chr8 : 56,544,095 56,544,813
718 bp 91 TFs 0 linked genes
This 718 bp open chromatin element has no linked target genes and is bound by 91 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr8:56,539,095 – 56,549,813
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
91 transcription factors
Source
Cell type
BACH1 1 dataset
ChIP H1 ENCFF282VDB 321 bp overlap
BCL6B 3 datasets
Motif DE_12h DE_12h-BCL6B_MA0731.1 17 bp overlap
Motif DE_24h DE_24h-BCL6B_MA0731.1 17 bp overlap
Motif DE_36h DE_36h-BCL6B_MA0731.1 17 bp overlap
BRD4 2 datasets
ChIP MV4-11_DMSO GSE120715.BRD4.MV4-11_DMSO 220 bp overlap
ChIP SEM GSE83671.BRD4.SEM 639 bp overlap
CTCF 1 dataset
ChIP leukemia GSE142161.CTCF.leukemia 138 bp overlap
Ddit3::Cebpa 3 datasets
Motif DE_12h DE_12h-Ddit3Cebpa_MA0019.2 10 bp overlap
Motif DE_24h DE_24h-Ddit3Cebpa_MA0019.2 10 bp overlap
Motif DE_36h DE_36h-Ddit3Cebpa_MA0019.2 10 bp overlap
Dux 3 datasets
Motif DE_12h DE_12h-Dux_MA0611.3 11 bp overlap
Motif DE_24h DE_24h-Dux_MA0611.3 11 bp overlap
Motif DE_36h DE_36h-Dux_MA0611.3 11 bp overlap
EBF1 4 datasets
Motif DE_12h DE_12h-EBF1_MA0154.5 11 bp overlap
Motif DE_24h DE_24h-EBF1_MA0154.5 11 bp overlap
Motif DE_36h DE_36h-EBF1_MA0154.5 11 bp overlap
ChIP NALM-6 GSE126300.EBF1.NALM-6 440 bp overlap
EOMES 5 datasets
Motif DE_12h DE_12h-EOMES_MA0800.2 9 bp overlap
Motif DE_12h DE_12h-EOMES_MA0800.2 9 bp overlap
Motif DE_24h DE_24h-EOMES_MA0800.2 9 bp overlap
Motif DE_36h DE_36h-EOMES_MA0800.2 9 bp overlap
Motif DE_36h DE_36h-EOMES_MA0800.2 9 bp overlap
ESR1 1 dataset
ChIP MCF-7_E2 ERP000901.ESR1.MCF-7_E2 132 bp overlap
EZH2 1 dataset
ChIP HCT-116 ENCSR046HGP.EZH2.HCT-116 238 bp overlap
Ebf4 3 datasets
Motif DE_12h DE_12h-Ebf4_MA2122.1 11 bp overlap
Motif DE_24h DE_24h-Ebf4_MA2122.1 11 bp overlap
Motif DE_36h DE_36h-Ebf4_MA2122.1 11 bp overlap
FOS 3 datasets
Motif DE_12h DE_12h-FOS_MA0476.2 8 bp overlap
Motif DE_24h DE_24h-FOS_MA0476.2 8 bp overlap
Motif DE_36h DE_36h-FOS_MA0476.2 8 bp overlap
FOSL1::JUND 3 datasets
Motif DE_12h DE_12h-FOSL1JUND_MA1142.2 8 bp overlap
Motif DE_24h DE_24h-FOSL1JUND_MA1142.2 8 bp overlap
Motif DE_36h DE_36h-FOSL1JUND_MA1142.2 8 bp overlap
FOXD3 1 dataset
Motif DE_12h DE_12h-FOXD3_MA0041.3 14 bp overlap
FOXH1 3 datasets
Motif DE_12h DE_12h-FOXH1_MA0479.2 8 bp overlap
Motif DE_24h DE_24h-FOXH1_MA0479.2 8 bp overlap
Motif DE_36h DE_36h-FOXH1_MA0479.2 8 bp overlap
GATA1::TAL1 3 datasets
Motif DE_12h DE_12h-GATA1TAL1_MA0140.3 17 bp overlap
Motif DE_24h DE_24h-GATA1TAL1_MA0140.3 17 bp overlap
Motif DE_36h DE_36h-GATA1TAL1_MA0140.3 17 bp overlap
GATA6 5 datasets
ChIP DE_D1 S09-DE-d1-GATA6-exp1 458 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 268 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 534 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 591 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 600 bp overlap
Hnf1A 6 datasets
Motif DE_12h DE_12h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_12h DE_12h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_24h DE_24h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_24h DE_24h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_36h DE_36h-Hnf1A_MA1991.2 10 bp overlap
Motif DE_36h DE_36h-Hnf1A_MA1991.2 10 bp overlap
IKZF1 1 dataset
ChIP pre-B-cell GSE107886.IKZF1.pre-B-cell 517 bp overlap
JUN 4 datasets
ChIP 786-O GSE86092.JUN.786-O 173 bp overlap
ChIP DE_D1 S08-DE-d1-JUN-exp1 311 bp overlap
ChIP DE_D1 S40-DE-d1-JUN-exp2 332 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.JUN.HUES-8_DE_ctrl 509 bp overlap
Jun 3 datasets
Motif DE_12h DE_12h-Jun_MA0489.3 8 bp overlap
Motif DE_24h DE_24h-Jun_MA0489.3 8 bp overlap
Motif DE_36h DE_36h-Jun_MA0489.3 8 bp overlap
LIN54 3 datasets
Motif DE_12h DE_12h-LIN54_MA0619.2 7 bp overlap
Motif DE_24h DE_24h-LIN54_MA0619.2 7 bp overlap
Motif DE_36h DE_36h-LIN54_MA0619.2 7 bp overlap
Lef1 3 datasets
Motif DE_12h DE_12h-Lef1_MA0768.3 8 bp overlap
Motif DE_24h DE_24h-Lef1_MA0768.3 8 bp overlap
Motif DE_36h DE_36h-Lef1_MA0768.3 8 bp overlap
MED 1 dataset
ChIP SEM GSE83671.MED.SEM 466 bp overlap
MED1 1 dataset
ChIP MCF-7_SHRARS_E2 GSE60270.MED1.MCF-7_SHRARS_E2 158 bp overlap
MGA 2 datasets
Motif DE_12h DE_12h-MGA_MA0801.1 8 bp overlap
Motif DE_36h DE_36h-MGA_MA0801.1 8 bp overlap
MYOD1 1 dataset
ChIP RH4 GSE83726.MYOD1.RH4 165 bp overlap
MZF1 3 datasets
Motif DE_12h DE_12h-MZF1_MA0056.3 8 bp overlap
Motif DE_24h DE_24h-MZF1_MA0056.3 8 bp overlap
Motif DE_36h DE_36h-MZF1_MA0056.3 8 bp overlap
Mecom 1 dataset
Motif DE_12h DE_12h-Mecom_MA0029.2 11 bp overlap
NANOG 3 datasets
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 455 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 203 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 198 bp overlap
NKX2-3 3 datasets
Motif DE_12h DE_12h-NKX2-3_MA0672.2 8 bp overlap
Motif DE_24h DE_24h-NKX2-3_MA0672.2 8 bp overlap
Motif DE_36h DE_36h-NKX2-3_MA0672.2 8 bp overlap
NKX2-4 3 datasets
Motif DE_12h DE_12h-NKX2-4_MA2003.2 8 bp overlap
Motif DE_24h DE_24h-NKX2-4_MA2003.2 8 bp overlap
Motif DE_36h DE_36h-NKX2-4_MA2003.2 8 bp overlap
NKX2-8 3 datasets
Motif DE_12h DE_12h-NKX2-8_MA0673.2 8 bp overlap
Motif DE_24h DE_24h-NKX2-8_MA0673.2 8 bp overlap
Motif DE_36h DE_36h-NKX2-8_MA0673.2 8 bp overlap
PAX5 1 dataset
ChIP NALM-6 GSE126300.PAX5.NALM-6 172 bp overlap
POLR2A 1 dataset
ChIP SK-N-MC ENCFF088IVG 491 bp overlap
POU1F1 3 datasets
Motif DE_12h DE_12h-POU1F1_MA0784.3 14 bp overlap
Motif DE_24h DE_24h-POU1F1_MA0784.3 14 bp overlap
Motif DE_36h DE_36h-POU1F1_MA0784.3 14 bp overlap
POU2F1 3 datasets
Motif DE_12h DE_12h-POU2F1_MA0785.2 9 bp overlap
Motif DE_24h DE_24h-POU2F1_MA0785.2 9 bp overlap
Motif DE_36h DE_36h-POU2F1_MA0785.2 9 bp overlap
POU2F1::SOX2 3 datasets
Motif DE_12h DE_12h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif DE_24h DE_24h-POU2F1SOX2_MA1962.1 17 bp overlap
Motif DE_36h DE_36h-POU2F1SOX2_MA1962.1 17 bp overlap
POU2F2 3 datasets
Motif DE_12h DE_12h-POU2F2_MA0507.3 13 bp overlap
Motif DE_24h DE_24h-POU2F2_MA0507.3 13 bp overlap
Motif DE_36h DE_36h-POU2F2_MA0507.3 13 bp overlap
POU2F3 4 datasets
Motif DE_12h DE_12h-POU2F3_MA0627.3 9 bp overlap
Motif DE_24h DE_24h-POU2F3_MA0627.3 9 bp overlap
Motif DE_36h DE_36h-POU2F3_MA0627.3 9 bp overlap
ChIP NCI-H1048 GSE115123.POU2F3.NCI-H1048 219 bp overlap
POU3F1 3 datasets
Motif DE_12h DE_12h-POU3F1_MA0786.2 10 bp overlap
Motif DE_24h DE_24h-POU3F1_MA0786.2 10 bp overlap
Motif DE_36h DE_36h-POU3F1_MA0786.2 10 bp overlap
POU3F2 3 datasets
Motif DE_12h DE_12h-POU3F2_MA0787.1 12 bp overlap
Motif DE_24h DE_24h-POU3F2_MA0787.1 12 bp overlap
Motif DE_36h DE_36h-POU3F2_MA0787.1 12 bp overlap
POU3F4 3 datasets
Motif DE_12h DE_12h-POU3F4_MA0789.1 9 bp overlap
Motif DE_24h DE_24h-POU3F4_MA0789.1 9 bp overlap
Motif DE_36h DE_36h-POU3F4_MA0789.1 9 bp overlap
POU5F1 7 datasets
Motif DE_12h DE_12h-POU5F1_MA1115.2 7 bp overlap
Motif DE_24h DE_24h-POU5F1_MA1115.2 7 bp overlap
Motif DE_36h DE_36h-POU5F1_MA1115.2 7 bp overlap
ChIP DE_D1 DED1-OCT4_Batch_II 538 bp overlap
ChIP HUES-8 GSE109524.POU5F1.HUES-8 397 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 507 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 219 bp overlap
POU5F1B 3 datasets
Motif DE_12h DE_12h-POU5F1B_MA0792.1 9 bp overlap
Motif DE_24h DE_24h-POU5F1B_MA0792.1 9 bp overlap
Motif DE_36h DE_36h-POU5F1B_MA0792.1 9 bp overlap
POU5F1_M 1 dataset
ChIP DE_D1 DED1-OCT4-M_Batch_II 630 bp overlap
PRDM9 3 datasets
Motif DE_12h DE_12h-PRDM9_MA1723.2 20 bp overlap
Motif DE_24h DE_24h-PRDM9_MA1723.2 20 bp overlap
Motif DE_36h DE_36h-PRDM9_MA1723.2 20 bp overlap
Pou5f1::Sox2 3 datasets
Motif DE_12h DE_12h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_24h DE_24h-Pou5f1Sox2_MA0142.1 15 bp overlap
Motif DE_36h DE_36h-Pou5f1Sox2_MA0142.1 15 bp overlap
RBPJ 3 datasets
Motif DE_12h DE_12h-RBPJ_MA1116.2 6 bp overlap
Motif DE_24h DE_24h-RBPJ_MA1116.2 6 bp overlap
Motif DE_36h DE_36h-RBPJ_MA1116.2 6 bp overlap
REL 3 datasets
Motif DE_12h DE_12h-REL_MA0101.1 10 bp overlap
Motif DE_24h DE_24h-REL_MA0101.1 10 bp overlap
Motif DE_36h DE_36h-REL_MA0101.1 10 bp overlap
RREB1 3 datasets
Motif DE_12h DE_12h-RREB1_MA0073.2 19 bp overlap
Motif DE_24h DE_24h-RREB1_MA0073.2 19 bp overlap
Motif DE_36h DE_36h-RREB1_MA0073.2 19 bp overlap
SMAD2 4 datasets
Motif DE_12h DE_12h-SMAD2_MA1964.2 6 bp overlap
Motif DE_24h DE_24h-SMAD2_MA1964.2 6 bp overlap
Motif DE_36h DE_36h-SMAD2_MA1964.2 6 bp overlap
ChIP hESC_YAP-_activinA_15h GSE99202.SMAD2.hESC_YAP-_activinA_15h 398 bp overlap
SMAD2-3 2 datasets
ChIP HUES-8_DE_JNKi GSE109524.SMAD2-3.HUES-8_DE_JNKi 584 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.SMAD2-3.HUES-8_DE_ctrl 610 bp overlap
SMAD2_3 3 datasets
ChIP DE_D1 S07-DE-d1-SMAD2_3-exp1 414 bp overlap
ChIP DE_D1 S12-DE-d1-SMAD2_3-exp1 481 bp overlap
ChIP DE_D1 S39-DE-d1-SMAD2_3-exp2 614 bp overlap
SMARCA4 2 datasets
ChIP 501-mel_SHSOX10 GSE61965.SMARCA4.501-mel_SHSOX10 240 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 169 bp overlap
SMARCC1 2 datasets
ChIP DE_D1 S15-DE-d1-BAF155-exp1 243 bp overlap
ChIP hiPSC GSE124903.SMARCC1.hiPSC 183 bp overlap
SOX2 1 dataset
ChIP hiPSC GSE56567.SOX2.hiPSC 358 bp overlap
SPIB 3 datasets
Motif DE_12h DE_12h-SPIB_MA0081.3 13 bp overlap
Motif DE_24h DE_24h-SPIB_MA0081.3 13 bp overlap
Motif DE_36h DE_36h-SPIB_MA0081.3 13 bp overlap
SREBF1 3 datasets
Motif DE_12h DE_12h-SREBF1_MA0595.1 10 bp overlap
Motif DE_24h DE_24h-SREBF1_MA0595.1 10 bp overlap
Motif DE_36h DE_36h-SREBF1_MA0595.1 10 bp overlap
SREBF2 3 datasets
Motif DE_12h DE_12h-SREBF2_MA0596.1 10 bp overlap
Motif DE_24h DE_24h-SREBF2_MA0596.1 10 bp overlap
Motif DE_36h DE_36h-SREBF2_MA0596.1 10 bp overlap
Spi1 3 datasets
Motif DE_12h DE_12h-Spi1_MA0080.7 13 bp overlap
Motif DE_24h DE_24h-Spi1_MA0080.7 13 bp overlap
Motif DE_36h DE_36h-Spi1_MA0080.7 13 bp overlap
Stat5a::Stat5b 3 datasets
Motif DE_12h DE_12h-Stat5aStat5b_MA0519.2 9 bp overlap
Motif DE_24h DE_24h-Stat5aStat5b_MA0519.2 9 bp overlap
Motif DE_36h DE_36h-Stat5aStat5b_MA0519.2 9 bp overlap
Stat5b 3 datasets
Motif DE_12h DE_12h-Stat5b_MA1625.2 9 bp overlap
Motif DE_24h DE_24h-Stat5b_MA1625.2 9 bp overlap
Motif DE_36h DE_36h-Stat5b_MA1625.2 9 bp overlap
TBR1 5 datasets
Motif DE_12h DE_12h-TBR1_MA0802.2 9 bp overlap
Motif DE_12h DE_12h-TBR1_MA0802.2 9 bp overlap
Motif DE_24h DE_24h-TBR1_MA0802.2 9 bp overlap
Motif DE_36h DE_36h-TBR1_MA0802.2 9 bp overlap
Motif DE_36h DE_36h-TBR1_MA0802.2 9 bp overlap
TBX1 2 datasets
Motif DE_12h DE_12h-TBX1_MA0805.1 8 bp overlap
Motif DE_36h DE_36h-TBX1_MA0805.1 8 bp overlap
TBX15 2 datasets
Motif DE_12h DE_12h-TBX15_MA0803.1 8 bp overlap
Motif DE_36h DE_36h-TBX15_MA0803.1 8 bp overlap
TBX18 2 datasets
Motif DE_12h DE_12h-TBX18_MA1565.2 9 bp overlap
Motif DE_36h DE_36h-TBX18_MA1565.2 9 bp overlap
TBX2 2 datasets
Motif DE_12h DE_12h-TBX2_MA0688.2 9 bp overlap
Motif DE_36h DE_36h-TBX2_MA0688.2 9 bp overlap
TBX20 5 datasets
Motif DE_12h DE_12h-TBX20_MA0689.1 11 bp overlap
Motif DE_12h DE_12h-TBX20_MA0689.1 11 bp overlap
Motif DE_24h DE_24h-TBX20_MA0689.1 11 bp overlap
Motif DE_36h DE_36h-TBX20_MA0689.1 11 bp overlap
Motif DE_36h DE_36h-TBX20_MA0689.1 11 bp overlap
TBX21 5 datasets
Motif DE_12h DE_12h-TBX21_MA0690.3 10 bp overlap
Motif DE_12h DE_12h-TBX21_MA0690.3 10 bp overlap
Motif DE_24h DE_24h-TBX21_MA0690.3 10 bp overlap
Motif DE_36h DE_36h-TBX21_MA0690.3 10 bp overlap
Motif DE_36h DE_36h-TBX21_MA0690.3 10 bp overlap
TBX3 5 datasets
Motif DE_12h DE_12h-TBX3_MA1566.3 9 bp overlap
Motif DE_12h DE_12h-TBX3_MA1566.3 9 bp overlap
Motif DE_24h DE_24h-TBX3_MA1566.3 9 bp overlap
Motif DE_36h DE_36h-TBX3_MA1566.3 9 bp overlap
Motif DE_36h DE_36h-TBX3_MA1566.3 9 bp overlap
TBX4 2 datasets
Motif DE_12h DE_12h-TBX4_MA0806.1 8 bp overlap
Motif DE_36h DE_36h-TBX4_MA0806.1 8 bp overlap
TCF7L1 3 datasets
Motif DE_12h DE_12h-TCF7L1_MA1421.1 12 bp overlap
Motif DE_24h DE_24h-TCF7L1_MA1421.1 12 bp overlap
Motif DE_36h DE_36h-TCF7L1_MA1421.1 12 bp overlap
TFAP2A 3 datasets
Motif DE_12h DE_12h-TFAP2A_MA0810.2 11 bp overlap
Motif DE_24h DE_24h-TFAP2A_MA0810.2 11 bp overlap
Motif DE_36h DE_36h-TFAP2A_MA0810.2 11 bp overlap
TFAP2B 3 datasets
Motif DE_12h DE_12h-TFAP2B_MA0811.2 11 bp overlap
Motif DE_24h DE_24h-TFAP2B_MA0811.2 11 bp overlap
Motif DE_36h DE_36h-TFAP2B_MA0811.2 11 bp overlap
TFAP2C 3 datasets
Motif DE_12h DE_12h-TFAP2C_MA0524.3 11 bp overlap
Motif DE_24h DE_24h-TFAP2C_MA0524.3 11 bp overlap
Motif DE_36h DE_36h-TFAP2C_MA0524.3 11 bp overlap
THAP1 3 datasets
Motif DE_12h DE_12h-THAP1_MA0597.3 8 bp overlap
Motif DE_24h DE_24h-THAP1_MA0597.3 8 bp overlap
Motif DE_36h DE_36h-THAP1_MA0597.3 8 bp overlap
TRIM28 1 dataset
ChIP WIBR3_NAIVE GSE84382.TRIM28.WIBR3_NAIVE 138 bp overlap
Tbx6 6 datasets
Motif DE_12h DE_12h-Tbx6_MA1567.3 9 bp overlap
Motif DE_12h DE_12h-Tbx6_MA1567.3 9 bp overlap
Motif DE_12h DE_12h-Tbx6_MA1567.3 9 bp overlap
Motif DE_24h DE_24h-Tbx6_MA1567.3 9 bp overlap
Motif DE_36h DE_36h-Tbx6_MA1567.3 9 bp overlap
Motif DE_36h DE_36h-Tbx6_MA1567.3 9 bp overlap
Vdr 1 dataset
Motif DE_12h DE_12h-Vdr_MA0693.4 7 bp overlap
ZBTB7C 1 dataset
Motif DE_12h DE_12h-ZBTB7C_MA0695.2 8 bp overlap
ZKSCAN3 3 datasets
Motif DE_12h DE_12h-ZKSCAN3_MA1973.2 14 bp overlap
Motif DE_24h DE_24h-ZKSCAN3_MA1973.2 14 bp overlap
Motif DE_36h DE_36h-ZKSCAN3_MA1973.2 14 bp overlap
ZNF143 1 dataset
ChIP FLP143HA_T4 GSE39263.ZNF143.FLP143HA_T4 145 bp overlap
ZNF324 3 datasets
Motif DE_12h DE_12h-ZNF324_MA1977.2 14 bp overlap
Motif DE_24h DE_24h-ZNF324_MA1977.2 14 bp overlap
Motif DE_36h DE_36h-ZNF324_MA1977.2 14 bp overlap
ZNF354C 3 datasets
Motif DE_12h DE_12h-ZNF354C_MA0130.1 6 bp overlap
Motif DE_24h DE_24h-ZNF354C_MA0130.1 6 bp overlap
Motif DE_36h DE_36h-ZNF354C_MA0130.1 6 bp overlap
ZNF454 3 datasets
Motif DE_12h DE_12h-ZNF454_MA1712.2 17 bp overlap
Motif DE_24h DE_24h-ZNF454_MA1712.2 17 bp overlap
Motif DE_36h DE_36h-ZNF454_MA1712.2 17 bp overlap
ZNF677 4 datasets
Motif DE_12h DE_12h-ZNF677_MA2101.1 12 bp overlap
Motif DE_12h DE_12h-ZNF677_MA2101.1 12 bp overlap
Motif DE_24h DE_24h-ZNF677_MA2101.1 12 bp overlap
Motif DE_36h DE_36h-ZNF677_MA2101.1 12 bp overlap
ZNF75A 3 datasets
Motif DE_12h DE_12h-ZNF75A_MA2097.1 12 bp overlap
Motif DE_24h DE_24h-ZNF75A_MA2097.1 12 bp overlap
Motif DE_36h DE_36h-ZNF75A_MA2097.1 12 bp overlap
ZNF75D 3 datasets
Motif DE_12h DE_12h-ZNF75D_MA1601.2 12 bp overlap
Motif DE_24h DE_24h-ZNF75D_MA1601.2 12 bp overlap
Motif DE_36h DE_36h-ZNF75D_MA1601.2 12 bp overlap