chr4 : 13,127,433 13,128,022
589 bp 47 TFs 0 linked genes
This 589 bp open chromatin element has no linked target genes and is bound by 47 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr4:13,122,433 – 13,133,022
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
47 transcription factors
Source
Cell type
AR 22 datasets
ChIP LNCaP GSE110655.AR.LNCaP 151 bp overlap
ChIP LNCaP-clone-FGC_DHT-ABBV-744 GSE118247.AR.LNCaP-clone-FGC_DHT-ABBV-744 208 bp overlap
ChIP LNCaP-clone-FGC_DHT-DMSO GSE118247.AR.LNCaP-clone-FGC_DHT-DMSO 214 bp overlap
ChIP LNCaP_DSG GSE114737.AR.LNCaP_DSG 167 bp overlap
ChIP LNCaP_FOXA1 ERP003503.AR.LNCaP_FOXA1 164 bp overlap
ChIP LNCaP_SHGATA2_R1881 GSE69043.AR.LNCaP_SHGATA2_R1881 113 bp overlap
ChIP LNCaP_Talen_DHT GSE89938.AR.LNCaP_Talen_DHT 199 bp overlap
ChIP LNCaP_androgen-N_hypoxia-Y GSE114732.AR.LNCaP_androgen-N_hypoxia-Y 156 bp overlap
ChIP LNCaP_r1881 GSE94682.AR.LNCaP_r1881 371 bp overlap
ChIP LNCaP_sgCHD1 GSE117430.AR.LNCaP_sgCHD1 240 bp overlap
ChIP VCaP_DHTTHZ1 GSE125245.AR.VCaP_DHTTHZ1 142 bp overlap
ChIP VCaP_R1881_30M GSE84432.AR.VCaP_R1881_30M 175 bp overlap
ChIP prostate-cancer_PDX_167 GSE130408.AR.prostate-cancer_PDX_167 107 bp overlap
ChIP prostate-cancer_PDX_189-4 GSE130408.AR.prostate-cancer_PDX_189-4 67 bp overlap
ChIP prostate-cancer_PDX_35 GSE130408.AR.prostate-cancer_PDX_35 277 bp overlap
ChIP prostate-cancer_PDX_58 GSE130408.AR.prostate-cancer_PDX_58 67 bp overlap
ChIP prostate-cancer_PDX_70 GSE130408.AR.prostate-cancer_PDX_70 173 bp overlap
ChIP prostate-cancer_PDX_81 GSE130408.AR.prostate-cancer_PDX_81 86 bp overlap
ChIP prostate-cancer_PDX_92 GSE130408.AR.prostate-cancer_PDX_92 81 bp overlap
ChIP prostate_1816_T GSE130408.AR.prostate_1816_T 200 bp overlap
ChIP prostate_4024_T GSE130408.AR.prostate_4024_T 225 bp overlap
ChIP prostate_P23_T GSE130408.AR.prostate_P23_T 220 bp overlap
ARID1A 1 dataset
ChIP LNCaP_r1881 GSE94682.ARID1A.LNCaP_r1881 213 bp overlap
BRD3 1 dataset
ChIP HEK293T GSE39579.BRD3.HEK293T 208 bp overlap
BRD4 3 datasets
ChIP CHL-1 GSE95585.BRD4.CHL-1 137 bp overlap
ChIP K-562_iBET-IFNG GSE138084.BRD4.K-562_iBET-IFNG 180 bp overlap
ChIP LNCaP-C4-2_F133V GSE88871.BRD4.LNCaP-C4-2_F133V 199 bp overlap
CDX2 1 dataset
ChIP LS180_125 GSE31939.CDX2.LS180_125 126 bp overlap
CREB5 1 dataset
ChIP LNCaP GSE137775.CREB5.LNCaP 187 bp overlap
CTCF 35 datasets
ChIP 786-O_NORMOXIA GSE78113.CTCF.786-O_NORMOXIA 259 bp overlap
ChIP A-549 ENCSR000DPF.CTCF.A-549 138 bp overlap
ChIP D54 ENCSR000DKN.CTCF.D54 174 bp overlap
ChIP FT282 GSE131931.CTCF.FT282 206 bp overlap
ChIP FT282_EV GSE131931.CTCF.FT282_EV 180 bp overlap
ChIP GM23338 ENCFF772DML 201 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 187 bp overlap
ChIP HAP1_clone21_CTCF-CRISPR GSE152721.CTCF.HAP1_clone21_CTCF-CRISPR 214 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 229 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 233 bp overlap
ChIP Hep-G2 ENCSR000BIE.CTCF.Hep-G2 108 bp overlap
ChIP Hep-G2 ENCSR000DLS.CTCF.Hep-G2 125 bp overlap
ChIP HepG2 ENCFF757EKU 351 bp overlap
ChIP K-562 ENCSR000DMA.CTCF.K-562 106 bp overlap
ChIP K-562_15776 GSE131349.CTCF.K-562_15776 124 bp overlap
ChIP K-562_MonoC GSE137216.CTCF.K-562_MonoC 251 bp overlap
ChIP K562 ENCFF082GOI 185 bp overlap
ChIP KARPAS422 ENCSR113REG.CTCF.KARPAS422 211 bp overlap
ChIP Kelly_resistant GSE103030.CTCF.Kelly_resistant 154 bp overlap
ChIP Kelly_sensitive GSE103030.CTCF.Kelly_sensitive 138 bp overlap
ChIP MDA-MB-435_Asynchronous GSE102019.CTCF.MDA-MB-435_Asynchronous 231 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 155 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.CTCF.THP-1_Pam3csk-150m-Flavo-030m 162 bp overlap
ChIP THP-1_Pam3csk-360m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-360m-Flavo-240m 160 bp overlap
ChIP brain ENCFF163BBN 544 bp overlap
ChIP hESC_D80 GSE116862.CTCF.hESC_D80 224 bp overlap
ChIP heart_left-ventricle ENCSR791AYW.CTCF.heart_left-ventricle 228 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 126 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 181 bp overlap
ChIP islet ERP004003.CTCF.islet 201 bp overlap
ChIP retinal pigment epithelial cell ENCFF810AAG 257 bp overlap
ChIP thyroid gland ENCFF631QRY 457 bp overlap
ChIP thyroid-gland ENCSR492ZIW.CTCF.thyroid-gland 437 bp overlap
ChIP thyroid-gland ENCSR033KMZ.CTCF.thyroid-gland 263 bp overlap
ChIP thyroid-gland ENCSR145HBC.CTCF.thyroid-gland 238 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF364PUR 201 bp overlap
ESR1 2 datasets
ChIP MCF-7_E2+4OHT_SRC-3 GSE119702.ESR1.MCF-7_E2+4OHT_SRC-3 169 bp overlap
ChIP MCF-7_SRC3_OHT GSE119702.ESR1.MCF-7_SRC3_OHT 169 bp overlap
FEZF1 2 datasets
ChIP HEK293 ENCFF528YED 390 bp overlap
ChIP HEK293 ENCSR827NWO.FEZF1.HEK293 291 bp overlap
FOXA1 71 datasets
ChIP 22Rv1 GSE96652.FOXA1.22Rv1 232 bp overlap
ChIP 22Rv1_CST GSE129951.FOXA1.22Rv1_CST 379 bp overlap
ChIP 22Rv1_CST GSE123618.FOXA1.22Rv1_CST 175 bp overlap
ChIP 22Rv1_CST_Crispr GSE123618.FOXA1.22Rv1_CST_Crispr 195 bp overlap
ChIP 22Rv1_CST_Crispr-36 GSE123618.FOXA1.22Rv1_CST_Crispr-36 198 bp overlap
ChIP 22Rv1_CST_Crispr-70 GSE123618.FOXA1.22Rv1_CST_Crispr-70 202 bp overlap
ChIP 22Rv1_CST_Crispr_WT3 GSE123618.FOXA1.22Rv1_CST_Crispr_WT3 221 bp overlap
ChIP 22Rv1_Dox GSE85558.FOXA1.22Rv1_Dox 214 bp overlap
ChIP 22Rv1_EtOH GSE80742.FOXA1.22Rv1_EtOH 259 bp overlap
ChIP 22Rv1_R1881 GSE80742.FOXA1.22Rv1_R1881 267 bp overlap
ChIP 22Rv1_TFS_Crispr_WT3 GSE123618.FOXA1.22Rv1_TFS_Crispr_WT3 187 bp overlap
ChIP 22Rv1_i176m_CST GSE123618.FOXA1.22Rv1_i176m_CST 198 bp overlap
ChIP 22Rv1_r261g_CST GSE123618.FOXA1.22Rv1_r261g_CST 240 bp overlap
ChIP HEK293_eGFP_TFS GSE123618.FOXA1.HEK293_eGFP_TFS 206 bp overlap
ChIP HEK293_i176m_TFS GSE123618.FOXA1.HEK293_i176m_TFS 177 bp overlap
ChIP HEK293_r261g_TFS GSE123618.FOXA1.HEK293_r261g_TFS 213 bp overlap
ChIP HepG2 ENCFF207NVJ 203 bp overlap
ChIP LAPC-4_CST_DHT GSE123618.FOXA1.LAPC-4_CST_DHT 184 bp overlap
ChIP LAPC-4_TFS_p358fs-V5 GSE123618.FOXA1.LAPC-4_TFS_p358fs-V5 190 bp overlap
ChIP LNCaP GSE64656.FOXA1.LNCaP 221 bp overlap
ChIP LNCaP GSE52725.FOXA1.LNCaP 169 bp overlap
ChIP LNCaP-C4-2B GSE40050.FOXA1.LNCaP-C4-2B 212 bp overlap
ChIP LNCaP-C4-2B_CST GSE123618.FOXA1.LNCaP-C4-2B_CST 220 bp overlap
ChIP LNCaP-C4-2B_DHT GSE40050.FOXA1.LNCaP-C4-2B_DHT 237 bp overlap
ChIP LNCaP-C4-2B_TFS GSE123618.FOXA1.LNCaP-C4-2B_TFS 180 bp overlap
ChIP LNCaP_1F5 GSE30623.FOXA1.LNCaP_1F5 250 bp overlap
ChIP LNCaP_1F5_SIFOXA1 GSE30623.FOXA1.LNCaP_1F5_SIFOXA1 129 bp overlap
ChIP LNCaP_DHT GSE28264.FOXA1.LNCaP_DHT 158 bp overlap
ChIP LNCaP_DMSO GSE114274.FOXA1.LNCaP_DMSO 190 bp overlap
ChIP LNCaP_DSG GSE114737.FOXA1.LNCaP_DSG 229 bp overlap
ChIP LNCaP_ETOH24H GSE58428.FOXA1.LNCaP_ETOH24H 297 bp overlap
ChIP LNCaP_F266S_shFOXA1 GSE128883.FOXA1.LNCaP_F266S_shFOXA1 185 bp overlap
ChIP LNCaP_FA GSE114737.FOXA1.LNCaP_FA 176 bp overlap
ChIP LNCaP_G87R_shFOXA1 GSE128883.FOXA1.LNCaP_G87R_shFOXA1 261 bp overlap
ChIP LNCaP_SHFOXA1_R1881 GSE37345.FOXA1.LNCaP_SHFOXA1_R1881 196 bp overlap
ChIP LNCaP_TFS GSE123618.FOXA1.LNCaP_TFS 186 bp overlap
ChIP LNCaP_UPF1069 GSE114274.FOXA1.LNCaP_UPF1069 169 bp overlap
ChIP LNCaP_VEH GSE114266.FOXA1.LNCaP_VEH 231 bp overlap
ChIP LS180 GSE140533.FOXA1.LS180 351 bp overlap
ChIP MCF-7 ERP000380.FOXA1.MCF-7 168 bp overlap
ChIP MCF-7 ERP001226.FOXA1.MCF-7 138 bp overlap
ChIP MCF-7-TAMR-1_4OH-Tam GSE75201.FOXA1.MCF-7-TAMR-1_4OH-Tam 102 bp overlap
ChIP MCF-7_4OH-Tam GSE75201.FOXA1.MCF-7_4OH-Tam 148 bp overlap
ChIP MCF-7_E2 GSE23852.FOXA1.MCF-7_E2 107 bp overlap
ChIP MCF-7_E2 GSE59530.FOXA1.MCF-7_E2 153 bp overlap
ChIP MCF-7_E2_TNF GSE59530.FOXA1.MCF-7_E2_TNF 183 bp overlap
ChIP MCF-7_ETOH GSE23852.FOXA1.MCF-7_ETOH 120 bp overlap
ChIP MCF-7_JC4693 GSE126004.FOXA1.MCF-7_JC4693 202 bp overlap
ChIP MCF-7_TNF GSE59530.FOXA1.MCF-7_TNF 137 bp overlap
ChIP MCF-7_estrogen_ab2 GSE112969.FOXA1.MCF-7_estrogen_ab2 202 bp overlap
ChIP MCF-7_vehicle_ab1 GSE112969.FOXA1.MCF-7_vehicle_ab1 238 bp overlap
ChIP MCF-7_vehicle_ab2 GSE112969.FOXA1.MCF-7_vehicle_ab2 207 bp overlap
ChIP T-47D_shRNF2_8h GSE137579.FOXA1.T-47D_shRNF2_8h 134 bp overlap
ChIP VCaP_ETOH24H GSE58428.FOXA1.VCaP_ETOH24H 151 bp overlap
ChIP ZR751 GSE72249.FOXA1.ZR751 217 bp overlap
ChIP ZR751_DEX GSE72249.FOXA1.ZR751_DEX 128 bp overlap
ChIP ZR751_E2 GSE72249.FOXA1.ZR751_E2 202 bp overlap
ChIP ZR751_E2_TAM ERP000380.FOXA1.ZR751_E2_TAM 146 bp overlap
ChIP breast-cancer_ENOB-2849 GSE128018.FOXA1.breast-cancer_ENOB-2849 201 bp overlap
ChIP breast-cancer_Veh-2856 GSE128018.FOXA1.breast-cancer_Veh-2856 242 bp overlap
ChIP breast-cancer_Veh-2860 GSE128018.FOXA1.breast-cancer_Veh-2860 437 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA1.pancreatic-progenitor_PP1 290 bp overlap
ChIP primary-prostate-cancer_G2_DSG GSE114737.FOXA1.primary-prostate-cancer_G2_DSG 321 bp overlap
ChIP primary-prostate-cancer_P2_DSG GSE114737.FOXA1.primary-prostate-cancer_P2_DSG 376 bp overlap
ChIP prostate-cancer_PDX_189-3 GSE130408.FOXA1.prostate-cancer_PDX_189-3 257 bp overlap
ChIP prostate_2030_T GSE130408.FOXA1.prostate_2030_T 173 bp overlap
ChIP prostate_2078_T GSE130408.FOXA1.prostate_2078_T 143 bp overlap
ChIP prostate_2483_T GSE130408.FOXA1.prostate_2483_T 197 bp overlap
ChIP prostate_P23_T GSE130408.FOXA1.prostate_P23_T 205 bp overlap
ChIP prostate_P27_T GSE130408.FOXA1.prostate_P27_T 164 bp overlap
ChIP prostate_P5_T GSE130408.FOXA1.prostate_P5_T 309 bp overlap
FOXA2 5 datasets
ChIP Caco-2 GSE66218.FOXA2.Caco-2 133 bp overlap
ChIP DE DE-FOXA2-1 389 bp overlap
ChIP DE DE-FOXA2-2 366 bp overlap
ChIP HepG2 ENCFF533COJ 212 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.FOXA2.pancreatic-progenitor_PP1 281 bp overlap
GABPA 1 dataset
ChIP VCaP_ETOH GSE49091.GABPA.VCaP_ETOH 146 bp overlap
GATA2 3 datasets
ChIP LNCaP GSE38391.GATA2.LNCaP 217 bp overlap
ChIP LNCaP GSE52725.GATA2.LNCaP 217 bp overlap
ChIP LNCaP_CSFCS GSE69043.GATA2.LNCaP_CSFCS 156 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 448 bp overlap
ChIP DE DE-GATA4-2 257 bp overlap
ChIP foregut GSE117136.GATA4.foregut 290 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 427 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA4.pancreatic-progenitor_PP1 366 bp overlap
GATA6 11 datasets
ChIP AGS GSE51705.GATA6.AGS 208 bp overlap
ChIP AGS GSE51936.GATA6.AGS 79 bp overlap
ChIP DE DE-GATA6-1 400 bp overlap
ChIP DE DE-GATA6-2 469 bp overlap
ChIP PATU8988 GSE47535.GATA6.PATU8988 292 bp overlap
ChIP YCC-3 GSE51705.GATA6.YCC-3 130 bp overlap
ChIP foregut GSE117136.GATA6.foregut 278 bp overlap
ChIP foregut_KO GSE117136.GATA6.foregut_KO 288 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA6.pancreatic-progenitor 272 bp overlap
ChIP pancreatic-progenitor_KO GSE117136.GATA6.pancreatic-progenitor_KO 236 bp overlap
ChIP pancreatic-progenitor_PP1 GSE104840.GATA6.pancreatic-progenitor_PP1 331 bp overlap
GLIS1 1 dataset
ChIP HEK293 ENCSR482BBZ.GLIS1.HEK293 221 bp overlap
HOXB13 9 datasets
ChIP prostate-cancer_PDX_167 GSE130408.HOXB13.prostate-cancer_PDX_167 105 bp overlap
ChIP prostate-cancer_PDX_189-4 GSE130408.HOXB13.prostate-cancer_PDX_189-4 135 bp overlap
ChIP prostate-cancer_PDX_70 GSE130408.HOXB13.prostate-cancer_PDX_70 174 bp overlap
ChIP prostate_2480_T GSE130408.HOXB13.prostate_2480_T 261 bp overlap
ChIP prostate_2483_T GSE130408.HOXB13.prostate_2483_T 210 bp overlap
ChIP prostate_P13_T GSE130408.HOXB13.prostate_P13_T 177 bp overlap
ChIP prostate_P19_T GSE130408.HOXB13.prostate_P19_T 253 bp overlap
ChIP prostate_P29_T GSE130408.HOXB13.prostate_P29_T 194 bp overlap
ChIP prostate_P5_T GSE130408.HOXB13.prostate_P5_T 276 bp overlap
KDM1A 1 dataset
ChIP pancreatic-progenitor_PP1 GSE104840.KDM1A.pancreatic-progenitor_PP1 207 bp overlap
MAFB 1 dataset
ChIP islet ERP004003.MAFB.islet 318 bp overlap
MYCN 1 dataset
ChIP LNCaP_DHT GSE117304.MYCN.LNCaP_DHT 210 bp overlap
MZF1 1 dataset
ChIP HEK293 ENCSR298QUH.MZF1.HEK293 293 bp overlap
NCOA2 1 dataset
ChIP MCF-7 ERP000901.NCOA2.MCF-7 133 bp overlap
NKX3-1 1 dataset
ChIP islet ERP004003.NKX3-1.islet 166 bp overlap
NR3C1 1 dataset
ChIP breast_tumor_Male_15 GSE104399.NR3C1.breast_tumor_Male_15 233 bp overlap
OSR2 3 datasets
ChIP HEK293 ENCFF875BDB 365 bp overlap
ChIP HEK293 ENCSR324LTM.OSR2.HEK293 261 bp overlap
ChIP HEK293 GSE76494.OSR2.HEK293 218 bp overlap
POU2F1 2 datasets
ChIP 22Rv1_DHT GSE123565.POU2F1.22Rv1_DHT 236 bp overlap
ChIP 22Rv1_Veh GSE123565.POU2F1.22Rv1_Veh 247 bp overlap
POU2F3 1 dataset
ChIP NCI-H1048 GSE115123.POU2F3.NCI-H1048 180 bp overlap
PROX1 1 dataset
ChIP HepG2 ENCFF016ZJS 368 bp overlap
RAD21 11 datasets
ChIP H1 ENCFF698EWO 241 bp overlap
ChIP H1 ENCFF967OJF 245 bp overlap
ChIP HAP1 GSE152721.RAD21.HAP1 192 bp overlap
ChIP HEK293_siCtrl GSE130135.RAD21.HEK293_siCtrl 456 bp overlap
ChIP Hep-G2 ENCSR054FKH.RAD21.Hep-G2 135 bp overlap
ChIP HepG2 ENCFF906QIS 225 bp overlap
ChIP MCF-7 ERP000209.RAD21.MCF-7 156 bp overlap
ChIP RH4 GSE83726.RAD21.RH4 168 bp overlap
ChIP SK-N-SH ENCSR000EHX.RAD21.SK-N-SH 131 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-000m-Flavo-000m 178 bp overlap
ChIP THP-1_Pam3csk-360m-Flavo-240m GSE103477.RAD21.THP-1_Pam3csk-360m-Flavo-240m 232 bp overlap
RELA 1 dataset
ChIP LNCaP_SIFOXA1_TNFA GSE83860.RELA.LNCaP_SIFOXA1_TNFA 139 bp overlap
RXR 1 dataset
ChIP LS180 GSE31939.RXR.LS180 112 bp overlap
SMARCA4 2 datasets
ChIP 22Rv1_PTEN-KO GSE115615.SMARCA4.22Rv1_PTEN-KO 206 bp overlap
ChIP HS-SY-2_HA-tagged GSE108926.SMARCA4.HS-SY-2_HA-tagged 589 bp overlap
SNAI2 1 dataset
ChIP SMS-CTR_T48 GSE137168.SNAI2.SMS-CTR_T48 191 bp overlap
SPI1 1 dataset
ChIP K-562 GSE70482.SPI1.K-562 151 bp overlap
STAG1 4 datasets
ChIP Hep-G2 ERP000209.STAG1.Hep-G2 175 bp overlap
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 230 bp overlap
ChIP MCF-7 ERP000209.STAG1.MCF-7 142 bp overlap
ChIP MCF-7_E2 ERP000209.STAG1.MCF-7_E2 158 bp overlap
TLE3 2 datasets
ChIP LNCaP GSE94682.TLE3.LNCaP 208 bp overlap
ChIP LNCaP_r1881 GSE94682.TLE3.LNCaP_r1881 218 bp overlap
TRIM28 2 datasets
ChIP HEK293 ENCSR618HNF.TRIM28.HEK293 376 bp overlap
ChIP U2OS ENCSR000EYC.TRIM28.U2OS 376 bp overlap
XBP1 1 dataset
ChIP LNCaP_Tg GSE121880.XBP1.LNCaP_Tg 222 bp overlap
ZMYM3 1 dataset
ChIP K562 ENCFF361LXT 95 bp overlap
ZNF654 2 datasets
ChIP HEK293 ENCFF636WIC 371 bp overlap
ChIP HEK293 ENCSR504VDV.ZNF654.HEK293 232 bp overlap
ZNF770 1 dataset
ChIP HEK293 GSE76494.ZNF770.HEK293 222 bp overlap
ZXDB 1 dataset
ChIP HEK293 ENCSR559IOZ.ZXDB.HEK293 331 bp overlap