chr4 : 92,848,141 – 92,849,226
1,085 bp 88 TFs 0 linked genes
This 1.1 kb open chromatin element has no linked target genes and is bound by 88 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:92,843,141 – 92,854,226
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
88 transcription factors
Source
Cell type
Arid3a 3 datasets
Motif DE_48h DE_48h-Arid3a_MA0151.1 6 bp overlap
Motif DE_60h DE_60h-Arid3a_MA0151.1 6 bp overlap
Motif DE_72h DE_72h-Arid3a_MA0151.1 6 bp overlap
CEBPA 2 datasets
Motif DE_60h DE_60h-CEBPA_MA0102.5 10 bp overlap
Motif DE_72h DE_72h-CEBPA_MA0102.5 10 bp overlap
CEBPD 2 datasets
Motif DE_60h DE_60h-CEBPD_MA0836.3 8 bp overlap
Motif DE_72h DE_72h-CEBPD_MA0836.3 8 bp overlap
CRX 1 dataset
ChIP retina_Hu6 GSE137311.CRX.retina_Hu6 335 bp overlap
DUX4 1 dataset
ChIP WA01 GSE94322.DUX4.WA01 313 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 150 bp overlap
FIGLA 6 datasets
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
Motif DE_48h DE_48h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
Motif DE_60h DE_60h-FIGLA_MA0820.2 6 bp overlap
Motif DE_72h DE_72h-FIGLA_MA0820.2 6 bp overlap
Motif DE_72h DE_72h-FIGLA_MA0820.2 6 bp overlap
FOXA2 2 datasets
ChIP DE DE-FOXA2-1 1080 bp overlap
ChIP DE DE-FOXA2-2 1085 bp overlap
GATA4 6 datasets
ChIP DE DE-GATA4-1 145 bp overlap
ChIP DE DE-GATA4-1 706 bp overlap
ChIP DE DE-GATA4-2 160 bp overlap
ChIP DE DE-GATA4-2 788 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 69 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 303 bp overlap
GATA6 6 datasets
ChIP DE DE-GATA6-1 130 bp overlap
ChIP DE DE-GATA6-1 756 bp overlap
ChIP DE DE-GATA6-2 172 bp overlap
ChIP DE DE-GATA6-2 823 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 339 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 449 bp overlap
HMBOX1 3 datasets
Motif DE_48h DE_48h-HMBOX1_MA0895.2 7 bp overlap
Motif DE_60h DE_60h-HMBOX1_MA0895.2 7 bp overlap
Motif DE_72h DE_72h-HMBOX1_MA0895.2 7 bp overlap
HNF1A 5 datasets
Motif DE_48h DE_48h-HNF1A_MA0046.3 13 bp overlap
Motif DE_60h DE_60h-HNF1A_MA0046.3 13 bp overlap
Motif DE_72h DE_72h-HNF1A_MA0046.3 13 bp overlap
ChIP HEE_1 GSE76376.HNF1A.HEE_1 382 bp overlap
ChIP HEE_5 GSE76376.HNF1A.HEE_5 233 bp overlap
HNF1B 5 datasets
Motif DE_48h DE_48h-HNF1B_MA0153.2 13 bp overlap
Motif DE_60h DE_60h-HNF1B_MA0153.2 13 bp overlap
Motif DE_72h DE_72h-HNF1B_MA0153.2 13 bp overlap
ChIP PDAC GSE64557.HNF1B.PDAC 438 bp overlap
ChIP foregut GSE117136.HNF1B.foregut 687 bp overlap
HNF4A 5 datasets
Motif DE_48h DE_48h-HNF4A_MA0114.5 9 bp overlap
Motif DE_60h DE_60h-HNF4A_MA0114.5 9 bp overlap
Motif DE_60h DE_60h-HNF4A_MA0114.5 9 bp overlap
Motif DE_72h DE_72h-HNF4A_MA0114.5 9 bp overlap
Motif DE_72h DE_72h-HNF4A_MA0114.5 9 bp overlap
HNF4G 5 datasets
Motif DE_48h DE_48h-HNF4G_MA0484.3 9 bp overlap
Motif DE_60h DE_60h-HNF4G_MA0484.3 9 bp overlap
Motif DE_60h DE_60h-HNF4G_MA0484.3 9 bp overlap
Motif DE_72h DE_72h-HNF4G_MA0484.3 9 bp overlap
Motif DE_72h DE_72h-HNF4G_MA0484.3 9 bp overlap
HOXA4 3 datasets
Motif DE_48h DE_48h-HOXA4_MA1496.2 7 bp overlap
Motif DE_60h DE_60h-HOXA4_MA1496.2 7 bp overlap
Motif DE_72h DE_72h-HOXA4_MA1496.2 7 bp overlap
HOXB4 6 datasets
Motif DE_48h DE_48h-HOXB4_MA1499.2 6 bp overlap
Motif DE_48h DE_48h-HOXB4_MA1499.2 6 bp overlap
Motif DE_60h DE_60h-HOXB4_MA1499.2 6 bp overlap
Motif DE_60h DE_60h-HOXB4_MA1499.2 6 bp overlap
Motif DE_72h DE_72h-HOXB4_MA1499.2 6 bp overlap
Motif DE_72h DE_72h-HOXB4_MA1499.2 6 bp overlap
HOXC4 6 datasets
Motif DE_48h DE_48h-HOXC4_MA1504.2 6 bp overlap
Motif DE_48h DE_48h-HOXC4_MA1504.2 6 bp overlap
Motif DE_60h DE_60h-HOXC4_MA1504.2 6 bp overlap
Motif DE_60h DE_60h-HOXC4_MA1504.2 6 bp overlap
Motif DE_72h DE_72h-HOXC4_MA1504.2 6 bp overlap
Motif DE_72h DE_72h-HOXC4_MA1504.2 6 bp overlap
HOXD4 6 datasets
Motif DE_48h DE_48h-HOXD4_MA1507.2 6 bp overlap
Motif DE_48h DE_48h-HOXD4_MA1507.2 6 bp overlap
Motif DE_60h DE_60h-HOXD4_MA1507.2 6 bp overlap
Motif DE_60h DE_60h-HOXD4_MA1507.2 6 bp overlap
Motif DE_72h DE_72h-HOXD4_MA1507.2 6 bp overlap
Motif DE_72h DE_72h-HOXD4_MA1507.2 6 bp overlap
Hnf1A 3 datasets
Motif DE_48h DE_48h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_60h DE_60h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_72h DE_72h-Hnf1A_MA1991.2 10 bp overlap
Isl1 3 datasets
Motif DE_48h DE_48h-Isl1_MA1608.2 7 bp overlap
Motif DE_60h DE_60h-Isl1_MA1608.2 7 bp overlap
Motif DE_72h DE_72h-Isl1_MA1608.2 7 bp overlap
Lef1 3 datasets
Motif DE_48h DE_48h-Lef1_MA0768.3 8 bp overlap
Motif DE_60h DE_60h-Lef1_MA0768.3 8 bp overlap
Motif DE_72h DE_72h-Lef1_MA0768.3 8 bp overlap
MAF 3 datasets
Motif DE_48h DE_48h-MAF_MA1520.2 13 bp overlap
Motif DE_60h DE_60h-MAF_MA1520.2 13 bp overlap
Motif DE_72h DE_72h-MAF_MA1520.2 13 bp overlap
MAFA 3 datasets
Motif DE_48h DE_48h-MAFA_MA1521.2 13 bp overlap
Motif DE_60h DE_60h-MAFA_MA1521.2 13 bp overlap
Motif DE_72h DE_72h-MAFA_MA1521.2 13 bp overlap
MEF2C 3 datasets
Motif DE_48h DE_48h-MEF2C_MA0497.2 11 bp overlap
Motif DE_60h DE_60h-MEF2C_MA0497.2 11 bp overlap
Motif DE_72h DE_72h-MEF2C_MA0497.2 11 bp overlap
MEIS1 6 datasets
Motif DE_48h DE_48h-MEIS1_MA0498.3 5 bp overlap
Motif DE_48h DE_48h-MEIS1_MA1639.2 9 bp overlap
Motif DE_60h DE_60h-MEIS1_MA0498.3 5 bp overlap
Motif DE_60h DE_60h-MEIS1_MA1639.2 9 bp overlap
Motif DE_72h DE_72h-MEIS1_MA0498.3 5 bp overlap
Motif DE_72h DE_72h-MEIS1_MA1639.2 9 bp overlap
MEIS2 3 datasets
Motif DE_48h DE_48h-MEIS2_MA1640.2 9 bp overlap
Motif DE_60h DE_60h-MEIS2_MA1640.2 9 bp overlap
Motif DE_72h DE_72h-MEIS2_MA1640.2 9 bp overlap
MGA 3 datasets
Motif DE_48h DE_48h-MGA_MA0801.1 8 bp overlap
Motif DE_60h DE_60h-MGA_MA0801.1 8 bp overlap
Motif DE_72h DE_72h-MGA_MA0801.1 8 bp overlap
MGA::EVX1 3 datasets
Motif DE_48h DE_48h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_60h DE_60h-MGAEVX1_MA1960.2 11 bp overlap
Motif DE_72h DE_72h-MGAEVX1_MA1960.2 11 bp overlap
NFIB 2 datasets
Motif DE_60h DE_60h-NFIB_MA1643.2 17 bp overlap
Motif DE_72h DE_72h-NFIB_MA1643.2 17 bp overlap
NKX2-2 3 datasets
Motif DE_48h DE_48h-NKX2-2_MA1645.2 8 bp overlap
Motif DE_60h DE_60h-NKX2-2_MA1645.2 8 bp overlap
Motif DE_72h DE_72h-NKX2-2_MA1645.2 8 bp overlap
NKX6-1 3 datasets
Motif DE_48h DE_48h-NKX6-1_MA0674.2 7 bp overlap
Motif DE_60h DE_60h-NKX6-1_MA0674.2 7 bp overlap
Motif DE_72h DE_72h-NKX6-1_MA0674.2 7 bp overlap
NKX6-3 3 datasets
Motif DE_48h DE_48h-NKX6-3_MA1530.2 8 bp overlap
Motif DE_60h DE_60h-NKX6-3_MA1530.2 8 bp overlap
Motif DE_72h DE_72h-NKX6-3_MA1530.2 8 bp overlap
NR1H4::RXRA 2 datasets
Motif DE_60h DE_60h-NR1H4RXRA_MA1146.2 13 bp overlap
Motif DE_72h DE_72h-NR1H4RXRA_MA1146.2 13 bp overlap
NR2C1 2 datasets
Motif DE_60h DE_60h-NR2C1_MA1535.2 6 bp overlap
Motif DE_72h DE_72h-NR2C1_MA1535.2 6 bp overlap
NR2C2 2 datasets
Motif DE_60h DE_60h-NR2C2_MA1536.2 6 bp overlap
Motif DE_72h DE_72h-NR2C2_MA1536.2 6 bp overlap
NR2F1 2 datasets
Motif DE_60h DE_60h-NR2F1_MA1538.1 15 bp overlap
Motif DE_72h DE_72h-NR2F1_MA1538.1 15 bp overlap
NR2F6 2 datasets
Motif DE_60h DE_60h-NR2F6_MA1539.1 15 bp overlap
Motif DE_72h DE_72h-NR2F6_MA1539.1 15 bp overlap
NR4A1 3 datasets
Motif DE_48h DE_48h-NR4A1_MA1112.3 8 bp overlap
Motif DE_60h DE_60h-NR4A1_MA1112.3 8 bp overlap
Motif DE_72h DE_72h-NR4A1_MA1112.3 8 bp overlap
NR4A2 3 datasets
Motif DE_48h DE_48h-NR4A2_MA0160.3 8 bp overlap
Motif DE_60h DE_60h-NR4A2_MA0160.3 8 bp overlap
Motif DE_72h DE_72h-NR4A2_MA0160.3 8 bp overlap
NR4A2::RXRA 2 datasets
Motif DE_60h DE_60h-NR4A2RXRA_MA1147.2 13 bp overlap
Motif DE_72h DE_72h-NR4A2RXRA_MA1147.2 13 bp overlap
Nr1H2 2 datasets
Motif DE_60h DE_60h-Nr1H2_MA1996.2 6 bp overlap
Motif DE_72h DE_72h-Nr1H2_MA1996.2 6 bp overlap
Nr1H4 2 datasets
Motif DE_60h DE_60h-Nr1H4_MA1110.3 6 bp overlap
Motif DE_72h DE_72h-Nr1H4_MA1110.3 6 bp overlap
Nr1h3 2 datasets
Motif DE_60h DE_60h-Nr1h3_MA2337.1 6 bp overlap
Motif DE_72h DE_72h-Nr1h3_MA2337.1 6 bp overlap
OTX2 1 dataset
ChIP retina_Hu7 GSE137311.OTX2.retina_Hu7 290 bp overlap
PBX1 6 datasets
Motif DE_48h DE_48h-PBX1_MA0070.2 9 bp overlap
Motif DE_48h DE_48h-PBX1_MA0070.2 9 bp overlap
Motif DE_60h DE_60h-PBX1_MA0070.2 9 bp overlap
Motif DE_60h DE_60h-PBX1_MA0070.2 9 bp overlap
Motif DE_72h DE_72h-PBX1_MA0070.2 9 bp overlap
Motif DE_72h DE_72h-PBX1_MA0070.2 9 bp overlap
PDX1 2 datasets
ChIP hiPSC GSE125768.PDX1.hiPSC 329 bp overlap
ChIP hiPSC_derived_PPs GSE106949.PDX1.hiPSC_derived_PPs 152 bp overlap
PHOX2B 3 datasets
Motif DE_48h DE_48h-PHOX2B_MA0681.3 12 bp overlap
Motif DE_60h DE_60h-PHOX2B_MA0681.3 12 bp overlap
Motif DE_72h DE_72h-PHOX2B_MA0681.3 12 bp overlap
PLAG1 2 datasets
Motif DE_60h DE_60h-PLAG1_MA0163.1 14 bp overlap
Motif DE_72h DE_72h-PLAG1_MA0163.1 14 bp overlap
POLR2A 1 dataset
ChIP neural cell ENCFF604SPB 195 bp overlap
POU2F1::SOX2 3 datasets
Motif DE_48h DE_48h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif DE_60h DE_60h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif DE_72h DE_72h-POU2F1SOX2_MA1962.1 17 bp overlap
POU4F2 3 datasets
Motif DE_48h DE_48h-POU4F2_MA0683.2 15 bp overlap
Motif DE_60h DE_60h-POU4F2_MA0683.2 15 bp overlap
Motif DE_72h DE_72h-POU4F2_MA0683.2 15 bp overlap
POU6F1 3 datasets
Motif DE_48h DE_48h-POU6F1_MA1549.2 7 bp overlap
Motif DE_60h DE_60h-POU6F1_MA1549.2 7 bp overlap
Motif DE_72h DE_72h-POU6F1_MA1549.2 7 bp overlap
POU6F2 3 datasets
Motif DE_48h DE_48h-POU6F2_MA0793.2 9 bp overlap
Motif DE_60h DE_60h-POU6F2_MA0793.2 9 bp overlap
Motif DE_72h DE_72h-POU6F2_MA0793.2 9 bp overlap
PROP1 3 datasets
Motif DE_48h DE_48h-PROP1_MA0715.1 11 bp overlap
Motif DE_60h DE_60h-PROP1_MA0715.1 11 bp overlap
Motif DE_72h DE_72h-PROP1_MA0715.1 11 bp overlap
Pou5f1::Sox2 3 datasets
Motif DE_48h DE_48h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_60h DE_60h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_72h DE_72h-Pou5f1Sox2_MA0142.1 15 bp overlap
Ptf1A 3 datasets
Motif DE_48h DE_48h-Ptf1A_MA1620.2 8 bp overlap
Motif DE_60h DE_60h-Ptf1A_MA1620.2 8 bp overlap
Motif DE_72h DE_72h-Ptf1A_MA1620.2 8 bp overlap
RARB 2 datasets
Motif DE_60h DE_60h-RARB_MA1552.2 13 bp overlap
Motif DE_72h DE_72h-RARB_MA1552.2 13 bp overlap
RORB 2 datasets
Motif DE_60h DE_60h-RORB_MA1150.2 10 bp overlap
Motif DE_72h DE_72h-RORB_MA1150.2 10 bp overlap
Rarb 3 datasets
Motif DE_48h DE_48h-Rarb_MA0858.1 17 bp overlap
Motif DE_60h DE_60h-Rarb_MA0858.1 17 bp overlap
Motif DE_72h DE_72h-Rarb_MA0858.1 17 bp overlap
SMAD2-3 1 dataset
ChIP HUES-8_DE GSE109524.SMAD2-3.HUES-8_DE 822 bp overlap
SMAD2_3 1 dataset
ChIP DE_D2 S01-DE-d2-SMAD2_3-exp1 263 bp overlap
SOX14 5 datasets
Motif DE_48h DE_48h-SOX14_MA1562.2 9 bp overlap
Motif DE_60h DE_60h-SOX14_MA1562.2 9 bp overlap
Motif DE_60h DE_60h-SOX14_MA1562.2 9 bp overlap
Motif DE_72h DE_72h-SOX14_MA1562.2 9 bp overlap
Motif DE_72h DE_72h-SOX14_MA1562.2 9 bp overlap
SOX15 3 datasets
Motif DE_48h DE_48h-SOX15_MA1152.2 7 bp overlap
Motif DE_60h DE_60h-SOX15_MA1152.2 7 bp overlap
Motif DE_72h DE_72h-SOX15_MA1152.2 7 bp overlap
SOX17 1 dataset
ChIP DE_D2 DED2-SOX17_Batch_II 230 bp overlap
SOX17_M 1 dataset
ChIP DE_D2 DED2-SOX17-M_Batch_II 337 bp overlap
SOX18 2 datasets
Motif DE_60h DE_60h-SOX18_MA1563.2 8 bp overlap
Motif DE_72h DE_72h-SOX18_MA1563.2 8 bp overlap
SOX8 2 datasets
Motif DE_60h DE_60h-SOX8_MA0868.3 7 bp overlap
Motif DE_72h DE_72h-SOX8_MA0868.3 7 bp overlap
SOX9 2 datasets
Motif DE_60h DE_60h-SOX9_MA0077.2 8 bp overlap
Motif DE_72h DE_72h-SOX9_MA0077.2 8 bp overlap
SREBF1 3 datasets
Motif DE_48h DE_48h-SREBF1_MA0595.1 10 bp overlap
Motif DE_60h DE_60h-SREBF1_MA0595.1 10 bp overlap
Motif DE_72h DE_72h-SREBF1_MA0595.1 10 bp overlap
SREBF2 3 datasets
Motif DE_48h DE_48h-SREBF2_MA0596.1 10 bp overlap
Motif DE_60h DE_60h-SREBF2_MA0596.1 10 bp overlap
Motif DE_72h DE_72h-SREBF2_MA0596.1 10 bp overlap
Sox11 2 datasets
Motif DE_60h DE_60h-Sox11_MA0869.3 8 bp overlap
Motif DE_72h DE_72h-Sox11_MA0869.3 8 bp overlap
Sox17 2 datasets
Motif DE_60h DE_60h-Sox17_MA0078.3 10 bp overlap
Motif DE_72h DE_72h-Sox17_MA0078.3 10 bp overlap
Sox5 2 datasets
Motif DE_60h DE_60h-Sox5_MA0087.3 8 bp overlap
Motif DE_72h DE_72h-Sox5_MA0087.3 8 bp overlap
Sox6 2 datasets
Motif DE_60h DE_60h-Sox6_MA0515.1 10 bp overlap
Motif DE_72h DE_72h-Sox6_MA0515.1 10 bp overlap
Sox7 2 datasets
Motif DE_60h DE_60h-Sox7_MA2095.1 10 bp overlap
Motif DE_72h DE_72h-Sox7_MA2095.1 10 bp overlap
TBX5 3 datasets
Motif DE_48h DE_48h-TBX5_MA0807.1 8 bp overlap
Motif DE_60h DE_60h-TBX5_MA0807.1 8 bp overlap
Motif DE_72h DE_72h-TBX5_MA0807.1 8 bp overlap
TCF7 3 datasets
Motif DE_48h DE_48h-TCF7_MA0769.3 7 bp overlap
Motif DE_60h DE_60h-TCF7_MA0769.3 7 bp overlap
Motif DE_72h DE_72h-TCF7_MA0769.3 7 bp overlap
Thap11 3 datasets
Motif DE_48h DE_48h-Thap11_MA1573.2 14 bp overlap
Motif DE_60h DE_60h-Thap11_MA1573.2 14 bp overlap
Motif DE_72h DE_72h-Thap11_MA1573.2 14 bp overlap
VENTX 3 datasets
Motif DE_48h DE_48h-VENTX_MA0724.1 9 bp overlap
Motif DE_60h DE_60h-VENTX_MA0724.1 9 bp overlap
Motif DE_72h DE_72h-VENTX_MA0724.1 9 bp overlap
Yy1 2 datasets
Motif DE_60h DE_60h-Yy1_MA0095.4 8 bp overlap
Motif DE_72h DE_72h-Yy1_MA0095.4 8 bp overlap
ZBTB12 3 datasets
Motif DE_48h DE_48h-ZBTB12_MA1649.2 7 bp overlap
Motif DE_60h DE_60h-ZBTB12_MA1649.2 7 bp overlap
Motif DE_72h DE_72h-ZBTB12_MA1649.2 7 bp overlap
ZBTB18 3 datasets
Motif DE_48h DE_48h-ZBTB18_MA0698.2 11 bp overlap
Motif DE_60h DE_60h-ZBTB18_MA0698.2 11 bp overlap
Motif DE_72h DE_72h-ZBTB18_MA0698.2 11 bp overlap
ZBTB26 3 datasets
Motif DE_48h DE_48h-ZBTB26_MA1579.2 8 bp overlap
Motif DE_60h DE_60h-ZBTB26_MA1579.2 8 bp overlap
Motif DE_72h DE_72h-ZBTB26_MA1579.2 8 bp overlap
ZEB1 6 datasets
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
Motif DE_48h DE_48h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
Motif DE_60h DE_60h-ZEB1_MA0103.4 6 bp overlap
Motif DE_72h DE_72h-ZEB1_MA0103.4 6 bp overlap
Motif DE_72h DE_72h-ZEB1_MA0103.4 6 bp overlap
ZFP14 2 datasets
Motif DE_60h DE_60h-ZFP14_MA1972.1 15 bp overlap
Motif DE_72h DE_72h-ZFP14_MA1972.1 15 bp overlap
ZNF136 3 datasets
Motif DE_48h DE_48h-ZNF136_MA1588.1 15 bp overlap
Motif DE_60h DE_60h-ZNF136_MA1588.1 15 bp overlap
Motif DE_72h DE_72h-ZNF136_MA1588.1 15 bp overlap
ZNF707 3 datasets
Motif DE_48h DE_48h-ZNF707_MA1715.1 15 bp overlap
Motif DE_60h DE_60h-ZNF707_MA1715.1 15 bp overlap
Motif DE_72h DE_72h-ZNF707_MA1715.1 15 bp overlap
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