chr4 : 154,277,040 154,277,533
493 bp 85 TFs 0 linked genes
This 493 bp open chromatin element has no linked target genes and is bound by 85 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:154,272,040 – 154,282,533
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
85 transcription factors
Source
Cell type
ARNT::HIF1A 2 datasets
Motif DE_12h DE_12h-ARNTHIF1A_MA0259.2 5 bp overlap
Motif ES_0h ES_0h-ARNTHIF1A_MA0259.2 5 bp overlap
ASH2L 1 dataset
ChIP WA01 ENCSR850KIP.ASH2L.WA01 493 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 289 bp overlap
BARX1 2 datasets
Motif DE_12h DE_12h-BARX1_MA0875.2 6 bp overlap
Motif ES_0h ES_0h-BARX1_MA0875.2 6 bp overlap
BCL11A 2 datasets
ChIP WA01 ENCSR000BIP.BCL11A.WA01 183 bp overlap
ChIP WA01 ENCSR000BMJ.BCL11A.WA01 184 bp overlap
BSX 2 datasets
Motif DE_12h DE_12h-BSX_MA0876.2 6 bp overlap
Motif ES_0h ES_0h-BSX_MA0876.2 6 bp overlap
CDX2 2 datasets
Motif DE_12h DE_12h-CDX2_MA0465.3 8 bp overlap
Motif ES_0h ES_0h-CDX2_MA0465.3 8 bp overlap
CHD7 3 datasets
ChIP H1 ENCFF126NLU 493 bp overlap
ChIP WA01 ENCSR000AVA.CHD7.WA01 170 bp overlap
ChIP hiPSC_derived_lt-NES GSE108506.CHD7.hiPSC_derived_lt-NES 493 bp overlap
CREB1 1 dataset
ChIP WA01 ENCSR000BSN.CREB1.WA01 161 bp overlap
CTBP2 1 dataset
ChIP WA01 ENCSR000EUO.CTBP2.WA01 222 bp overlap
CTCF 67 datasets
Motif DE_12h DE_12h-CTCF_MA1929.2 31 bp overlap
Motif ES_0h ES_0h-CTCF_MA1929.2 31 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 192 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 161 bp overlap
ChIP GM23338 ENCFF531QOI 209 bp overlap
ChIP GM23338 ENCFF772DML 140 bp overlap
ChIP H1 ENCFF414GZI 217 bp overlap
ChIP H1 ENCFF764RHO 161 bp overlap
ChIP H9 ENCFF152GTF 306 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 291 bp overlap
ChIP HAP1_2kb GSE152721.CTCF.HAP1_2kb 301 bp overlap
ChIP HAP1_WT GSE126634.CTCF.HAP1_WT 321 bp overlap
ChIP HAP1_WaplKO-33_SCC4KO GSE94992.CTCF.HAP1_WaplKO-33_SCC4KO 220 bp overlap
ChIP HAP1_clone21 GSE152721.CTCF.HAP1_clone21 248 bp overlap
ChIP HAP1_clone21_CTCF-CRISPR GSE152721.CTCF.HAP1_clone21_CTCF-CRISPR 287 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 262 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_2 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_2 272 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 317 bp overlap
ChIP HAP1_clone25 GSE152721.CTCF.HAP1_clone25 290 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 241 bp overlap
ChIP HUES-64 GSE97394.CTCF.HUES-64 187 bp overlap
ChIP HUES-64_DNMT-KO GSE97394.CTCF.HUES-64_DNMT-KO 290 bp overlap
ChIP Hep-G2 ENCSR000DLS.CTCF.Hep-G2 79 bp overlap
ChIP NPC GSE115407.CTCF.NPC 293 bp overlap
ChIP SK-MEL-147 GSE94488.CTCF.SK-MEL-147 190 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 232 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 266 bp overlap
ChIP astrocyte ENCSR000AOO.CTCF.astrocyte 223 bp overlap
ChIP astrocyte of the cerebellum ENCFF511OCS 283 bp overlap
ChIP astrocyte_cerebellum ENCSR000DSZ.CTCF.astrocyte_cerebellum 291 bp overlap
ChIP astrocyte_spinal_cord ENCSR000DSU.CTCF.astrocyte_spinal_cord 132 bp overlap
ChIP brain ENCFF685VRG 366 bp overlap
ChIP endodermal cell ENCFF471YCZ 288 bp overlap
ChIP epithelial_kidney ENCSR000DVH.CTCF.epithelial_kidney 250 bp overlap
ChIP epithelial_mammary ENCSR000DUS.CTCF.epithelial_mammary 130 bp overlap
ChIP epithelial_retinal_pigment ENCSR000DVI.CTCF.epithelial_retinal_pigment 194 bp overlap
ChIP hESC GSE20650.CTCF.hESC 108 bp overlap
ChIP hESC_D0 GSE116862.CTCF.hESC_D0 227 bp overlap
ChIP hESC_D2 GSE116862.CTCF.hESC_D2 197 bp overlap
ChIP hESC_NAIVE GSE69646.CTCF.hESC_NAIVE 249 bp overlap
ChIP hiPSC GSE115407.CTCF.hiPSC 307 bp overlap
ChIP hiPSC_Angelman GSE117283.CTCF.hiPSC_Angelman 232 bp overlap
ChIP hiPSC_CVI GSE106870.CTCF.hiPSC_CVI 257 bp overlap
ChIP hiPSC_HUES9 GSE106870.CTCF.hiPSC_HUES9 120 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 227 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 188 bp overlap
ChIP hiPSC_IIA11 GSE106870.CTCF.hiPSC_IIA11 241 bp overlap
ChIP hiPSC_IIIB12 GSE106870.CTCF.hiPSC_IIIB12 130 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 217 bp overlap
ChIP hiPSC_Prader-Willi GSE117283.CTCF.hiPSC_Prader-Willi 231 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 234 bp overlap
ChIP hiPSC_parental-line GSE106870.CTCF.hiPSC_parental-line 280 bp overlap
ChIP keratinocyte ENCSR000DWX.CTCF.keratinocyte 258 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 110 bp overlap
ChIP kidney epithelial cell ENCFF173LWY 292 bp overlap
ChIP mammary epithelial cell ENCFF164SPU 255 bp overlap
ChIP mammary epithelial cell ENCFF873ERE 271 bp overlap
ChIP neural progenitor cell ENCFF420RBO 270 bp overlap
ChIP neural progenitor cell ENCFF581WPG 254 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 299 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 163 bp overlap
ChIP primary-glioblastoma_G567 GSE121601.CTCF.primary-glioblastoma_G567 144 bp overlap
ChIP primary-glioblastoma_G583 GSE121601.CTCF.primary-glioblastoma_G583 245 bp overlap
ChIP progenitor cell of endocrine pancreas ENCFF492KHV 367 bp overlap
ChIP retinal pigment epithelial cell ENCFF810AAG 257 bp overlap
ChIP smooth-muscle-cell ENCSR261VAS.CTCF.smooth-muscle-cell 167 bp overlap
ChIP type B pancreatic cell ENCFF910FNQ 231 bp overlap
CTCFL 2 datasets
Motif DE_12h DE_12h-CTCFL_MA1102.3 8 bp overlap
Motif ES_0h ES_0h-CTCFL_MA1102.3 8 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF460KDD 215 bp overlap
DLX1 2 datasets
Motif DE_12h DE_12h-DLX1_MA0879.3 6 bp overlap
Motif ES_0h ES_0h-DLX1_MA0879.3 6 bp overlap
DLX6 2 datasets
Motif DE_12h DE_12h-DLX6_MA0882.2 6 bp overlap
Motif ES_0h ES_0h-DLX6_MA0882.2 6 bp overlap
Dlx2 2 datasets
Motif DE_12h DE_12h-Dlx2_MA0885.3 8 bp overlap
Motif ES_0h ES_0h-Dlx2_MA0885.3 8 bp overlap
Dlx3 2 datasets
Motif DE_12h DE_12h-Dlx3_MA0880.2 6 bp overlap
Motif ES_0h ES_0h-Dlx3_MA0880.2 6 bp overlap
Dlx4 2 datasets
Motif DE_12h DE_12h-Dlx4_MA0881.2 6 bp overlap
Motif ES_0h ES_0h-Dlx4_MA0881.2 6 bp overlap
Dlx5 2 datasets
Motif DE_12h DE_12h-Dlx5_MA1476.3 8 bp overlap
Motif ES_0h ES_0h-Dlx5_MA1476.3 8 bp overlap
EN2 2 datasets
Motif DE_12h DE_12h-EN2_MA0642.3 7 bp overlap
Motif ES_0h ES_0h-EN2_MA0642.3 7 bp overlap
EP300 2 datasets
ChIP H1 ENCFF927IYK 297 bp overlap
ChIP WA01 ENCSR000BKK.EP300.WA01 221 bp overlap
FOXP1 1 dataset
ChIP H9 GSE31006.FOXP1.H9 203 bp overlap
GBX1 2 datasets
Motif DE_12h DE_12h-GBX1_MA0889.2 7 bp overlap
Motif ES_0h ES_0h-GBX1_MA0889.2 7 bp overlap
GBX2 2 datasets
Motif DE_12h DE_12h-GBX2_MA0890.2 6 bp overlap
Motif ES_0h ES_0h-GBX2_MA0890.2 6 bp overlap
HDAC2 1 dataset
ChIP WA01 ENCSR000BNR.HDAC2.WA01 193 bp overlap
HESX1 2 datasets
Motif DE_12h DE_12h-HESX1_MA0894.2 6 bp overlap
Motif ES_0h ES_0h-HESX1_MA0894.2 6 bp overlap
HOXA7 2 datasets
Motif DE_12h DE_12h-HOXA7_MA1498.3 6 bp overlap
Motif ES_0h ES_0h-HOXA7_MA1498.3 6 bp overlap
IRF3 1 dataset
Motif ES_0h ES_0h-IRF3_MA1418.2 17 bp overlap
JUN 1 dataset
ChIP HUES-8 GSE109524.JUN.HUES-8 310 bp overlap
JUND 2 datasets
ChIP H1 ENCFF010YXS 311 bp overlap
ChIP WA01 ENCSR000BKP.JUND.WA01 128 bp overlap
LBX1 2 datasets
Motif DE_12h DE_12h-LBX1_MA0618.2 7 bp overlap
Motif ES_0h ES_0h-LBX1_MA0618.2 7 bp overlap
LBX2 2 datasets
Motif DE_12h DE_12h-LBX2_MA0699.2 6 bp overlap
Motif ES_0h ES_0h-LBX2_MA0699.2 6 bp overlap
LHX2 2 datasets
Motif DE_12h DE_12h-LHX2_MA0700.3 6 bp overlap
Motif ES_0h ES_0h-LHX2_MA0700.3 6 bp overlap
LHX9 2 datasets
Motif DE_12h DE_12h-LHX9_MA0701.3 7 bp overlap
Motif ES_0h ES_0h-LHX9_MA0701.3 7 bp overlap
MAX 1 dataset
ChIP WA01 ENCSR000EUP.MAX.WA01 200 bp overlap
MSX1 2 datasets
Motif DE_12h DE_12h-MSX1_MA0666.3 6 bp overlap
Motif ES_0h ES_0h-MSX1_MA0666.3 6 bp overlap
MSX2 2 datasets
Motif DE_12h DE_12h-MSX2_MA0708.3 6 bp overlap
Motif ES_0h ES_0h-MSX2_MA0708.3 6 bp overlap
Msx3 2 datasets
Motif DE_12h DE_12h-Msx3_MA0709.2 6 bp overlap
Motif ES_0h ES_0h-Msx3_MA0709.2 6 bp overlap
NANOG 10 datasets
ChIP GM23338 ENCFF065NZG 311 bp overlap
ChIP GM23338 ENCSR061DGF.NANOG.GM23338 233 bp overlap
ChIP H1 ENCFF747ZPQ 241 bp overlap
ChIP HUES-8 GSE109524.NANOG.HUES-8 485 bp overlap
ChIP WA01 ERP004238.NANOG.WA01 334 bp overlap
ChIP WA01 ENCSR000BMT.NANOG.WA01 231 bp overlap
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 330 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 430 bp overlap
ChIP WA09_heat-shock GSE105028.NANOG.WA09_heat-shock 377 bp overlap
ChIP hESC GSE18292.NANOG.hESC 177 bp overlap
NFATC3 1 dataset
Motif ES_0h ES_0h-NFATC3_MA0625.3 6 bp overlap
NFATC4 1 dataset
Motif ES_0h ES_0h-NFATC4_MA1525.3 9 bp overlap
Nfat5 1 dataset
Motif ES_0h ES_0h-Nfat5_MA0606.3 8 bp overlap
Nfatc1 1 dataset
Motif ES_0h ES_0h-Nfatc1_MA0624.3 6 bp overlap
Nfatc2 1 dataset
Motif ES_0h ES_0h-Nfatc2_MA0152.3 8 bp overlap
Nobox 2 datasets
Motif DE_12h DE_12h-Nobox_MA0125.2 6 bp overlap
Motif ES_0h ES_0h-Nobox_MA0125.2 6 bp overlap
POU5F1 3 datasets
ChIP HUES-8 GSE109524.POU5F1.HUES-8 434 bp overlap
ChIP hESC_PRIMED GSE69646.POU5F1.hESC_PRIMED 384 bp overlap
ChIP hiPSC GSE56567.POU5F1.hiPSC 148 bp overlap
PRDM1 1 dataset
ChIP fetal_testis GSE100639.PRDM1.fetal_testis 159 bp overlap
PRRX2 2 datasets
Motif DE_12h DE_12h-PRRX2_MA0075.4 7 bp overlap
Motif ES_0h ES_0h-PRRX2_MA0075.4 7 bp overlap
RAD21 20 datasets
ChIP H1 ENCFF698EWO 224 bp overlap
ChIP H1 ENCFF967OJF 124 bp overlap
ChIP HAP1 GSE152721.RAD21.HAP1 332 bp overlap
ChIP HAP1_CTCF GSE126634.RAD21.HAP1_CTCF 324 bp overlap
ChIP HAP1_clone25 GSE152721.RAD21.HAP1_clone25 277 bp overlap
ChIP HCT-116 GSE121355.RAD21.HCT-116 269 bp overlap
ChIP Ishikawa ENCFF570JVV 225 bp overlap
ChIP Ishikawa ENCSR000BTU.RAD21.Ishikawa 200 bp overlap
ChIP K-562 ENCSR000BKV.RAD21.K-562 142 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-020m-Flavo-000m 160 bp overlap
ChIP WA09 GSE105028.RAD21.WA09 267 bp overlap
ChIP WA09_heat-shock GSE105028.RAD21.WA09_heat-shock 276 bp overlap
ChIP hiPSC_CVI GSE106870.RAD21.hiPSC_CVI 265 bp overlap
ChIP hiPSC_HUES9 GSE106870.RAD21.hiPSC_HUES9 174 bp overlap
ChIP hiPSC_IB12 GSE106870.RAD21.hiPSC_IB12 240 bp overlap
ChIP hiPSC_IIA12 GSE106870.RAD21.hiPSC_IIA12 197 bp overlap
ChIP hiPSC_IIIB12 GSE106870.RAD21.hiPSC_IIIB12 254 bp overlap
ChIP hiPSC_IV8 GSE106870.RAD21.hiPSC_IV8 253 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.RAD21.hiPSC_failed-genome-editing 278 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 295 bp overlap
RARA 1 dataset
ChIP hiPSC_D4 GSE109172.RARA.hiPSC_D4 309 bp overlap
RAX 2 datasets
Motif DE_12h DE_12h-RAX_MA0718.2 6 bp overlap
Motif ES_0h ES_0h-RAX_MA0718.2 6 bp overlap
RUNX1 1 dataset
ChIP keratinocyte GSE98483.RUNX1.keratinocyte 210 bp overlap
SCRT1 2 datasets
Motif DE_12h DE_12h-SCRT1_MA0743.3 10 bp overlap
Motif ES_0h ES_0h-SCRT1_MA0743.3 10 bp overlap
SCRT2 2 datasets
Motif DE_12h DE_12h-SCRT2_MA0744.3 10 bp overlap
Motif ES_0h ES_0h-SCRT2_MA0744.3 10 bp overlap
SMAD1 1 dataset
ChIP BG03 GSE36578.SMAD1.BG03 165 bp overlap
SMAD3 1 dataset
ChIP BG03 GSE36578.SMAD3.BG03 121 bp overlap
SMARCA4 4 datasets
ChIP NPC_siCT GSE122631.SMARCA4.NPC_siCT 398 bp overlap
ChIP NPC_siSOX GSE122631.SMARCA4.NPC_siSOX 350 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 216 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 224 bp overlap
SMARCB1 1 dataset
ChIP hiPSC_WT GSE124903.SMARCB1.hiPSC_WT 267 bp overlap
SMARCC1 2 datasets
ChIP hiPSC GSE124903.SMARCC1.hiPSC 157 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 178 bp overlap
SOX10 2 datasets
Motif DE_12h DE_12h-SOX10_MA0442.3 6 bp overlap
Motif ES_0h ES_0h-SOX10_MA0442.3 6 bp overlap
SOX2 8 datasets
ChIP H9 GSE46837.SOX2.H9 224 bp overlap
ChIP HNSC GSE69479.SOX2.HNSC 493 bp overlap
ChIP LK2 GSE137459.SOX2.LK2 368 bp overlap
ChIP LK2_DNp63 GSE137459.SOX2.LK2_DNp63 284 bp overlap
ChIP LK2_GFP GSE137459.SOX2.LK2_GFP 400 bp overlap
ChIP hESC GSE69479.SOX2.hESC 225 bp overlap
ChIP hESC GSE18292.SOX2.hESC 95 bp overlap
ChIP hiPSC GSE56567.SOX2.hiPSC 322 bp overlap
SP1 2 datasets
ChIP H1 ENCFF263FUH 321 bp overlap
ChIP WA01 ENCSR000BIR.SP1.WA01 341 bp overlap
SP4 1 dataset
ChIP WA01 ENCSR000BQV.SP4.WA01 266 bp overlap
STAT3 1 dataset
ChIP MDA-MB-157_EtOH GSE85579.STAT3.MDA-MB-157_EtOH 280 bp overlap
Sox11 2 datasets
Motif DE_12h DE_12h-Sox11_MA0869.3 8 bp overlap
Motif ES_0h ES_0h-Sox11_MA0869.3 8 bp overlap
Sox17 2 datasets
Motif DE_12h DE_12h-Sox17_MA0078.3 10 bp overlap
Motif ES_0h ES_0h-Sox17_MA0078.3 10 bp overlap
Sox6 2 datasets
Motif DE_12h DE_12h-Sox6_MA0515.1 10 bp overlap
Motif ES_0h ES_0h-Sox6_MA0515.1 10 bp overlap
Sox7 2 datasets
Motif DE_12h DE_12h-Sox7_MA2095.1 10 bp overlap
Motif ES_0h ES_0h-Sox7_MA2095.1 10 bp overlap
Stat2 3 datasets
Motif DE_12h DE_12h-Stat2_MA1623.2 10 bp overlap
Motif ES_0h ES_0h-Stat2_MA1623.2 10 bp overlap
Motif ES_0h ES_0h-Stat2_MA1623.2 10 bp overlap
TCF12 1 dataset
ChIP WA01 ENCSR000BIT.TCF12.WA01 186 bp overlap
TCF3 1 dataset
ChIP NPC GSE154479.TCF3.NPC 220 bp overlap
TEAD1 2 datasets
ChIP WTC11 ENCFF502QUV 405 bp overlap
ChIP pancreas_12D ERP008682.TEAD1.pancreas_12D 165 bp overlap
TEAD4 5 datasets
ChIP ESC S26-ESC-d0-TEAD4-exp1 439 bp overlap
ChIP H1 ENCFF778PAX 201 bp overlap
ChIP Ishikawa ENCSR000BSW.TEAD4.Ishikawa 175 bp overlap
ChIP WTC11 ENCFF114TZS 341 bp overlap
ChIP hESC GSE99202.TEAD4.hESC 282 bp overlap
TP53 1 dataset
ChIP hESC_DIFF GSE39912.TP53.hESC_DIFF 264 bp overlap
TRIM28 1 dataset
ChIP hESC GSE115387.TRIM28.hESC 244 bp overlap
Thap11 2 datasets
Motif DE_12h DE_12h-Thap11_MA1573.2 14 bp overlap
Motif ES_0h ES_0h-Thap11_MA1573.2 14 bp overlap
YY1 2 datasets
ChIP H1 ENCFF524BTL 322 bp overlap
ChIP WA01 ENCSR000BKD.YY1.WA01 240 bp overlap
ZIM3 2 datasets
Motif DE_12h DE_12h-ZIM3_MA1709.2 11 bp overlap
Motif ES_0h ES_0h-ZIM3_MA1709.2 11 bp overlap
ZNF317 2 datasets
Motif DE_12h DE_12h-ZNF317_MA1593.2 8 bp overlap
Motif ES_0h ES_0h-ZNF317_MA1593.2 8 bp overlap
ZNF423 1 dataset
ChIP WTC11 ENCFF574PBR 317 bp overlap
ZNF462 2 datasets
ChIP GM23338 ENCFF896CCA 251 bp overlap
ChIP GM23338 ENCSR334UWP.ZNF462.GM23338 268 bp overlap
ZNF766 2 datasets
Motif DE_12h DE_12h-ZNF766_MA2098.1 9 bp overlap
Motif ES_0h ES_0h-ZNF766_MA2098.1 9 bp overlap
ZNF768 2 datasets
Motif DE_12h DE_12h-ZNF768_MA1731.2 9 bp overlap
Motif ES_0h ES_0h-ZNF768_MA1731.2 9 bp overlap
ZNF8 2 datasets
Motif DE_12h DE_12h-ZNF8_MA1718.1 20 bp overlap
Motif ES_0h ES_0h-ZNF8_MA1718.1 20 bp overlap