chr2 : 156,986,409 156,986,626
217 bp 43 TFs 0 linked genes
This 217 bp open chromatin element has no linked target genes and is bound by 43 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr2:156,981,409 – 156,991,626
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
43 transcription factors
Source
Cell type
AR 4 datasets
ChIP LAPC-4_R1881 GSE148358.AR.LAPC-4_R1881 118 bp overlap
ChIP prostate-cancer_PDX_35 GSE130408.AR.prostate-cancer_PDX_35 63 bp overlap
ChIP prostate-cancer_PDX_58 GSE130408.AR.prostate-cancer_PDX_58 75 bp overlap
ChIP prostate-cancer_PDX_78 GSE130408.AR.prostate-cancer_PDX_78 73 bp overlap
BRD4 1 dataset
ChIP hESC GSE33281.BRD4.hESC 99 bp overlap
CBX3 3 datasets
ChIP HCT-116 ENCSR000BUH.CBX3.HCT-116 128 bp overlap
ChIP HCT116 ENCFF947BOL 217 bp overlap
ChIP HCT116 ENCFF947BOL 79 bp overlap
CDX2 2 datasets
ChIP LS180 GSE31939.CDX2.LS180 162 bp overlap
ChIP LS180_125 GSE31939.CDX2.LS180_125 116 bp overlap
CTCF 135 datasets
ChIP A-549 ENCSR000BHW.CTCF.A-549 120 bp overlap
ChIP A-549 ENCSR000DNA.CTCF.A-549 132 bp overlap
ChIP A-549 ENCSR000DPF.CTCF.A-549 215 bp overlap
ChIP A549 ENCFF034FVO 217 bp overlap
ChIP C4-2B ENCFF821XVN 217 bp overlap
ChIP Caco-2 ENCFF934QYS 211 bp overlap
ChIP GM04648 GSE148179.CTCF.GM04648 151 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 162 bp overlap
ChIP GM12869 ENCSR000DRJ.CTCF.GM12869 98 bp overlap
ChIP GM12878 ENCFF511URZ 201 bp overlap
ChIP GM12878 ENCFF635MMB 164 bp overlap
ChIP GM12878 GSE76922.CTCF.GM12878 217 bp overlap
ChIP GM23338 ENCFF531QOI 131 bp overlap
ChIP GM23338 ENCFF772DML 159 bp overlap
ChIP H1 ENCFF230QSV 165 bp overlap
ChIP H1 ENCFF414GZI 186 bp overlap
ChIP H1 ENCFF764RHO 87 bp overlap
ChIP H9 ENCFF152GTF 217 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 217 bp overlap
ChIP HAP1_2kb GSE152721.CTCF.HAP1_2kb 214 bp overlap
ChIP HAP1_WT GSE126634.CTCF.HAP1_WT 217 bp overlap
ChIP HAP1_clone21 GSE152721.CTCF.HAP1_clone21 217 bp overlap
ChIP HAP1_clone21_CTCF-CRISPR GSE152721.CTCF.HAP1_clone21_CTCF-CRISPR 217 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 217 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_2 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_2 217 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 217 bp overlap
ChIP HAP1_clone25 GSE152721.CTCF.HAP1_clone25 217 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 217 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 217 bp overlap
ChIP HCT-116 ENCSR000DTO.CTCF.HCT-116 217 bp overlap
ChIP HCT-116 ENCSR000BSE.CTCF.HCT-116 188 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.CTCF.HCT-116_RAD21-mAC 217 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.CTCF.HCT-116_RAD21-mAC_500uM_auxin 217 bp overlap
ChIP HCT116 ENCFF003KHP 217 bp overlap
ChIP HCT116 ENCFF209YMI 217 bp overlap
ChIP HCT116 ENCFF373YMA 217 bp overlap
ChIP HEC-1-B_R1-mutation GSE140868.CTCF.HEC-1-B_R1-mutation 54 bp overlap
ChIP HEC-1-B_WT GSE140868.CTCF.HEC-1-B_WT 106 bp overlap
ChIP HEC-1-B_ac1-CBS-deletion GSE140868.CTCF.HEC-1-B_ac1-CBS-deletion 58 bp overlap
ChIP HEK293 ENCSR000DTW.CTCF.HEK293 115 bp overlap
ChIP HUES-64 GSE97394.CTCF.HUES-64 217 bp overlap
ChIP HUES-64_DNMT-KO GSE97394.CTCF.HUES-64_DNMT-KO 217 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 217 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 193 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 180 bp overlap
ChIP HeLa-S3 ENCFF565UFR 138 bp overlap
ChIP HeLa-S3 ENCSR000AOA.CTCF.HeLa-S3 203 bp overlap
ChIP HeLa-S3 ENCSR000DLO.CTCF.HeLa-S3 121 bp overlap
ChIP HeLa-S3 ENCSR000DUB.CTCF.HeLa-S3 102 bp overlap
ChIP HeLa-S3_biotin GSE108869.CTCF.HeLa-S3_biotin 209 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 168 bp overlap
ChIP HeLa-S3_unsynchro GSE108173.CTCF.HeLa-S3_unsynchro 136 bp overlap
ChIP Hep-G2 ENCSR000BIE.CTCF.Hep-G2 170 bp overlap
ChIP Hep-G2 ENCSR000DLS.CTCF.Hep-G2 126 bp overlap
ChIP Hep-G2 ENCSR000DUG.CTCF.Hep-G2 120 bp overlap
ChIP HepG2 ENCFF127KUP 217 bp overlap
ChIP HepG2 ENCFF348BUL 180 bp overlap
ChIP ID00014 GSE76922.CTCF.ID00014 217 bp overlap
ChIP ID00016 GSE76922.CTCF.ID00016 200 bp overlap
ChIP K-562 ENCSR000AKO.CTCF.K-562 217 bp overlap
ChIP K-562 GSE70764.CTCF.K-562 184 bp overlap
ChIP K-562 ENCSR000EGM.CTCF.K-562 134 bp overlap
ChIP K-562 ENCSR000BPJ.CTCF.K-562 204 bp overlap
ChIP K-562 ENCSR000DMA.CTCF.K-562 152 bp overlap
ChIP K-562_13004 GSE131349.CTCF.K-562_13004 129 bp overlap
ChIP K-562_14259 GSE131349.CTCF.K-562_14259 99 bp overlap
ChIP K-562_MonoC GSE137216.CTCF.K-562_MonoC 217 bp overlap
ChIP K-562_MonoN GSE137216.CTCF.K-562_MonoN 181 bp overlap
ChIP K-562_RF2-insertion GSE140868.CTCF.K-562_RF2-insertion 157 bp overlap
ChIP K-562_RabbitC GSE137216.CTCF.K-562_RabbitC 217 bp overlap
ChIP K-562_WT GSE140868.CTCF.K-562_WT 208 bp overlap
ChIP K-562_sgGal4 GSE132212.CTCF.K-562_sgGal4 217 bp overlap
ChIP K562 ENCFF082GOI 181 bp overlap
ChIP K562 ENCFF111MGE 211 bp overlap
ChIP K562 ENCFF400DFR 184 bp overlap
ChIP LNCaP_Ctrl GSE125639.CTCF.LNCaP_Ctrl 167 bp overlap
ChIP MCF-7 ENCFF139NQI 217 bp overlap
ChIP MCF-7 ENCFF162GNE 187 bp overlap
ChIP MCF-7 ENCFF198DQX 188 bp overlap
ChIP MCF-7 ENCFF414SZG 83 bp overlap
ChIP MCF-7 ENCFF424NQR 168 bp overlap
ChIP MCF-7 ENCFF494VXA 188 bp overlap
ChIP MCF-7 ENCFF844STM 168 bp overlap
ChIP MCF-7 ENCFF954TUV 166 bp overlap
ChIP MCF-7 GSE137216.CTCF.MCF-7 213 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 217 bp overlap
ChIP MCF-7 ENCSR000AHD.CTCF.MCF-7 182 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 184 bp overlap
ChIP MCF-7 GSE70764.CTCF.MCF-7 176 bp overlap
ChIP MCF-7 ENCSR000DWH.CTCF.MCF-7 206 bp overlap
ChIP MCF-7 ENCSR000DMS.CTCF.MCF-7 152 bp overlap
ChIP MCF-7 ERP000380.CTCF.MCF-7 117 bp overlap
ChIP MCF-7L_t1 GSE108787.CTCF.MCF-7L_t1 177 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 193 bp overlap
ChIP MCF-7_E2 ERP000380.CTCF.MCF-7_E2 127 bp overlap
ChIP MCF-7_NORMOXIA GSE78113.CTCF.MCF-7_NORMOXIA 217 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 140 bp overlap
ChIP OCI-Ly7 ENCSR027HML.CTCF.OCI-Ly7 198 bp overlap
ChIP PC-3 ENCFF487TUI 217 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 217 bp overlap
ChIP Peyer's patch ENCFF701KWW 141 bp overlap
ChIP Peyers-patch ENCSR419ANE.CTCF.Peyers-patch 122 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 144 bp overlap
ChIP SK-MEL-147 GSE94488.CTCF.SK-MEL-147 107 bp overlap
ChIP SK-N-AS GSE101295.CTCF.SK-N-AS 124 bp overlap
ChIP SK-N-SH ENCSR000BLX.CTCF.SK-N-SH 99 bp overlap
ChIP SUM159 GSE46055.CTCF.SUM159 88 bp overlap
ChIP T-47D GSE111923.CTCF.T-47D 189 bp overlap
ChIP THP-1_Pam3csk-030m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-030m-Flavo-000m 155 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 193 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 153 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 203 bp overlap
ChIP ZR751 ERP000783.CTCF.ZR751 106 bp overlap
ChIP colon_transverse ENCSR102CSD.CTCF.colon_transverse 161 bp overlap
ChIP endodermal cell ENCFF471YCZ 217 bp overlap
ChIP endodermal cell ENCFF471YCZ 153 bp overlap
ChIP hESC GSE20650.CTCF.hESC 99 bp overlap
ChIP hESC_D0 GSE116862.CTCF.hESC_D0 190 bp overlap
ChIP hESC_NAIVE GSE69646.CTCF.hESC_NAIVE 217 bp overlap
ChIP hepatocyte ENCSR252QYR.CTCF.hepatocyte 124 bp overlap
ChIP hiPSC GSE115407.CTCF.hiPSC 217 bp overlap
ChIP hiPSC_Angelman GSE117283.CTCF.hiPSC_Angelman 217 bp overlap
ChIP hiPSC_CVI GSE106870.CTCF.hiPSC_CVI 128 bp overlap
ChIP hiPSC_HUES9 GSE106870.CTCF.hiPSC_HUES9 169 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 145 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 163 bp overlap
ChIP hiPSC_IIA11 GSE106870.CTCF.hiPSC_IIA11 164 bp overlap
ChIP hiPSC_IID12 GSE106870.CTCF.hiPSC_IID12 130 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 173 bp overlap
ChIP hiPSC_Prader-Willi GSE117283.CTCF.hiPSC_Prader-Willi 180 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 159 bp overlap
ChIP hiPSC_parental-line GSE106870.CTCF.hiPSC_parental-line 217 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 163 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 217 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 142 bp overlap
DAXX 1 dataset
ChIP PC-3 GSE68647.DAXX.PC-3 50 bp overlap
DMRTC2 2 datasets
Motif DE_12h DE_12h-DMRTC2_MA1479.2 11 bp overlap
Motif ES_0h ES_0h-DMRTC2_MA1479.2 11 bp overlap
DUX4 1 dataset
ChIP WA01 GSE94322.DUX4.WA01 57 bp overlap
ESR1 11 datasets
ChIP MCF-7 GSE119057.ESR1.MCF-7 158 bp overlap
ChIP MCF-7_ER_t10 GSE119057.ESR1.MCF-7_ER_t10 157 bp overlap
ChIP MCF-7_ER_t20 GSE119057.ESR1.MCF-7_ER_t20 174 bp overlap
ChIP MCF-7_ER_t30 GSE119057.ESR1.MCF-7_ER_t30 162 bp overlap
ChIP MCF-7_ER_t50 GSE119057.ESR1.MCF-7_ER_t50 186 bp overlap
ChIP MCF-7_ER_t60 GSE119057.ESR1.MCF-7_ER_t60 150 bp overlap
ChIP MCF-7_ER_t70 GSE119057.ESR1.MCF-7_ER_t70 174 bp overlap
ChIP MCF-7_ER_t80 GSE119057.ESR1.MCF-7_ER_t80 149 bp overlap
ChIP MCF-7_ER_t90 GSE119057.ESR1.MCF-7_ER_t90 168 bp overlap
ChIP MCF-7_jc5849 GSE126004.ESR1.MCF-7_jc5849 114 bp overlap
ChIP MCF-7_jc5850 GSE126004.ESR1.MCF-7_jc5850 93 bp overlap
FOXA1 3 datasets
ChIP LNCaP_VEH GSE114266.FOXA1.LNCaP_VEH 139 bp overlap
ChIP prostate-cancer_PDX_189-3 GSE130408.FOXA1.prostate-cancer_PDX_189-3 214 bp overlap
ChIP prostate_P7_T GSE130408.FOXA1.prostate_P7_T 90 bp overlap
FOXP1 1 dataset
ChIP H9 GSE31006.FOXP1.H9 157 bp overlap
HNF4A 2 datasets
ChIP Caco-2_DIFF GSE23436.HNF4A.Caco-2_DIFF 148 bp overlap
ChIP IM95 GSE114018.HNF4A.IM95 133 bp overlap
HOXB13 2 datasets
ChIP LNCaP GSE96652.HOXB13.LNCaP 105 bp overlap
ChIP prostate_P7_T GSE130408.HOXB13.prostate_P7_T 74 bp overlap
MAX 3 datasets
ChIP HCT-116 ENCSR000BSH.MAX.HCT-116 141 bp overlap
ChIP HCT116 ENCFF810LEN 217 bp overlap
ChIP MCF-7 ENCSR000BUL.MAX.MCF-7 108 bp overlap
MCRS1 1 dataset
ChIP Huh-7 GSE97411.MCRS1.Huh-7 217 bp overlap
MXI1 2 datasets
Motif DE_12h DE_12h-MXI1_MA1108.3 6 bp overlap
Motif ES_0h ES_0h-MXI1_MA1108.3 6 bp overlap
MYC 1 dataset
ChIP MCF-7 ENCSR000DMQ.MYC.MCF-7 117 bp overlap
NKX2-2 2 datasets
Motif DE_12h DE_12h-NKX2-2_MA1645.2 8 bp overlap
Motif ES_0h ES_0h-NKX2-2_MA1645.2 8 bp overlap
NKX2-5 2 datasets
Motif DE_12h DE_12h-NKX2-5_MA0063.3 7 bp overlap
Motif ES_0h ES_0h-NKX2-5_MA0063.3 7 bp overlap
PHIP 1 dataset
ChIP HCT-116_ab833 GSE101646.PHIP.HCT-116_ab833 155 bp overlap
POU2F2 1 dataset
ChIP HNPC_UNDIF GSE74814.POU2F2.HNPC_UNDIF 151 bp overlap
RAD21 33 datasets
ChIP A-549 ENCSR000BUC.RAD21.A-549 146 bp overlap
ChIP A549 ENCFF047SFC 201 bp overlap
ChIP H1 ENCFF698EWO 115 bp overlap
ChIP H1 ENCFF967OJF 215 bp overlap
ChIP HAP1 GSE152721.RAD21.HAP1 217 bp overlap
ChIP HAP1_CTCF GSE126634.RAD21.HAP1_CTCF 217 bp overlap
ChIP HAP1_clone25 GSE152721.RAD21.HAP1_clone25 139 bp overlap
ChIP HCT-116 GSE121355.RAD21.HCT-116 217 bp overlap
ChIP HCT-116 ENCSR000BSB.RAD21.HCT-116 199 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.RAD21.HCT-116_RAD21-mAC 213 bp overlap
ChIP HCT116 ENCFF568PEO 217 bp overlap
ChIP HEC-1-B_F-insertion GSE140868.RAD21.HEC-1-B_F-insertion 104 bp overlap
ChIP Hep-G2 ENCSR054FKH.RAD21.Hep-G2 148 bp overlap
ChIP HepG2 ENCFF360ZSW 217 bp overlap
ChIP HepG2 ENCFF906QIS 187 bp overlap
ChIP Ishikawa ENCSR000BTU.RAD21.Ishikawa 106 bp overlap
ChIP K-562 ENCSR000BKV.RAD21.K-562 171 bp overlap
ChIP K-562_RF2-insertion GSE140868.RAD21.K-562_RF2-insertion 99 bp overlap
ChIP K562 ENCFF634XYR 217 bp overlap
ChIP MCF-7 ENCFF694KOM 217 bp overlap
ChIP MCF-7 ENCFF724VCQ 179 bp overlap
ChIP MCF-7 GSE72082.RAD21.MCF-7 166 bp overlap
ChIP MCF-7 GSE137216.RAD21.MCF-7 172 bp overlap
ChIP MCF-7 ENCSR000BTQ.RAD21.MCF-7 142 bp overlap
ChIP MCF-7 ENCSR703TNG.RAD21.MCF-7 208 bp overlap
ChIP WA09 GSE105028.RAD21.WA09 207 bp overlap
ChIP WA09_heat-shock GSE105028.RAD21.WA09_heat-shock 205 bp overlap
ChIP hiPSC_CVI GSE106870.RAD21.hiPSC_CVI 166 bp overlap
ChIP hiPSC_IIA11 GSE106870.RAD21.hiPSC_IIA11 195 bp overlap
ChIP hiPSC_IIA12 GSE106870.RAD21.hiPSC_IIA12 217 bp overlap
ChIP hiPSC_IIIB12 GSE106870.RAD21.hiPSC_IIIB12 217 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.RAD21.hiPSC_failed-genome-editing 217 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 200 bp overlap
RARA 1 dataset
ChIP hiPSC_D3 GSE109172.RARA.hiPSC_D3 61 bp overlap
RBPJ 2 datasets
Motif DE_12h DE_12h-RBPJ_MA1116.2 6 bp overlap
Motif ES_0h ES_0h-RBPJ_MA1116.2 6 bp overlap
SMAD1 1 dataset
ChIP BG03 GSE36578.SMAD1.BG03 163 bp overlap
SMAD3 1 dataset
ChIP BG03 GSE36578.SMAD3.BG03 126 bp overlap
SMC1 2 datasets
ChIP HCAEC GSE101921.SMC1.HCAEC 206 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.SMC1.HCT-116_RAD21-mAC 153 bp overlap
SMC1A 2 datasets
ChIP Hep-G2 GSE76893.SMC1A.Hep-G2 174 bp overlap
ChIP MCF-7 GSE76893.SMC1A.MCF-7 141 bp overlap
SMC3 1 dataset
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.SMC3.HeLa-Kyoto_WAPL_PDS-depleted 213 bp overlap
STAG1 4 datasets
ChIP HeLa GSE126990.STAG1.HeLa 144 bp overlap
ChIP HeLa-Kyoto GSE138405.STAG1.HeLa-Kyoto 144 bp overlap
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 118 bp overlap
ChIP MCF-7 ERP000209.STAG1.MCF-7 143 bp overlap
TBX19 2 datasets
Motif DE_12h DE_12h-TBX19_MA0804.2 17 bp overlap
Motif ES_0h ES_0h-TBX19_MA0804.2 17 bp overlap
TBXT 2 datasets
Motif DE_12h DE_12h-TBXT_MA0009.2 16 bp overlap
Motif ES_0h ES_0h-TBXT_MA0009.2 16 bp overlap
TERF1 1 dataset
ChIP LCL GSE55053.TERF1.LCL 51 bp overlap
TLE3 2 datasets
ChIP LNCaP GSE94682.TLE3.LNCaP 60 bp overlap
ChIP LNCaP_r1881 GSE94682.TLE3.LNCaP_r1881 128 bp overlap
USF1 2 datasets
ChIP HCT116 ENCFF330PYP 217 bp overlap
ChIP WA01 ENCSR000BIU.USF1.WA01 123 bp overlap
ZBTB2 2 datasets
ChIP GM12878 GSE97661.ZBTB2.GM12878 123 bp overlap
ChIP Hep-G2 GSE97661.ZBTB2.Hep-G2 129 bp overlap
ZNF24 2 datasets
Motif DE_12h DE_12h-ZNF24_MA1124.1 13 bp overlap
Motif ES_0h ES_0h-ZNF24_MA1124.1 13 bp overlap
ZNF417 2 datasets
Motif DE_12h DE_12h-ZNF417_MA1727.2 7 bp overlap
Motif ES_0h ES_0h-ZNF417_MA1727.2 7 bp overlap
Zfp335 2 datasets
Motif DE_12h DE_12h-Zfp335_MA2002.2 7 bp overlap
Motif ES_0h ES_0h-Zfp335_MA2002.2 7 bp overlap
Zfx 2 datasets
Motif DE_12h DE_12h-Zfx_MA0146.3 10 bp overlap
Motif ES_0h ES_0h-Zfx_MA0146.3 10 bp overlap