chr5 : 165,757,525 165,758,404
879 bp 56 TFs 0 linked genes
This 879 bp open chromatin element has no linked target genes and is bound by 56 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr5:165,752,525 – 165,763,404
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
56 transcription factors
Source
Cell type
AR 65 datasets
ChIP DUCAP_ANDROGEN GSE70679.AR.DUCAP_ANDROGEN 259 bp overlap
ChIP LAPC-4_R1881 GSE148358.AR.LAPC-4_R1881 223 bp overlap
ChIP LNCaP GSE80256.AR.LNCaP 267 bp overlap
ChIP LNCaP GSE117430.AR.LNCaP 265 bp overlap
ChIP LNCaP GSE85558.AR.LNCaP 209 bp overlap
ChIP LNCaP ERP003503.AR.LNCaP 228 bp overlap
ChIP LNCaP GSE110655.AR.LNCaP 199 bp overlap
ChIP LNCaP ERP001226.AR.LNCaP 141 bp overlap
ChIP LNCaP-abl GSE39459.AR.LNCaP-abl 222 bp overlap
ChIP LNCaP_1F5 GSE30623.AR.LNCaP_1F5 240 bp overlap
ChIP LNCaP_1F5_SIFOXA1 GSE30623.AR.LNCaP_1F5_SIFOXA1 257 bp overlap
ChIP LNCaP_Bag1L_KO_Bag1L_Rescue_DHT_4h GSE89938.AR.LNCaP_Bag1L_KO_Bag1L_Rescue_DHT_4h 209 bp overlap
ChIP LNCaP_DHT GSE125245.AR.LNCaP_DHT 133 bp overlap
ChIP LNCaP_DHT GSE43720.AR.LNCaP_DHT 237 bp overlap
ChIP LNCaP_DHT24H GSE58428.AR.LNCaP_DHT24H 324 bp overlap
ChIP LNCaP_DSG GSE114737.AR.LNCaP_DSG 250 bp overlap
ChIP LNCaP_ETOH GSE69043.AR.LNCaP_ETOH 224 bp overlap
ChIP LNCaP_FOXA1 ERP003503.AR.LNCaP_FOXA1 264 bp overlap
ChIP LNCaP_GFP_shFOXA1_Ethanol GSE128883.AR.LNCaP_GFP_shFOXA1_Ethanol 247 bp overlap
ChIP LNCaP_R1881 GSE62492.AR.LNCaP_R1881 281 bp overlap
ChIP LNCaP_R1881 GSE69043.AR.LNCaP_R1881 253 bp overlap
ChIP LNCaP_SHCTR_DHT GSE62492.AR.LNCaP_SHCTR_DHT 320 bp overlap
ChIP LNCaP_SHFOXA1_R1881 GSE37345.AR.LNCaP_SHFOXA1_R1881 181 bp overlap
ChIP LNCaP_SHFOXP1_DHT GSE62492.AR.LNCaP_SHFOXP1_DHT 249 bp overlap
ChIP LNCaP_SHGATA2_ETOH GSE69043.AR.LNCaP_SHGATA2_ETOH 140 bp overlap
ChIP LNCaP_SHGATA2_R1881 GSE69043.AR.LNCaP_SHGATA2_R1881 279 bp overlap
ChIP LNCaP_Talen_DHT GSE89938.AR.LNCaP_Talen_DHT 272 bp overlap
ChIP LNCaP_androgen-N_hypoxia-Y GSE114732.AR.LNCaP_androgen-N_hypoxia-Y 398 bp overlap
ChIP LNCaP_androgen-Y_hypoxia-N GSE114732.AR.LNCaP_androgen-Y_hypoxia-N 212 bp overlap
ChIP LNCaP_androgen-Y_hypoxia-Y GSE114732.AR.LNCaP_androgen-Y_hypoxia-Y 227 bp overlap
ChIP LNCaP_r1881 GSE94682.AR.LNCaP_r1881 426 bp overlap
ChIP LNCaP_sgCHD1 GSE117430.AR.LNCaP_sgCHD1 366 bp overlap
ChIP LNCaP_sgCtrl GSE117430.AR.LNCaP_sgCtrl 376 bp overlap
ChIP LTAD_siControl GSE94577.AR.LTAD_siControl 198 bp overlap
ChIP MDA-MB-453_R1881_SICTR GSE70161.AR.MDA-MB-453_R1881_SICTR 190 bp overlap
ChIP MDA-MB-453_R1881_SIPIAS1 GSE70161.AR.MDA-MB-453_R1881_SIPIAS1 177 bp overlap
ChIP PC-3_R1881 GSE54110.AR.PC-3_R1881 231 bp overlap
ChIP VCaP_DHAT_2H GSE28950.AR.VCaP_DHAT_2H 256 bp overlap
ChIP VCaP_DHT24H_SHFOXA1 GSE58428.AR.VCaP_DHT24H_SHFOXA1 318 bp overlap
ChIP VCaP_R1881 GSE32892.AR.VCaP_R1881 202 bp overlap
ChIP VCaP_R1881_30M GSE84432.AR.VCaP_R1881_30M 276 bp overlap
ChIP VCaP_R1881_4H GSE84432.AR.VCaP_R1881_4H 297 bp overlap
ChIP VCaP_SH1_DHT GSE79128.AR.VCaP_SH1_DHT 349 bp overlap
ChIP VCaP_SH1_R1881 GSE79128.AR.VCaP_SH1_R1881 304 bp overlap
ChIP VCaP_shERG GSE110655.AR.VCaP_shERG 217 bp overlap
ChIP prostate GSE65478.AR.prostate 288 bp overlap
ChIP prostate GSE56288.AR.prostate 189 bp overlap
ChIP prostate-cancer GSE136128.AR.prostate-cancer 153 bp overlap
ChIP prostate-cancer_C4-2-CON GSE136128.AR.prostate-cancer_C4-2-CON 296 bp overlap
ChIP prostate-cancer_PDX_167 GSE130408.AR.prostate-cancer_PDX_167 227 bp overlap
ChIP prostate-cancer_PDX_189-4 GSE130408.AR.prostate-cancer_PDX_189-4 166 bp overlap
ChIP prostate-cancer_PDX_70 GSE130408.AR.prostate-cancer_PDX_70 187 bp overlap
ChIP prostate-cancer_PDX_78 GSE130408.AR.prostate-cancer_PDX_78 174 bp overlap
ChIP prostate-cancer_PDX_92 GSE130408.AR.prostate-cancer_PDX_92 157 bp overlap
ChIP prostate_1239_T GSE130408.AR.prostate_1239_T 109 bp overlap
ChIP prostate_1816_T GSE130408.AR.prostate_1816_T 287 bp overlap
ChIP prostate_1853_T GSE130408.AR.prostate_1853_T 236 bp overlap
ChIP prostate_2752_T GSE130408.AR.prostate_2752_T 364 bp overlap
ChIP prostate_2815_T GSE130408.AR.prostate_2815_T 502 bp overlap
ChIP prostate_4024_T GSE130408.AR.prostate_4024_T 434 bp overlap
ChIP prostate_P13_T GSE130408.AR.prostate_P13_T 233 bp overlap
ChIP prostate_P23_T GSE130408.AR.prostate_P23_T 211 bp overlap
ChIP prostate_P25 GSE130408.AR.prostate_P25 275 bp overlap
ChIP prostate_P29_T GSE130408.AR.prostate_P29_T 173 bp overlap
ChIP prostate_P5_T GSE130408.AR.prostate_P5_T 239 bp overlap
ARID1A 3 datasets
ChIP HAP1 GSE108387.ARID1A.HAP1 334 bp overlap
ChIP MCF-7_Fulvestrant GSE123284.ARID1A.MCF-7_Fulvestrant 275 bp overlap
ChIP NGP GSE134626.ARID1A.NGP 338 bp overlap
ATRX 1 dataset
ChIP metastatic-neuroblastoma_CHLA90 GSE100148.ATRX.metastatic-neuroblastoma_CHLA90 496 bp overlap
BAF155 1 dataset
ChIP VCaP_shARID1A GSE110655.BAF155.VCaP_shARID1A 160 bp overlap
BRD4 5 datasets
ChIP LNCaP-C4-2_F133V GSE88871.BRD4.LNCaP-C4-2_F133V 769 bp overlap
ChIP LNCaP-C4-2_F133V_JQ1 GSE88871.BRD4.LNCaP-C4-2_F133V_JQ1 253 bp overlap
ChIP LNCaP-C4-2_F133V_JQ1 GSE88871.BRD4.LNCaP-C4-2_F133V_JQ1 189 bp overlap
ChIP LNCaP-C4-2_OE GSE88871.BRD4.LNCaP-C4-2_OE 838 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 255 bp overlap
Bcl11B 1 dataset
Motif DE_36h DE_36h-Bcl11B_MA1989.2 9 bp overlap
CHD8 1 dataset
ChIP T-47D_R5020_45 GSE62428.CHD8.T-47D_R5020_45 253 bp overlap
CREB1 1 dataset
ChIP LNCaP-abl_SHCTR GSE63034.CREB1.LNCaP-abl_SHCTR 402 bp overlap
CTCF 4 datasets
ChIP HAP1_clone21_CTCF-CRISPR GSE152721.CTCF.HAP1_clone21_CTCF-CRISPR 546 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 175 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 167 bp overlap
ChIP T-47D_NaCl-triptolide GSE111923.CTCF.T-47D_NaCl-triptolide 310 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF031ISE 105 bp overlap
DUX4 1 dataset
ChIP WA01 GSE94322.DUX4.WA01 428 bp overlap
EOMES 1 dataset
ChIP hESC GSE26097.EOMES.hESC 491 bp overlap
ESR1 2 datasets
ChIP T-47D_PROG GSE68355.ESR1.T-47D_PROG 263 bp overlap
ChIP T-47D_R5020 GSE68355.ESR1.T-47D_R5020 230 bp overlap
FOXA1 11 datasets
ChIP LNCaP_R1881 GSE69043.FOXA1.LNCaP_R1881 188 bp overlap
ChIP LNCaP_SHGATA2_R1881 GSE69043.FOXA1.LNCaP_SHGATA2_R1881 134 bp overlap
ChIP LNCaP_VEH GSE114266.FOXA1.LNCaP_VEH 208 bp overlap
ChIP T-47D_E2_TAM ERP000380.FOXA1.T-47D_E2_TAM 240 bp overlap
ChIP T-47D_JC4748 GSE126004.FOXA1.T-47D_JC4748 298 bp overlap
ChIP primary-prostate-cancer_P2_DSG GSE114737.FOXA1.primary-prostate-cancer_P2_DSG 316 bp overlap
ChIP prostate-cancer_PDX_136 GSE130408.FOXA1.prostate-cancer_PDX_136 199 bp overlap
ChIP prostate_2480 GSE130408.FOXA1.prostate_2480 343 bp overlap
ChIP prostate_P13_T GSE130408.FOXA1.prostate_P13_T 228 bp overlap
ChIP prostate_P1_T GSE130408.FOXA1.prostate_P1_T 197 bp overlap
ChIP prostate_P23_T GSE130408.FOXA1.prostate_P23_T 269 bp overlap
FOXA2 1 dataset
ChIP BJ1-hTERT_FOXA2_GATA4_Coexp GSE92491.FOXA2.BJ1-hTERT_FOXA2_GATA4_Coexp 300 bp overlap
GABPA 1 dataset
ChIP VCaP_ETOH GSE49091.GABPA.VCaP_ETOH 143 bp overlap
GATA2 9 datasets
Motif DE_36h DE_36h-GATA2_MA0036.4 7 bp overlap
ChIP LNCaP GSE38391.GATA2.LNCaP 294 bp overlap
ChIP LNCaP GSE52725.GATA2.LNCaP 294 bp overlap
ChIP LNCaP_CSFCS GSE69043.GATA2.LNCaP_CSFCS 304 bp overlap
ChIP LNCaP_FBS GSE69043.GATA2.LNCaP_FBS 290 bp overlap
ChIP SH-SY5Y ENCSR000EYB.GATA2.SH-SY5Y 343 bp overlap
ChIP TF1 GSE73207.GATA2.TF1 207 bp overlap
ChIP VCaP GSE125236.GATA2.VCaP 209 bp overlap
ChIP VCaP_JQ1 GSE125236.GATA2.VCaP_JQ1 213 bp overlap
GATA3 8 datasets
ChIP MCF-7 ENCFF437NQS 371 bp overlap
ChIP MCF-7 GSE51274.GATA3.MCF-7 450 bp overlap
ChIP MCF-7 GSE133072.GATA3.MCF-7 175 bp overlap
ChIP MCF-7 GSE122847.GATA3.MCF-7 331 bp overlap
ChIP MCF-7_E2 GSE40129.GATA3.MCF-7_E2 129 bp overlap
ChIP T-47D GSE51274.GATA3.T-47D 457 bp overlap
ChIP T-47D ENCSR000BMX.GATA3.T-47D 362 bp overlap
ChIP T-47D_sc GSE122847.GATA3.T-47D_sc 178 bp overlap
GATA3_Cter 2 datasets
ChIP T-47D_CR3flp GSE99479.GATA3_Cter.T-47D_CR3flp 253 bp overlap
ChIP T-47D_flp-ctrl GSE99479.GATA3_Cter.T-47D_flp-ctrl 231 bp overlap
GATA3_Nter 2 datasets
ChIP T-47D_CR3flp GSE99479.GATA3_Nter.T-47D_CR3flp 438 bp overlap
ChIP T-47D_flp-ctrl GSE99479.GATA3_Nter.T-47D_flp-ctrl 424 bp overlap
GATA4 5 datasets
ChIP DE DE-GATA4-1 619 bp overlap
ChIP DE DE-GATA4-2 535 bp overlap
Motif DE_36h DE_36h-GATA4_MA0482.3 8 bp overlap
ChIP Hep-G2 GSE135714.GATA4.Hep-G2 566 bp overlap
ChIP pancreatic-progenitor GSE117136.GATA4.pancreatic-progenitor 647 bp overlap
GATA5 1 dataset
Motif DE_36h DE_36h-GATA5_MA0766.3 8 bp overlap
GATA6 10 datasets
ChIP DE DE-GATA6-1 454 bp overlap
ChIP DE DE-GATA6-2 492 bp overlap
Motif DE_36h DE_36h-GATA6_MA1104.3 8 bp overlap
ChIP DE_D1 S09-DE-d1-GATA6-exp1 349 bp overlap
ChIP DE_D1 S14-DE-d1-GATA6-exp1 571 bp overlap
ChIP DE_D1 S41-DE-d1-GATA6-exp2 299 bp overlap
ChIP DE_D2 S55-DE-d2-GATA6-exp2 708 bp overlap
ChIP HUES-8_DE_JNKi GSE109524.GATA6.HUES-8_DE_JNKi 626 bp overlap
ChIP HUES-8_DE_ctrl GSE109524.GATA6.HUES-8_DE_ctrl 457 bp overlap
ChIP endoderm_KO GSE117136.GATA6.endoderm_KO 611 bp overlap
Gata3 1 dataset
Motif DE_36h DE_36h-Gata3_MA0037.5 8 bp overlap
HOXB13 20 datasets
ChIP prostate-cancer_PDX_167 GSE130408.HOXB13.prostate-cancer_PDX_167 287 bp overlap
ChIP prostate_2030_T GSE130408.HOXB13.prostate_2030_T 286 bp overlap
ChIP prostate_2078_T GSE130408.HOXB13.prostate_2078_T 256 bp overlap
ChIP prostate_2480 GSE130408.HOXB13.prostate_2480 315 bp overlap
ChIP prostate_2480_T GSE130408.HOXB13.prostate_2480_T 275 bp overlap
ChIP prostate_2483 GSE130408.HOXB13.prostate_2483 248 bp overlap
ChIP prostate_2483_T GSE130408.HOXB13.prostate_2483_T 208 bp overlap
ChIP prostate_2484 GSE130408.HOXB13.prostate_2484 268 bp overlap
ChIP prostate_2484_T GSE130408.HOXB13.prostate_2484_T 177 bp overlap
ChIP prostate_P13 GSE130408.HOXB13.prostate_P13 148 bp overlap
ChIP prostate_P13_T GSE130408.HOXB13.prostate_P13_T 235 bp overlap
ChIP prostate_P19 GSE130408.HOXB13.prostate_P19 288 bp overlap
ChIP prostate_P19_T GSE130408.HOXB13.prostate_P19_T 474 bp overlap
ChIP prostate_P1_T GSE130408.HOXB13.prostate_P1_T 279 bp overlap
ChIP prostate_P23_T GSE130408.HOXB13.prostate_P23_T 272 bp overlap
ChIP prostate_P25 GSE130408.HOXB13.prostate_P25 276 bp overlap
ChIP prostate_P27_T GSE130408.HOXB13.prostate_P27_T 227 bp overlap
ChIP prostate_P29 GSE130408.HOXB13.prostate_P29 245 bp overlap
ChIP prostate_P29_T GSE130408.HOXB13.prostate_P29_T 275 bp overlap
ChIP prostate_P5_T GSE130408.HOXB13.prostate_P5_T 205 bp overlap
KDM5B 1 dataset
ChIP SUM185_SHCTCF GSE46055.KDM5B.SUM185_SHCTCF 215 bp overlap
KLF4 1 dataset
ChIP HAP1 GSE130417.KLF4.HAP1 193 bp overlap
NANOG 3 datasets
ChIP WA01_3IL ERP004238.NANOG.WA01_3IL 400 bp overlap
ChIP WA09 GSE105028.NANOG.WA09 219 bp overlap
ChIP hESC GSE18292.NANOG.hESC 102 bp overlap
NR3C1 3 datasets
ChIP LNCaP_1F5_SIFOXA1 GSE30623.NR3C1.LNCaP_1F5_SIFOXA1 116 bp overlap
ChIP SUM159PT GSE152203.NR3C1.SUM159PT 156 bp overlap
ChIP T47D-A1-2_Dex GSE112491.NR3C1.T47D-A1-2_Dex 120 bp overlap
PGR 10 datasets
ChIP MCF-7_PROG GSE68355.PGR.MCF-7_PROG 252 bp overlap
ChIP T-47D GSE31129.PGR.T-47D 372 bp overlap
ChIP T-47D-A_E2_R5020 GSE80358.PGR.T-47D-A_E2_R5020 474 bp overlap
ChIP T-47D_E2PG GSE68356.PGR.T-47D_E2PG 275 bp overlap
ChIP T-47D_PG GSE68356.PGR.T-47D_PG 294 bp overlap
ChIP T-47D_PROG GSE68355.PGR.T-47D_PROG 388 bp overlap
ChIP T-47D_R5020 GSE68355.PGR.T-47D_R5020 356 bp overlap
ChIP T-47D_R5020-A1 GSE126859.PGR.T-47D_R5020-A1 415 bp overlap
ChIP T-47D_R5020-MTVL GSE126859.PGR.T-47D_R5020-MTVL 532 bp overlap
ChIP T-47D_progesterone GSE132649.PGR.T-47D_progesterone 375 bp overlap
POLR2A 1 dataset
ChIP prostate gland ENCFF881OMH 417 bp overlap
POU2F1 1 dataset
Motif DE_36h DE_36h-POU2F1_MA0785.2 9 bp overlap
POU2F2 1 dataset
Motif DE_36h DE_36h-POU2F2_MA0507.3 13 bp overlap
POU2F3 1 dataset
Motif DE_36h DE_36h-POU2F3_MA0627.3 9 bp overlap
POU3F1 1 dataset
Motif DE_36h DE_36h-POU3F1_MA0786.2 10 bp overlap
POU3F4 1 dataset
Motif DE_36h DE_36h-POU3F4_MA0789.1 9 bp overlap
POU5F1 2 datasets
ChIP NCCIT_SNF5 GSE36134.POU5F1.NCCIT_SNF5 546 bp overlap
ChIP hESC_NAIVE GSE69646.POU5F1.hESC_NAIVE 605 bp overlap
RAD21 4 datasets
ChIP HAP1 GSE126634.RAD21.HAP1 550 bp overlap
ChIP HAP1 GSE152721.RAD21.HAP1 495 bp overlap
ChIP HAP1_CTCF GSE126634.RAD21.HAP1_CTCF 521 bp overlap
ChIP HAP1_clone25 GSE152721.RAD21.HAP1_clone25 343 bp overlap
RELA 1 dataset
ChIP FaDu_LPS GSE132018.RELA.FaDu_LPS 127 bp overlap
SIX2 1 dataset
ChIP kidney_fetal_17w GSE75948.SIX2.kidney_fetal_17w 152 bp overlap
SMARCA4 3 datasets
ChIP NGP_ARID1A-mut2 GSE134626.SMARCA4.NGP_ARID1A-mut2 386 bp overlap
ChIP hiPSC GSE124903.SMARCA4.hiPSC 387 bp overlap
ChIP hiPSC_WT GSE124903.SMARCA4.hiPSC_WT 242 bp overlap
SMARCC1 2 datasets
ChIP hiPSC GSE124903.SMARCC1.hiPSC 206 bp overlap
ChIP hiPSC_WT GSE124903.SMARCC1.hiPSC_WT 203 bp overlap
SOX2 1 dataset
ChIP hiPSC GSE56567.SOX2.hiPSC 241 bp overlap
STAT3 19 datasets
Motif DE_36h DE_36h-STAT3_MA0144.3 9 bp overlap
ChIP HCC1187 GSE152203.STAT3.HCC1187 380 bp overlap
ChIP MCF-7_jc5834 GSE126004.STAT3.MCF-7_jc5834 374 bp overlap
ChIP MCF-7_jc5836 GSE126004.STAT3.MCF-7_jc5836 367 bp overlap
ChIP MCF-7_jc5838 GSE126004.STAT3.MCF-7_jc5838 377 bp overlap
ChIP MCF-7_jc5840 GSE126004.STAT3.MCF-7_jc5840 433 bp overlap
ChIP MCF-7_jc5842 GSE126004.STAT3.MCF-7_jc5842 266 bp overlap
ChIP MCF-7_jc5844 GSE126004.STAT3.MCF-7_jc5844 246 bp overlap
ChIP MCF-7_jc5848 GSE126004.STAT3.MCF-7_jc5848 275 bp overlap
ChIP T-47D_JC4735 GSE126004.STAT3.T-47D_JC4735 195 bp overlap
ChIP T-47D_JC4737 GSE126004.STAT3.T-47D_JC4737 193 bp overlap
ChIP T-47D_JC4741 GSE126004.STAT3.T-47D_JC4741 222 bp overlap
ChIP T-47D_JC4751 GSE126004.STAT3.T-47D_JC4751 447 bp overlap
ChIP T-47D_JC4753 GSE126004.STAT3.T-47D_JC4753 432 bp overlap
ChIP T-47D_JC4755 GSE126004.STAT3.T-47D_JC4755 232 bp overlap
ChIP T-47D_JC4757 GSE126004.STAT3.T-47D_JC4757 430 bp overlap
ChIP T-47D_JC5054 GSE126004.STAT3.T-47D_JC5054 427 bp overlap
ChIP T-47D_JC5056 GSE126004.STAT3.T-47D_JC5056 529 bp overlap
ChIP T-47D_JC5062 GSE126004.STAT3.T-47D_JC5062 423 bp overlap
T 1 dataset
ChIP H9_ENDODERM GSE60606.T.H9_ENDODERM 245 bp overlap
TEAD4 1 dataset
ChIP SNU-216 GSE44416.TEAD4.SNU-216 171 bp overlap
TRPS1 2 datasets
Motif DE_36h DE_36h-TRPS1_MA1970.2 8 bp overlap
ChIP T-47D GSE107013.TRPS1.T-47D 114 bp overlap
UBN1 1 dataset
ChIP HeLa GSE45024.UBN1.HeLa 272 bp overlap
ZNF418 1 dataset
Motif DE_36h DE_36h-ZNF418_MA1980.1 15 bp overlap
ZNF823 1 dataset
ChIP HEK293T GSE78099.ZNF823.HEK293T 307 bp overlap