0151.1 6 bp overlap
BARX1 2 datasets
Motif DE_36h DE_36h-BARX1_MA0875.2 6 bp overlap
Motif DE_60h DE_60h-BARX1_MA0875.2 6 bp overlap
BCL11A 1 dataset
ChIP CD34_Day3_30min GSE104676.BCL11A.CD34_Day3_30min 64 bp overlap
BRD4 1 dataset
ChIP Jurkat GSE83777.BRD4.Jurkat 307 bp overlap
BSX 2 datasets
Motif DE_36h DE_36h-BSX_MA0876.2 6 bp overlap
Motif DE_60h DE_60h-BSX_MA0876.2 6 bp overlap
CDK9-HEXIM1 1 dataset
ChIP A-375 GSE68047.CDK9-HEXIM1.A-375 144 bp overlap
CREBBP 1 dataset
ChIP monocyte_IFNg-LPS GSE131294.CREBBP.monocyte_IFNg-LPS 140 bp overlap
DLX1 2 datasets
Motif DE_36h DE_36h-DLX1_MA0879.3 6 bp overlap
Motif DE_60h DE_60h-DLX1_MA0879.3 6 bp overlap
DLX6 2 datasets
Motif DE_36h DE_36h-DLX6_MA0882.2 6 bp overlap
Motif DE_60h DE_60h-DLX6_MA0882.2 6 bp overlap
Dlx3 2 datasets
Motif DE_36h DE_36h-Dlx3_MA0880.2 6 bp overlap
Motif DE_60h DE_60h-Dlx3_MA0880.2 6 bp overlap
Dlx4 2 datasets
Motif DE_36h DE_36h-Dlx4_MA0881.2 6 bp overlap
Motif DE_60h DE_60h-Dlx4_MA0881.2 6 bp overlap
ELF1 2 datasets
Motif DE_36h DE_36h-ELF1_MA0473.4 9 bp overlap
Motif DE_60h DE_60h-ELF1_MA0473.4 9 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 290 bp overlap
EP300 1 dataset
ChIP AML GSE131939.EP300.AML 104 bp overlap
ERF::FOXI1 2 datasets
Motif DE_36h DE_36h-ERFFOXI1_MA1935.2 10 bp overlap
Motif DE_60h DE_60h-ERFFOXI1_MA1935.2 10 bp overlap
ERG 3 datasets
ChIP Jurkat GSE49091.ERG.Jurkat 137 bp overlap
ChIP ME-1 GSE46044.ERG.ME-1 270 bp overlap
ChIP TSU-1621MT_ATRA GSE60477.ERG.TSU-1621MT_ATRA 296 bp overlap
ETS1 4 datasets
ChIP CTV-1_DSG GSE128835.ETS1.CTV-1_DSG 194 bp overlap
ChIP CTV-1_DSG_SPI1-mut GSE128835.ETS1.CTV-1_DSG_SPI1-mut 263 bp overlap
Motif DE_36h DE_36h-ETS1_MA0098.4 9 bp overlap
Motif DE_60h DE_60h-ETS1_MA0098.4 9 bp overlap
ETS2 2 datasets
Motif DE_36h DE_36h-ETS2_MA1484.2 9 bp overlap
Motif DE_60h DE_60h-ETS2_MA1484.2 9 bp overlap
ETV1 2 datasets
Motif DE_36h DE_36h-ETV1_MA0761.3 9 bp overlap
Motif DE_60h DE_60h-ETV1_MA0761.3 9 bp overlap
ETV2::FOXI1 2 datasets
Motif DE_36h DE_36h-ETV2FOXI1_MA1942.2 11 bp overlap
Motif DE_60h DE_60h-ETV2FOXI1_MA1942.2 11 bp overlap
ETV5::FOXO1 2 datasets
Motif DE_36h DE_36h-ETV5FOXO1_MA1947.2 10 bp overlap
Motif DE_60h DE_60h-ETV5FOXO1_MA1947.2 10 bp overlap
EVI1 1 dataset
ChIP SKH1 GSE87283.EVI1.SKH1 311 bp overlap
EZH2 1 dataset
ChIP DND41 ENCSR000ASW.EZH2.DND41 239 bp overlap
Erg 2 datasets
Motif DE_36h DE_36h-Erg_MA0474.4 10 bp overlap
Motif DE_60h DE_60h-Erg_MA0474.4 10 bp overlap
FEZF2 3 datasets
Motif DE_36h DE_36h-FEZF2_MA2341.1 8 bp overlap
Motif DE_48h DE_48h-FEZF2_MA2341.1 8 bp overlap
Motif DE_60h DE_60h-FEZF2_MA2341.1 8 bp overlap
FLI1::FOXI1 2 datasets
Motif DE_36h DE_36h-FLI1FOXI1_MA1950.2 11 bp overlap
Motif DE_60h DE_60h-FLI1FOXI1_MA1950.2 11 bp overlap
FOXA2 3 datasets
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.FOXA2.BJ1-hTERT_FOXA2_GATA4_Coexp 155 bp overlap
ChIP DE DE-FOXA2-1 504 bp overlap
ChIP DE DE-FOXA2-2 482 bp overlap
FOXD3 3 datasets
Motif DE_36h DE_36h-FOXD3_MA0041.3 14 bp overlap
Motif DE_48h DE_48h-FOXD3_MA0041.3 14 bp overlap
Motif DE_60h DE_60h-FOXD3_MA0041.3 14 bp overlap
FOXJ2::ELF1 2 datasets
Motif DE_36h DE_36h-FOXJ2ELF1_MA1952.2 11 bp overlap
Motif DE_60h DE_60h-FOXJ2ELF1_MA1952.2 11 bp overlap
FOXO1::ELF1 2 datasets
Motif DE_36h DE_36h-FOXO1ELF1_MA1953.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1ELF1_MA1953.2 13 bp overlap
FOXO1::ELK1 2 datasets
Motif DE_36h DE_36h-FOXO1ELK1_MA1954.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1ELK1_MA1954.2 13 bp overlap
FOXO1::ELK3 2 datasets
Motif DE_36h DE_36h-FOXO1ELK3_MA1955.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1ELK3_MA1955.2 13 bp overlap
FOXO1::FLI1 2 datasets
Motif DE_36h DE_36h-FOXO1FLI1_MA1956.2 13 bp overlap
Motif DE_60h DE_60h-FOXO1FLI1_MA1956.2 13 bp overlap
GABPA 2 datasets
Motif DE_36h DE_36h-GABPA_MA0062.4 10 bp overlap
Motif DE_60h DE_60h-GABPA_MA0062.4 10 bp overlap
GATA1 10 datasets
ChIP CD34_Day7_30min GSE104676.GATA1.CD34_Day7_30min 55 bp overlap
ChIP CD34_ERYTH_BIO GSE29194.GATA1.CD34_ERYTH_BIO 254 bp overlap
ChIP CD34_ERYTH_BMP GSE29194.GATA1.CD34_ERYTH_BMP 147 bp overlap
ChIP HUDEP-2_30min GSE104676.GATA1.HUDEP-2_30min 128 bp overlap
ChIP erythroblast ENCFF867JAR 558 bp overlap
ChIP erythroblast ENCSR000EXP.GATA1.erythroblast 442 bp overlap
ChIP erythroid-progenitor GSE124163.GATA1.erythroid-progenitor 178 bp overlap
ChIP erythroid_Don001 GSE137982.GATA1.erythroid_Don001 446 bp overlap
ChIP erythroid_Don002 GSE137982.GATA1.erythroid_Don002 240 bp overlap
ChIP erythroid_Don003 GSE137982.GATA1.erythroid_Don003 306 bp overlap
GATA2 12 datasets
ChIP CD34_ACY957 GSE60792.GATA2.CD34_ACY957 415 bp overlap
ChIP CD34_DMSO GSE60792.GATA2.CD34_DMSO 229 bp overlap
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
Motif DE_48h DE_48h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
Motif DE_60h DE_60h-GATA2_MA0036.4 7 bp overlap
ChIP ME-1 GSE46044.GATA2.ME-1 319 bp overlap
ChIP SKH1 GSE87283.GATA2.SKH1 376 bp overlap
ChIP SKH1_RUNX1-EVI1_KD GSE87283.GATA2.SKH1_RUNX1-EVI1_KD 193 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 535 bp overlap
ChIP TSU-1621MT GSE60477.GATA2.TSU-1621MT 194 bp overlap
GATA3 2 datasets
ChIP Jurkat GSE76181.GATA3.Jurkat 293 bp overlap
ChIP Jurkat GSE120063.GATA3.Jurkat 266 bp overlap
GATA4 10 datasets
ChIP DE DE-GATA4-1 572 bp overlap
ChIP DE DE-GATA4-2 622 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
Motif DE_48h DE_48h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
Motif DE_60h DE_60h-GATA4_MA0482.3 8 bp overlap
ChIP foregut GSE117136.GATA4.foregut 454 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 273 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 272 bp overlap
GATA6 19 datasets
ChIP AGS GSE51705.GATA6.AGS 132 bp overlap
ChIP DE DE-GATA6-1 557 bp overlap
ChIP DE DE-GATA6-2 586 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
Motif DE_48h DE_48h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
Motif DE_60h DE_60h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 561 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 548 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 517 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 590 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 550 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 558 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 523 bp overlap
ChIP foregut GSE117136.GATA6.foregut 502 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 474 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 414 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 253 bp overlap
GBX2 2 datasets
Motif DE_36h DE_36h-GBX2_MA0890.2 6 bp overlap
Motif DE_60h DE_60h-GBX2_MA0890.2 6 bp overlap
GRHL2 2 datasets
Motif DE_36h DE_36h-GRHL2_MA1105.3 8 bp overlap
Motif DE_60h DE_60h-GRHL2_MA1105.3 8 bp overlap
Gata3 5 datasets
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
Motif DE_48h DE_48h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
Motif DE_60h DE_60h-Gata3_MA0037.5 8 bp overlap
HESX1 2 datasets
Motif DE_36h DE_36h-HESX1_MA0894.2 6 bp overlap
Motif DE_60h DE_60h-HESX1_MA0894.2 6 bp overlap
HOXA10 2 datasets
Motif DE_36h DE_36h-HOXA10_MA0899.2 9 bp overlap
Motif DE_60h DE_60h-HOXA10_MA0899.2 9 bp overlap
HOXA7 2 datasets
Motif DE_36h DE_36h-HOXA7_MA1498.3 6 bp overlap
Motif DE_60h DE_60h-HOXA7_MA1498.3 6 bp overlap
HOXB13 2 datasets
Motif DE_36h DE_36h-HOXB13_MA0901.3 9 bp overlap
Motif DE_60h DE_60h-HOXB13_MA0901.3 9 bp overlap
HOXD9 2 datasets
Motif DE_36h DE_36h-HOXD9_MA0913.3 9 bp overlap
Motif DE_60h DE_60h-HOXD9_MA0913.3 9 bp overlap
Hmx1 2 datasets
Motif DE_36h DE_36h-Hmx1_MA0896.2 9 bp overlap
Motif DE_60h DE_60h-Hmx1_MA0896.2 9 bp overlap
Hmx2 2 datasets
Motif DE_36h DE_36h-Hmx2_MA0897.2 15 bp overlap
Motif DE_60h DE_60h-Hmx2_MA0897.2 15 bp overlap
Hmx3 2 datasets
Motif DE_36h DE_36h-Hmx3_MA0898.2 9 bp overlap
Motif DE_60h DE_60h-Hmx3_MA0898.2 9 bp overlap
Hnf1A 1 dataset
Motif DE_60h DE_60h-Hnf1A_MA1991.2 10 bp overlap
Ikzf3 2 datasets
Motif DE_36h DE_36h-Ikzf3_MA1992.2 9 bp overlap
Motif DE_60h DE_60h-Ikzf3_MA1992.2 9 bp overlap
Irf1 1 dataset
Motif DE_60h DE_60h-Irf1_MA0050.4 11 bp overlap
KDM1A 1 dataset
ChIP Kasumi-1_GSK690 GSE71739.KDM1A.Kasumi-1_GSK690 194 bp overlap
KLF1 1 dataset
ChIP erythroid_R3R4 GSE43625.KLF1.erythroid_R3R4 80 bp overlap
LBX2 2 datasets
Motif DE_36h DE_36h-LBX2_MA0699.2 6 bp overlap
Motif DE_60h DE_60h-LBX2_MA0699.2 6 bp overlap
LHX2 2 datasets
Motif DE_36h DE_36h-LHX2_MA0700.3 6 bp overlap
Motif DE_60h DE_60h-LHX2_MA0700.3 6 bp overlap
LMO1 1 dataset
ChIP Jurkat GSE94391.LMO1.Jurkat 239 bp overlap
LMO2 1 dataset
ChIP TSU-1621MT GSE60477.LMO2.TSU-1621MT 271 bp overlap
MECOM 1 dataset
ChIP SKH1 GSE102697.MECOM.SKH1 158 bp overlap
MEF2C 3 datasets
Motif DE_36h DE_36h-MEF2C_MA0497.2 11 bp overlap
Motif DE_48h DE_48h-MEF2C_MA0497.2 11 bp overlap
Motif DE_60h DE_60h-MEF2C_MA0497.2 11 bp overlap
MEIS1 3 datasets
ChIP CHRF28811 ERR063469.MEIS1.CHRF28811 293 bp overlap
Motif DE_36h DE_36h-MEIS1_MA0498.3 5 bp overlap
Motif DE_60h DE_60h-MEIS1_MA0498.3 5 bp overlap
MSANTD3 2 datasets
Motif DE_36h DE_36h-MSANTD3_MA1523.2 7 bp overlap
Motif DE_60h DE_60h-MSANTD3_MA1523.2 7 bp overlap
MSX1 2 datasets
Motif DE_36h DE_36h-MSX1_MA0666.3 6 bp overlap
Motif DE_60h DE_60h-MSX1_MA0666.3 6 bp overlap
MSX2 2 datasets
Motif DE_36h DE_36h-MSX2_MA0708.3 6 bp overlap
Motif DE_60h DE_60h-MSX2_MA0708.3 6 bp overlap
MYB 4 datasets
ChIP DU528 GSE94000.MYB.DU528 344 bp overlap
ChIP Jurkat GSE59657.MYB.Jurkat 172 bp overlap
ChIP Loucy GSE94000.MYB.Loucy 315 bp overlap
ChIP PF-382 GSE94000.MYB.PF-382 251 bp overlap
Msx3 2 datasets
Motif DE_36h DE_36h-Msx3_MA0709.2 6 bp overlap
Motif DE_60h DE_60h-Msx3_MA0709.2 6 bp overlap
NCAPH2 1 dataset
ChIP IMR-90_alpha-amanitin-30min GSE118494.NCAPH2.IMR-90_alpha-amanitin-30min 280 bp overlap
NCOR1 1 dataset
ChIP LS180 GSE39277.NCOR1.LS180 95 bp overlap
NFE2 1 dataset
ChIP ProEs GSE59087.NFE2.ProEs 223 bp overlap
Nobox 2 datasets
Motif DE_36h DE_36h-Nobox_MA0125.2 6 bp overlap
Motif DE_60h DE_60h-Nobox_MA0125.2 6 bp overlap
Nr2e3 3 datasets
Motif DE_36h DE_36h-Nr2e3_MA0164.2 6 bp overlap
Motif DE_48h DE_48h-Nr2e3_MA0164.2 6 bp overlap
Motif DE_60h DE_60h-Nr2e3_MA0164.2 6 bp overlap
PBX3 2 datasets
Motif DE_36h DE_36h-PBX3_MA1114.2 11 bp overlap
Motif DE_60h DE_60h-PBX3_MA1114.2 11 bp overlap
PGR 2 datasets
Motif DE_36h DE_36h-PGR_MA2327.1 9 bp overlap
Motif DE_60h DE_60h-PGR_MA2327.1 9 bp overlap
RAX 2 datasets
Motif DE_36h DE_36h-RAX_MA0718.2 6 bp overlap
Motif DE_60h DE_60h-RAX_MA0718.2 6 bp overlap
RNF2 1 dataset
ChIP ME-1_KD GSE128771.RNF2.ME-1_KD 311 bp overlap
RUNX1 5 datasets
ChIP Jurkat GSE76181.RUNX1.Jurkat 190 bp overlap
ChIP ME-1 GSE46044.RUNX1.ME-1 238 bp overlap
ChIP ME-1_CBFB-MYH11-KD GSE117138.RUNX1.ME-1_CBFB-MYH11-KD 331 bp overlap
ChIP ME-1_Con GSE128771.RUNX1.ME-1_Con 331 bp overlap
ChIP THP-1 GSE79899.RUNX1.THP-1 241 bp overlap
RUNX2 1 dataset
ChIP PER-117 GSE151819.RUNX2.PER-117 264 bp overlap
SMAD1 1 dataset
ChIP CD34_PROG_BMP GSE29194.SMAD1.CD34_PROG_BMP 123 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 479 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 341 bp overlap
SMAD2_3 2 datasets
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 268 bp overlap
ChIP DE_D2 S53-DE-d2-SMAD2_3-exp2 335 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 225 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 238 bp overlap
SOX18 2 datasets
Motif DE_36h DE_36h-SOX18_MA1563.2 8 bp overlap
Motif DE_60h DE_60h-SOX18_MA1563.2 8 bp overlap
SPDEF 2 datasets
Motif DE_36h DE_36h-SPDEF_MA0686.2 10 bp overlap
Motif DE_60h DE_60h-SPDEF_MA0686.2 10 bp overlap
SPIB 2 datasets
Motif DE_36h DE_36h-SPIB_MA0081.3 13 bp overlap
Motif DE_60h DE_60h-SPIB_MA0081.3 13 bp overlap
SPIC 2 datasets
Motif DE_36h DE_36h-SPIC_MA0687.2 13 bp overlap
Motif DE_60h DE_60h-SPIC_MA0687.2 13 bp overlap
STAT1 2 datasets
ChIP SET-2_DMSO GSE100566.STAT1.SET-2_DMSO 170 bp overlap
ChIP SET-2_cortistatin-A GSE100566.STAT1.SET-2_cortistatin-A 343 bp overlap
STAT1_pS727 1 dataset
ChIP SET-2 GSE100566.STAT1_pS727.SET-2 256 bp overlap
Spi1 2 datasets
Motif DE_36h DE_36h-Spi1_MA0080.7 13 bp overlap
Motif DE_60h DE_60h-Spi1_MA0080.7 13 bp overlap
TAL1 5 datasets
ChIP CD34 GSE52924.TAL1.CD34 147 bp overlap
ChIP CHRF28811 ERP008568.TAL1.CHRF28811 253 bp overlap
ChIP MOLT-3 GSE59657.TAL1.MOLT-3 223 bp overlap
ChIP ProEs GSE59087.TAL1.ProEs 197 bp overlap
ChIP TSU-1621MT GSE60477.TAL1.TSU-1621MT 220 bp overlap
TCF12 2 datasets
ChIP Kasumi-1 GSE23730.TCF12.Kasumi-1 182 bp overlap
ChIP ME-1 GSE46044.TCF12.ME-1 274 bp overlap
TCF3 1 dataset
ChIP Kasumi-1 GSE43834.TCF3.Kasumi-1 220 bp overlap
TCF7L1 1 dataset
Motif DE_60h DE_60h-TCF7L1_MA1421.1 12 bp overlap
TRPS1 5 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
Motif DE_48h DE_48h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
Motif DE_60h DE_60h-TRPS1_MA1970.2 8 bp overlap
VENTX 2 datasets
Motif DE_36h DE_36h-VENTX_MA0724.1 9 bp overlap
Motif DE_60h DE_60h-VENTX_MA0724.1 9 bp overlap
YBX1 1 dataset
ChIP HEL GSE146717.YBX1.HEL 287 bp overlap
ZNF175 2 datasets
Motif DE_36h DE_36h-ZNF175_MA2332.1 9 bp overlap
Motif DE_60h DE_60h-ZNF175_MA2332.1 9 bp overlap
ZNF282 2 datasets
Motif DE_36h DE_36h-ZNF282_MA1154.2 15 bp overlap
Motif DE_60h DE_60h-ZNF282_MA1154.2 15 bp overlap
ZNF770 3 datasets
Motif DE_36h DE_36h-ZNF770_MA2099.1 8 bp overlap
Motif DE_48h DE_48h-ZNF770_MA2099.1 8 bp overlap
Motif DE_60h DE_60h-ZNF770_MA2099.1 8 bp overlap
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