chr5 : 33,890,772 33,890,937
165 bp 32 TFs 1 linked gene
This 165 bp open chromatin element is linked to ADAMTS12 and is bound by 32 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
ADAMTS12 1.1 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr5:33,885,772 – 33,895,937
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
32 transcription factors
Source
Cell type
ARID1A 3 datasets
ChIP MCF-7_4-hydroxytamoxifen GSE123284.ARID1A.MCF-7_4-hydroxytamoxifen 124 bp overlap
ChIP MCF-7_Fulvestrant GSE123284.ARID1A.MCF-7_Fulvestrant 72 bp overlap
ChIP MCF-7_JQ1 GSE123284.ARID1A.MCF-7_JQ1 112 bp overlap
BRD2 2 datasets
ChIP SK-MEL-147 GSE94488.BRD2.SK-MEL-147 165 bp overlap
ChIP SK-MEL-147_JQ1 GSE94488.BRD2.SK-MEL-147_JQ1 165 bp overlap
BRD4 4 datasets
ChIP HEK293_Ctrl GSE129407.BRD4.HEK293_Ctrl 144 bp overlap
ChIP Kelly_sensitive GSE103030.BRD4.Kelly_sensitive 150 bp overlap
ChIP MDA-MB-231_JQ1-neg_S_3f GSE136151.BRD4.MDA-MB-231_JQ1-neg_S_3f 67 bp overlap
ChIP SUM159PT_scramble GSE131097.BRD4.SUM159PT_scramble 165 bp overlap
BRD7 1 dataset
ChIP SUM159PT_JQ1 GSE131097.BRD7.SUM159PT_JQ1 162 bp overlap
CDK8 1 dataset
ChIP leiomyoma_PT886 GSE128230.CDK8.leiomyoma_PT886 76 bp overlap
CTCF 197 datasets
ChIP A-549 ENCSR000DPF.CTCF.A-549 165 bp overlap
ChIP A-549 ENCSR000AUF.CTCF.A-549 165 bp overlap
ChIP A-549 ENCSR000DNA.CTCF.A-549 116 bp overlap
ChIP A549 ENCFF034FVO 165 bp overlap
ChIP A549 ENCFF434LUY 165 bp overlap
ChIP AG04449 ENCFF248MBD 165 bp overlap
ChIP AG04450 ENCFF116DJL 165 bp overlap
ChIP AG09309 ENCFF478XPS 165 bp overlap
ChIP AG09319 ENCFF401ZTN 165 bp overlap
ChIP AG10803 ENCFF549AQK 165 bp overlap
ChIP BJ ENCFF434HEC 165 bp overlap
ChIP BJ ENCSR000DQI.CTCF.BJ 165 bp overlap
ChIP D54 ENCSR000DKN.CTCF.D54 120 bp overlap
ChIP FT282 GSE131931.CTCF.FT282 165 bp overlap
ChIP GM04025_B-lymphocytes GSE111170.CTCF.GM04025_B-lymphocytes 165 bp overlap
ChIP GM04604 GSE148179.CTCF.GM04604 165 bp overlap
ChIP GM04648 GSE148179.CTCF.GM04648 165 bp overlap
ChIP GM06077 GSE148179.CTCF.GM06077 165 bp overlap
ChIP GM06990 ENCSR000DQW.CTCF.GM06990 145 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 165 bp overlap
ChIP GM12801 ENCSR000DQY.CTCF.GM12801 127 bp overlap
ChIP GM12865 ENCFF067GFI 165 bp overlap
ChIP GM12865 ENCSR000DRE.CTCF.GM12865 144 bp overlap
ChIP GM12866 ENCSR000DRF.CTCF.GM12866 165 bp overlap
ChIP GM12867 ENCSR000DRH.CTCF.GM12867 163 bp overlap
ChIP GM12868 ENCSR000DRI.CTCF.GM12868 165 bp overlap
ChIP GM12869 ENCSR000DRJ.CTCF.GM12869 165 bp overlap
ChIP GM12870 ENCSR000DRK.CTCF.GM12870 165 bp overlap
ChIP GM12871 ENCSR000DRL.CTCF.GM12871 165 bp overlap
ChIP GM12872 ENCFF697BYI 165 bp overlap
ChIP GM12872 ENCSR000DRN.CTCF.GM12872 155 bp overlap
ChIP GM12873 ENCFF711LOS 128 bp overlap
ChIP GM12873 ENCSR000DRP.CTCF.GM12873 165 bp overlap
ChIP GM12874 ENCFF942MTD 165 bp overlap
ChIP GM12874 ENCSR000DRR.CTCF.GM12874 165 bp overlap
ChIP GM12875 ENCFF081UCQ 165 bp overlap
ChIP GM12875 ENCSR000DRU.CTCF.GM12875 165 bp overlap
ChIP GM12878 ENCFF217EAX 165 bp overlap
ChIP GM12878 ENCFF511URZ 165 bp overlap
ChIP GM12878 ENCSR000AKB.CTCF.GM12878 165 bp overlap
ChIP GM12878 ENCSR000DKV.CTCF.GM12878 152 bp overlap
ChIP GM12878 ENCSR000DRZ.CTCF.GM12878 102 bp overlap
ChIP GM17942 GSE76922.CTCF.GM17942 165 bp overlap
ChIP GM23338 ENCFF531QOI 165 bp overlap
ChIP GM23338 ENCFF832KWE 165 bp overlap
ChIP GSC23 GSE139416.CTCF.GSC23 95 bp overlap
ChIP H1 ENCFF230QSV 129 bp overlap
ChIP H1 ENCFF764RHO 165 bp overlap
ChIP H54 ENCFF255TVO 165 bp overlap
ChIP H9 ENCFF152GTF 165 bp overlap
ChIP HAP1_WT GSE126634.CTCF.HAP1_WT 165 bp overlap
ChIP HFF ENCSR000DUM.CTCF.HFF 165 bp overlap
ChIP HFFc6 ENCFF005CJI 165 bp overlap
ChIP IMR-90 ENCFF887MRH 165 bp overlap
ChIP IMR-90 ENCSR000EFI.CTCF.IMR-90 151 bp overlap
ChIP IMR-90_Ctrl GSE125639.CTCF.IMR-90_Ctrl 165 bp overlap
ChIP IMR-90_siRNA GSE125639.CTCF.IMR-90_siRNA 165 bp overlap
ChIP Kelly_sensitive GSE103030.CTCF.Kelly_sensitive 165 bp overlap
ChIP Kelly_shB4-res GSE115248.CTCF.Kelly_shB4-res 120 bp overlap
ChIP Kelly_shLUC-res GSE115248.CTCF.Kelly_shLUC-res 152 bp overlap
ChIP LNCaP_Ctrl GSE125639.CTCF.LNCaP_Ctrl 165 bp overlap
ChIP MCF-7 ENCFF139NQI 165 bp overlap
ChIP MCF-7 ENCFF198DQX 165 bp overlap
ChIP MCF-7 ENCFF414SZG 154 bp overlap
ChIP MCF-7 ENCFF424NQR 139 bp overlap
ChIP MCF-7 ENCFF494VXA 165 bp overlap
ChIP MCF-7 ENCFF844STM 139 bp overlap
ChIP MCF-7 ENCSR000DWH.CTCF.MCF-7 165 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 160 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 127 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 165 bp overlap
ChIP NPC GSE115407.CTCF.NPC 116 bp overlap
ChIP PC-9 ENCFF539ULB 165 bp overlap
ChIP RWPE2 ENCFF911IEE 165 bp overlap
ChIP RWPE2 ENCFF911IEE 165 bp overlap
ChIP SK-MEL-147 GSE94488.CTCF.SK-MEL-147 165 bp overlap
ChIP SK-N-SH ENCSR541AMF.CTCF.SK-N-SH 165 bp overlap
ChIP SLK_Ctrl GSE138105.CTCF.SLK_Ctrl 165 bp overlap
ChIP SLK_RAD21-KD GSE138105.CTCF.SLK_RAD21-KD 165 bp overlap
ChIP THP-1_macrophage_PMA GSE96800.CTCF.THP-1_macrophage_PMA 165 bp overlap
ChIP U2OS_interphase GSE141081.CTCF.U2OS_interphase 157 bp overlap
ChIP VCaP ENCSR265ARE.CTCF.VCaP 165 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 110 bp overlap
ChIP WI-38 ENCSR000DYB.CTCF.WI-38 165 bp overlap
ChIP WI38 ENCFF841AXJ 165 bp overlap
ChIP adrenal gland ENCFF596QXB 165 bp overlap
ChIP adrenal gland ENCFF678WUB 165 bp overlap
ChIP adrenal gland ENCFF886WNR 165 bp overlap
ChIP adrenal-gland ENCSR899JSO.CTCF.adrenal-gland 165 bp overlap
ChIP adrenal-gland ENCSR450BLH.CTCF.adrenal-gland 165 bp overlap
ChIP adrenal-gland ENCSR770IWO.CTCF.adrenal-gland 165 bp overlap
ChIP artery_tibial ENCSR699BEK.CTCF.artery_tibial 165 bp overlap
ChIP ascending-aorta ENCSR846JKO.CTCF.ascending-aorta 165 bp overlap
ChIP astrocyte ENCSR000AOO.CTCF.astrocyte 165 bp overlap
ChIP astrocyte of the cerebellum ENCFF511OCS 165 bp overlap
ChIP astrocyte of the spinal cord ENCFF213GKL 165 bp overlap
ChIP astrocyte_cerebellum ENCSR000DSZ.CTCF.astrocyte_cerebellum 165 bp overlap
ChIP astrocyte_spinal_cord ENCSR000DSU.CTCF.astrocyte_spinal_cord 165 bp overlap
ChIP brain ENCFF163BBN 165 bp overlap
ChIP brain ENCFF685VRG 165 bp overlap
ChIP brain microvascular endothelial cell ENCFF526VHE 154 bp overlap
ChIP cardiac fibroblast ENCFF326EDY 165 bp overlap
ChIP cardiac muscle cell ENCFF728JSA 165 bp overlap
ChIP cardiac muscle cell ENCFF728JSA 58 bp overlap
ChIP cardiac muscle cell ENCFF777TNC 157 bp overlap
ChIP cardiac-muscle ENCSR713SXF.CTCF.cardiac-muscle 165 bp overlap
ChIP cardiac_muscle ENCSR000DTI.CTCF.cardiac_muscle 165 bp overlap
ChIP chondrocyte ENCFF134ORZ 165 bp overlap
ChIP choroid plexus epithelial cell ENCFF407YNR 100 bp overlap
ChIP choroid-plexus_epithelial ENCSR000DTL.CTCF.choroid-plexus_epithelial 165 bp overlap
ChIP colon_transverse ENCSR769WKR.CTCF.colon_transverse 165 bp overlap
ChIP colon_transverse ENCSR236YGF.CTCF.colon_transverse 150 bp overlap
ChIP coronary-artery ENCSR175FLL.CTCF.coronary-artery 165 bp overlap
ChIP endodermal cell ENCFF471YCZ 165 bp overlap
ChIP endothelial cell ENCFF663LIE 140 bp overlap
ChIP endothelial_brain-microvascular ENCSR000DTA.CTCF.endothelial_brain-microvascular 165 bp overlap
ChIP epithelial cell of proximal tubule ENCFF763ZKS 165 bp overlap
ChIP epithelial_kidney ENCSR000DVH.CTCF.epithelial_kidney 165 bp overlap
ChIP epithelial_mammary ENCSR000DUS.CTCF.epithelial_mammary 137 bp overlap
ChIP epithelial_proximal_tubule ENCSR000DXD.CTCF.epithelial_proximal_tubule 165 bp overlap
ChIP epithelial_retinal_pigment ENCSR000DVI.CTCF.epithelial_retinal_pigment 165 bp overlap
ChIP esophagus muscularis mucosa ENCFF421MEH 165 bp overlap
ChIP esophagus muscularis mucosa ENCFF534UGM 124 bp overlap
ChIP esophagus-muscularis-mucosa ENCSR073TPC.CTCF.esophagus-muscularis-mucosa 165 bp overlap
ChIP esophagus_muscularis-mucosa ENCSR559KAB.CTCF.esophagus_muscularis-mucosa 165 bp overlap
ChIP esophagus_muscularis-mucosa ENCSR074SFL.CTCF.esophagus_muscularis-mucosa 123 bp overlap
ChIP fibroblast of dermis ENCFF986DNJ 165 bp overlap
ChIP fibroblast of lung ENCFF084DUH 165 bp overlap
ChIP fibroblast of lung ENCFF356FDN 165 bp overlap
ChIP fibroblast of lung ENCFF505HVQ 165 bp overlap
ChIP fibroblast of mammary gland ENCFF109AZU 165 bp overlap
ChIP fibroblast of pulmonary artery ENCFF742RSV 155 bp overlap
ChIP fibroblast of the aortic adventitia ENCFF639DMR 165 bp overlap
ChIP fibroblast of villous mesenchyme ENCFF345VQO 165 bp overlap
ChIP fibroblast_AORTIC_ADVENTITIA ENCSR000DPY.CTCF.fibroblast_AORTIC_ADVENTITIA 165 bp overlap
ChIP fibroblast_CARDIAC ENCSR000DTF.CTCF.fibroblast_CARDIAC 165 bp overlap
ChIP fibroblast_DERMAL ENCSR000APM.CTCF.fibroblast_DERMAL 165 bp overlap
ChIP fibroblast_FORESKIN ENCSR000DUH.CTCF.fibroblast_FORESKIN 165 bp overlap
ChIP fibroblast_FORESKIN ENCSR000DWQ.CTCF.fibroblast_FORESKIN 165 bp overlap
ChIP fibroblast_GINGIVA ENCSR000DPS.CTCF.fibroblast_GINGIVA 164 bp overlap
ChIP fibroblast_LUNG ENCSR000DPM.CTCF.fibroblast_LUNG 165 bp overlap
ChIP fibroblast_LUNG ENCSR000ANO.CTCF.fibroblast_LUNG 165 bp overlap
ChIP fibroblast_LUNG ENCSR000DWY.CTCF.fibroblast_LUNG 165 bp overlap
ChIP fibroblast_LUNG ENCSR000DVA.CTCF.fibroblast_LUNG 165 bp overlap
ChIP fibroblast_MAMMARY ENCSR000DUU.CTCF.fibroblast_MAMMARY 165 bp overlap
ChIP fibroblast_PEDAL_DIGIT_SKIN ENCSR000DPP.CTCF.fibroblast_PEDAL_DIGIT_SKIN 148 bp overlap
ChIP fibroblast_PULMONARY_ARTERY ENCSR000DUX.CTCF.fibroblast_PULMONARY_ARTERY 165 bp overlap
ChIP fibroblast_SKIN_ABDOMEN ENCSR000DPV.CTCF.fibroblast_SKIN_ABDOMEN 165 bp overlap
ChIP fibroblast_SKIN_LEG ENCSR000DPG.CTCF.fibroblast_SKIN_LEG 165 bp overlap
ChIP fibroblast_VILLOUS_MESENCHYME ENCSR000DVQ.CTCF.fibroblast_VILLOUS_MESENCHYME 165 bp overlap
ChIP foreskin fibroblast ENCFF219EBQ 165 bp overlap
ChIP foreskin fibroblast ENCFF671HLG 165 bp overlap
ChIP gastroesophageal sphincter ENCFF918KPI 165 bp overlap
ChIP gastroesophageal-sphincter ENCSR298ZPF.CTCF.gastroesophageal-sphincter 165 bp overlap
ChIP gastroesophageal-sphincter ENCSR186NVR.CTCF.gastroesophageal-sphincter 165 bp overlap
ChIP gastroesophageal-sphincter ENCSR206ETG.CTCF.gastroesophageal-sphincter 165 bp overlap
ChIP gastroesophageal-sphincter ENCSR661XNQ.CTCF.gastroesophageal-sphincter 159 bp overlap
ChIP glioblastoma_GBM3565 GSE139416.CTCF.glioblastoma_GBM3565 165 bp overlap
ChIP hESC_D0 GSE116862.CTCF.hESC_D0 165 bp overlap
ChIP hESC_D5 GSE116862.CTCF.hESC_D5 165 bp overlap
ChIP hepatocyte ENCSR252QYR.CTCF.hepatocyte 151 bp overlap
ChIP hiPSC_CVI GSE106870.CTCF.hiPSC_CVI 53 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 155 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 146 bp overlap
ChIP keratinocyte ENCSR000ALJ.CTCF.keratinocyte 159 bp overlap
ChIP kidney epithelial cell ENCFF173LWY 165 bp overlap
ChIP lung_left_upper-lobe ENCSR970UZD.CTCF.lung_left_upper-lobe 139 bp overlap
ChIP lymphoblast_mut GSE155324.CTCF.lymphoblast_mut 165 bp overlap
ChIP mammary epithelial cell ENCFF164SPU 165 bp overlap
ChIP mammary epithelial cell ENCFF873ERE 165 bp overlap
ChIP myoblast_skeletal_muscle ENCSR000ANE.CTCF.myoblast_skeletal_muscle 165 bp overlap
ChIP myotube ENCFF981UHL 165 bp overlap
ChIP myotube ENCSR000ANS.CTCF.myotube 165 bp overlap
ChIP nephron ENCFF589HXU 165 bp overlap
ChIP nephron progenitor cell ENCFF455DMI 165 bp overlap
ChIP neural ENCSR822CEA.CTCF.neural 165 bp overlap
ChIP neural progenitor cell ENCFF581WPG 165 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 164 bp overlap
ChIP osteoblast ENCFF491ZJZ 135 bp overlap
ChIP osteoblast ENCSR000APF.CTCF.osteoblast 165 bp overlap
ChIP osteocyte ENCFF929FPD 165 bp overlap
ChIP primary-glioblastoma_G567 GSE121601.CTCF.primary-glioblastoma_G567 158 bp overlap
ChIP progenitor cell of endocrine pancreas ENCFF492KHV 165 bp overlap
ChIP progenitor cell of endocrine pancreas ENCFF492KHV 122 bp overlap
ChIP retina_AB1-FW14 GSE86981.CTCF.retina_AB1-FW14 138 bp overlap
ChIP retina_AB1-FW16 GSE86981.CTCF.retina_AB1-FW16 165 bp overlap
ChIP retinal pigment epithelial cell ENCFF810AAG 165 bp overlap
ChIP skeletal muscle myoblast ENCFF813BQI 165 bp overlap
ChIP smooth muscle cell ENCFF656FBT 165 bp overlap
ChIP tibial-artery ENCSR079YAP.CTCF.tibial-artery 149 bp overlap
ChIP type B pancreatic cell ENCFF910FNQ 165 bp overlap
ChIP type B pancreatic cell ENCFF910FNQ 99 bp overlap
ChIP upper lobe of left lung ENCFF645BXH 165 bp overlap
ChIP uterus ENCFF466ZUR 165 bp overlap
ChIP uterus ENCFF837OEY 165 bp overlap
ChIP uterus ENCSR684PGO.CTCF.uterus 165 bp overlap
ChIP uterus ENCSR392SFJ.CTCF.uterus 165 bp overlap
Cebpa 1 dataset
ChIP BLaER1 ENCFF335XTP 165 bp overlap
EBF1 1 dataset
ChIP GM12878 ENCFF813OXE 165 bp overlap
ELF1 1 dataset
ChIP GM12878 ENCSR000BMB.ELF1.GM12878 130 bp overlap
ESR1 3 datasets
ChIP MCF-7_Veh GSE67295.ESR1.MCF-7_Veh 96 bp overlap
ChIP MCF-7_jc5850 GSE126004.ESR1.MCF-7_jc5850 89 bp overlap
ChIP MCF-7_parental GSE123284.ESR1.MCF-7_parental 112 bp overlap
EZH2 1 dataset
ChIP SK-N-MC ENCFF674XUJ 165 bp overlap
FOXA1 1 dataset
ChIP MCF-7_estrogen_ab2 GSE112969.FOXA1.MCF-7_estrogen_ab2 97 bp overlap
HDAC1 1 dataset
ChIP MCF-7_parental_4-hydroxytamoxifen GSE123284.HDAC1.MCF-7_parental_4-hydroxytamoxifen 61 bp overlap
HIF3A 1 dataset
ChIP Hep-3B2-1-7 GSE129491.HIF3A.Hep-3B2-1-7 165 bp overlap
IRF4 1 dataset
ChIP GM12878 ENCFF769ZDL 165 bp overlap
MCM3 1 dataset
ChIP K-562 ENCSR990AZC.MCM3.K-562 154 bp overlap
MLLT1 1 dataset
ChIP GM12878 ENCSR552XSN.MLLT1.GM12878 165 bp overlap
MXI1 1 dataset
ChIP IMR-90 ENCSR000EFE.MXI1.IMR-90 103 bp overlap
MYCN 1 dataset
ChIP Kelly_sensitive GSE115249.MYCN.Kelly_sensitive 119 bp overlap
POLR2A 2 datasets
ChIP GM12878 ENCFF521FXC 132 bp overlap
ChIP GM18505 ENCFF311CYB 165 bp overlap
RAD21 11 datasets
ChIP A-549 ENCSR000BUC.RAD21.A-549 122 bp overlap
ChIP A549 ENCFF047SFC 165 bp overlap
ChIP GM12878 ENCFF101UQZ 162 bp overlap
ChIP GM12878 ENCSR000BMY.RAD21.GM12878 157 bp overlap
ChIP H1 ENCFF698EWO 165 bp overlap
ChIP IMR-90 ENCFF752PTH 165 bp overlap
ChIP IMR-90 ENCSR000EFJ.RAD21.IMR-90 165 bp overlap
ChIP SLK_CTCF-KD GSE138105.RAD21.SLK_CTCF-KD 165 bp overlap
ChIP SLK_Ctrl GSE138105.RAD21.SLK_Ctrl 165 bp overlap
ChIP lymphoblast GSE155324.RAD21.lymphoblast 165 bp overlap
ChIP lymphoblast_mut GSE155324.RAD21.lymphoblast_mut 165 bp overlap
REST 1 dataset
ChIP WA01 ENCSR000BHM.REST.WA01 110 bp overlap
SIN3A 1 dataset
ChIP A-549 ENCSR513XQX.SIN3A.A-549 113 bp overlap
SMARCB1 2 datasets
ChIP MCF-7 GSE123284.SMARCB1.MCF-7 50 bp overlap
ChIP MCF-7_4-hydroxytamoxifen GSE123284.SMARCB1.MCF-7_4-hydroxytamoxifen 140 bp overlap
SMC3 2 datasets
ChIP IMR-90 ENCFF627LON 165 bp overlap
ChIP IMR-90 ENCSR000HPG.SMC3.IMR-90 162 bp overlap
STAT3 2 datasets
ChIP MCF-7 GSE152203.STAT3.MCF-7 61 bp overlap
ChIP MCF-7_jc5848 GSE126004.STAT3.MCF-7_jc5848 107 bp overlap
TAF1 2 datasets
ChIP H1 ENCFF478SZO 165 bp overlap
ChIP WA01 ENCSR000BHO.TAF1.WA01 131 bp overlap
TBP 1 dataset
ChIP hESC_10h GSE122298.TBP.hESC_10h 87 bp overlap
ZBTB2 1 dataset
ChIP GM12878 GSE97661.ZBTB2.GM12878 124 bp overlap
ZBTB7A 1 dataset
ChIP VCaP_FBS GSE123089.ZBTB7A.VCaP_FBS 165 bp overlap
ZNF121 1 dataset
ChIP WTC11 ENCFF291API 122 bp overlap
ZNF490 1 dataset
ChIP HepG2 ENCFF030RSJ 114 bp overlap