chr9 : 3,893,665 3,894,029
364 bp 22 TFs 1 linked gene
This 364 bp open chromatin element is linked to GLIS3-AS1 and is bound by 22 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
GLIS3-AS1 4.6 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr9:3,888,665 – 3,899,029
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
22 transcription factors
Source
Cell type
ASCL1 1 dataset
ChIP NCI-H889 GSE69394.ASCL1.NCI-H889 159 bp overlap
ATF3 1 dataset
ChIP WA01 ENCSR000BKC.ATF3.WA01 166 bp overlap
BRD4 1 dataset
ChIP CD4_JG1 GSE33281.BRD4.CD4_JG1 112 bp overlap
CREB1 1 dataset
ChIP LNCaP-abl_SHCTR GSE63034.CREB1.LNCaP-abl_SHCTR 78 bp overlap
CREBBP 1 dataset
ChIP MCF-7 ERP000901.CREBBP.MCF-7 220 bp overlap
CTCF 156 datasets
ChIP 22Rv1 ENCFF466OXN 364 bp overlap
ChIP C4-2B ENCFF821XVN 298 bp overlap
ChIP Caco-2 ENCFF934QYS 191 bp overlap
ChIP Caco-2 ENCSR000DQN.CTCF.Caco-2 143 bp overlap
ChIP GM04025_B-lymphocytes GSE111170.CTCF.GM04025_B-lymphocytes 181 bp overlap
ChIP GM04604 GSE148179.CTCF.GM04604 157 bp overlap
ChIP GM04648 GSE148179.CTCF.GM04648 172 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 149 bp overlap
ChIP GM12869 ENCSR000DRJ.CTCF.GM12869 81 bp overlap
ChIP GM12872 ENCSR000DRN.CTCF.GM12872 107 bp overlap
ChIP GM12874 ENCSR000DRR.CTCF.GM12874 102 bp overlap
ChIP GM23338 ENCFF531QOI 280 bp overlap
ChIP GP5D GSE51234.CTCF.GP5D 116 bp overlap
ChIP H1 ENCFF414GZI 179 bp overlap
ChIP H1 ENCFF764RHO 220 bp overlap
ChIP HAP1_WT GSE126634.CTCF.HAP1_WT 190 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_2 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_2 177 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 286 bp overlap
ChIP HCT-116 ENCSR000DTO.CTCF.HCT-116 158 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 190 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.CTCF.HCT-116_RAD21-mAC 202 bp overlap
ChIP HCT116 ENCFF003KHP 275 bp overlap
ChIP HCT116 ENCFF209YMI 208 bp overlap
ChIP HEC-1-B_WT GSE140868.CTCF.HEC-1-B_WT 281 bp overlap
ChIP HFFc6 ENCFF005CJI 344 bp overlap
ChIP HL-60 GSE131577.CTCF.HL-60 140 bp overlap
ChIP HMS001 GSE143026.CTCF.HMS001 91 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 178 bp overlap
ChIP Hep-G2 ENCSR000AMA.CTCF.Hep-G2 135 bp overlap
ChIP Hep-G2 ENCSR000DUG.CTCF.Hep-G2 116 bp overlap
ChIP Hep-G2 ERP000209.CTCF.Hep-G2 131 bp overlap
ChIP HepG2 ENCFF194VBQ 210 bp overlap
ChIP LNCaP ENCSR315NAC.CTCF.LNCaP 226 bp overlap
ChIP LNCaP clone FGC ENCFF519YVI 154 bp overlap
ChIP LNCaP clone FGC ENCFF957KCI 154 bp overlap
ChIP LNCaP-C4-2B ENCSR460LGH.CTCF.LNCaP-C4-2B 364 bp overlap
ChIP LNCaP_Ctrl GSE125639.CTCF.LNCaP_Ctrl 215 bp overlap
ChIP MCF-10A GSE98551.CTCF.MCF-10A 218 bp overlap
ChIP MCF-10AT1 GSE98551.CTCF.MCF-10AT1 206 bp overlap
ChIP MCF-10CA1a GSE98551.CTCF.MCF-10CA1a 185 bp overlap
ChIP MCF-7 ENCFF139NQI 107 bp overlap
ChIP MCF-7 ENCFF162GNE 180 bp overlap
ChIP MCF-7 ENCFF198DQX 102 bp overlap
ChIP MCF-7 ENCFF210JUZ 266 bp overlap
ChIP MCF-7 ENCFF210JUZ 320 bp overlap
ChIP MCF-7 ENCFF414SZG 164 bp overlap
ChIP MCF-7 ENCFF424NQR 153 bp overlap
ChIP MCF-7 ENCFF494VXA 102 bp overlap
ChIP MCF-7 ENCFF844STM 153 bp overlap
ChIP MCF-7 ENCFF954TUV 163 bp overlap
ChIP MCF-7 ENCSR000DWH.CTCF.MCF-7 233 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 273 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 170 bp overlap
ChIP MCF-7 GSE137216.CTCF.MCF-7 152 bp overlap
ChIP MCF-7 ENCSR000DMS.CTCF.MCF-7 144 bp overlap
ChIP MCF-7 ERP000380.CTCF.MCF-7 172 bp overlap
ChIP MCF-7 ENCSR000AHD.CTCF.MCF-7 169 bp overlap
ChIP MCF-7 GSE70764.CTCF.MCF-7 104 bp overlap
ChIP MCF-7L_t0 GSE108787.CTCF.MCF-7L_t0 254 bp overlap
ChIP MCF-7L_t1 GSE108787.CTCF.MCF-7L_t1 229 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 182 bp overlap
ChIP MCF-7_E2 ERP000380.CTCF.MCF-7_E2 168 bp overlap
ChIP MCF-7_TAM ERP000380.CTCF.MCF-7_TAM 139 bp overlap
ChIP MCF-7_TAM ERP000783.CTCF.MCF-7_TAM 112 bp overlap
ChIP MDA-MB-157 GSE116868.CTCF.MDA-MB-157 186 bp overlap
ChIP PC-3 ENCFF487TUI 269 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 279 bp overlap
ChIP Peyer's patch ENCFF701KWW 259 bp overlap
ChIP Peyer's patch ENCFF746TCR 240 bp overlap
ChIP Peyer's patch ENCFF828IDE 219 bp overlap
ChIP Peyers-patch ENCSR375VXU.CTCF.Peyers-patch 160 bp overlap
ChIP Peyers-patch ENCSR419ANE.CTCF.Peyers-patch 232 bp overlap
ChIP Peyers-patch ENCSR799WDT.CTCF.Peyers-patch 173 bp overlap
ChIP RWPE-2 ENCSR856JJB.CTCF.RWPE-2 285 bp overlap
ChIP SK-MEL-147 GSE94488.CTCF.SK-MEL-147 102 bp overlap
ChIP SLK_Ctrl GSE138105.CTCF.SLK_Ctrl 303 bp overlap
ChIP T-47D GSE111923.CTCF.T-47D 278 bp overlap
ChIP T-47D GSE120162.CTCF.T-47D 161 bp overlap
ChIP T-47D_NaCl-1h GSE111923.CTCF.T-47D_NaCl-1h 206 bp overlap
ChIP T-47D_NaCl-7-5min GSE111923.CTCF.T-47D_NaCl-7-5min 178 bp overlap
ChIP T-47D_NaCl-isotonic GSE111923.CTCF.T-47D_NaCl-isotonic 291 bp overlap
ChIP T-47D_serum GSE120162.CTCF.T-47D_serum 122 bp overlap
ChIP VCaP ENCFF858YQT 234 bp overlap
ChIP VCaP ENCFF858YQT 364 bp overlap
ChIP VCaP ENCSR265ARE.CTCF.VCaP 358 bp overlap
ChIP VCaP_ETOH GSE84432.CTCF.VCaP_ETOH 117 bp overlap
ChIP VCaP_R1881 GSE84432.CTCF.VCaP_R1881 154 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 108 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 125 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 160 bp overlap
ChIP WERI-Rb-1 ENCSR000DXW.CTCF.WERI-Rb-1 105 bp overlap
ChIP ZR751 ERP000783.CTCF.ZR751 135 bp overlap
ChIP body of pancreas ENCFF798MEO 218 bp overlap
ChIP bonchial_epithelial ENCSR000DXI.CTCF.bonchial_epithelial 155 bp overlap
ChIP bronchial epithelial cell ENCFF500SEA 208 bp overlap
ChIP cardiac fibroblast ENCFF326EDY 217 bp overlap
ChIP colon_transverse ENCSR833FWC.CTCF.colon_transverse 249 bp overlap
ChIP colon_transverse ENCSR102CSD.CTCF.colon_transverse 243 bp overlap
ChIP colon_transverse ENCSR769WKR.CTCF.colon_transverse 199 bp overlap
ChIP endodermal cell ENCFF471YCZ 228 bp overlap
ChIP epithelial cell of esophagus ENCFF946GGT 194 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 212 bp overlap
ChIP epithelial cell of proximal tubule ENCFF763ZKS 181 bp overlap
ChIP epithelial_esophagus ENCSR000DTR.CTCF.epithelial_esophagus 130 bp overlap
ChIP epithelial_kidney ENCSR000DVH.CTCF.epithelial_kidney 209 bp overlap
ChIP epithelial_mammary ENCSR000ALV.CTCF.epithelial_mammary 133 bp overlap
ChIP epithelial_mammary ENCSR000DUS.CTCF.epithelial_mammary 131 bp overlap
ChIP epithelial_proximal_tubule ENCSR000DXD.CTCF.epithelial_proximal_tubule 171 bp overlap
ChIP esophagus squamous epithelium ENCFF037IYT 251 bp overlap
ChIP esophagus_squamous-epithelium ENCSR450FRI.CTCF.esophagus_squamous-epithelium 163 bp overlap
ChIP fibroblast_CARDIAC ENCSR000DTF.CTCF.fibroblast_CARDIAC 128 bp overlap
ChIP fibroblast_VILLOUS_MESENCHYME ENCSR000DVQ.CTCF.fibroblast_VILLOUS_MESENCHYME 74 bp overlap
ChIP foreskin keratinocyte ENCFF980OWR 241 bp overlap
ChIP hepatocyte ENCSR252QYR.CTCF.hepatocyte 171 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 150 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 156 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 160 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 149 bp overlap
ChIP islet ERP004003.CTCF.islet 200 bp overlap
ChIP keratinocyte ENCSR000DWX.CTCF.keratinocyte 166 bp overlap
ChIP keratinocyte GSE154221.CTCF.keratinocyte 223 bp overlap
ChIP keratinocyte ENCSR000ALJ.CTCF.keratinocyte 178 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 109 bp overlap
ChIP lower lobe of right lung ENCFF092XHT 331 bp overlap
ChIP lung ENCSR463XCZ.CTCF.lung 211 bp overlap
ChIP lung_left_upper-lobe ENCSR964BKO.CTCF.lung_left_upper-lobe 167 bp overlap
ChIP mammary epithelial cell ENCFF164SPU 192 bp overlap
ChIP neural progenitor cell ENCFF581WPG 323 bp overlap
ChIP neural progenitor cell ENCFF581WPG 364 bp overlap
ChIP pancreas ENCSR000DND.CTCF.pancreas 133 bp overlap
ChIP pancreas_body ENCSR408XTO.CTCF.pancreas_body 163 bp overlap
ChIP pancreas_body ENCSR265PFQ.CTCF.pancreas_body 177 bp overlap
ChIP pancreas_body ENCSR484DDO.CTCF.pancreas_body 148 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d0 70 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d6 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d6 199 bp overlap
ChIP progenitor cell of endocrine pancreas ENCFF492KHV 291 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 295 bp overlap
ChIP testis ENCSR981CID.CTCF.testis 160 bp overlap
ChIP thyroid gland ENCFF163TUI 292 bp overlap
ChIP thyroid gland ENCFF204HWS 264 bp overlap
ChIP thyroid gland ENCFF300RYK 278 bp overlap
ChIP thyroid gland ENCFF300RYK 330 bp overlap
ChIP thyroid gland ENCFF631QRY 190 bp overlap
ChIP thyroid gland ENCFF631QRY 309 bp overlap
ChIP thyroid gland ENCFF748ICQ 211 bp overlap
ChIP thyroid-gland ENCSR145HBC.CTCF.thyroid-gland 214 bp overlap
ChIP thyroid-gland ENCSR492ZIW.CTCF.thyroid-gland 298 bp overlap
ChIP thyroid-gland ENCSR033KMZ.CTCF.thyroid-gland 274 bp overlap
ChIP thyroid-gland ENCSR955BIB.CTCF.thyroid-gland 364 bp overlap
ChIP thyroid-gland ENCSR744YJR.CTCF.thyroid-gland 218 bp overlap
ChIP transverse colon ENCFF077CMZ 294 bp overlap
ChIP transverse colon ENCFF594PFO 281 bp overlap
ChIP transverse colon ENCFF749DPF 303 bp overlap
ChIP upper lobe of left lung ENCFF374MAK 270 bp overlap
ChIP upper lobe of left lung ENCFF374MAK 305 bp overlap
ChIP upper lobe of left lung ENCFF654BFF 302 bp overlap
ESR1 12 datasets
ChIP Ishikawa_ETV4-KO1_Mili GSE129803.ESR1.Ishikawa_ETV4-KO1_Mili 226 bp overlap
ChIP Ishikawa_ETV4-KO2_Mili GSE129803.ESR1.Ishikawa_ETV4-KO2_Mili 277 bp overlap
ChIP MCF-7 GSE119057.ESR1.MCF-7 197 bp overlap
ChIP MCF-7_ER_t10 GSE119057.ESR1.MCF-7_ER_t10 216 bp overlap
ChIP MCF-7_ER_t20 GSE119057.ESR1.MCF-7_ER_t20 210 bp overlap
ChIP MCF-7_ER_t30 GSE119057.ESR1.MCF-7_ER_t30 199 bp overlap
ChIP MCF-7_ER_t40 GSE119057.ESR1.MCF-7_ER_t40 214 bp overlap
ChIP MCF-7_ER_t50 GSE119057.ESR1.MCF-7_ER_t50 197 bp overlap
ChIP MCF-7_ER_t60 GSE119057.ESR1.MCF-7_ER_t60 211 bp overlap
ChIP MCF-7_ER_t70 GSE119057.ESR1.MCF-7_ER_t70 203 bp overlap
ChIP MCF-7_ER_t80 GSE119057.ESR1.MCF-7_ER_t80 198 bp overlap
ChIP MCF-7_ER_t90 GSE119057.ESR1.MCF-7_ER_t90 198 bp overlap
FOS 1 dataset
ChIP MCF-7 ENCSR569XNP.FOS.MCF-7 225 bp overlap
HOXB13 1 dataset
ChIP G-401 GSE65381.HOXB13.G-401 333 bp overlap
JUN 1 dataset
ChIP MCF-7_abemaciclib GSE157385.JUN.MCF-7_abemaciclib 241 bp overlap
NKX3-1 1 dataset
ChIP islet ERP004003.NKX3-1.islet 130 bp overlap
NUTM1 1 dataset
ChIP NUT_DMSO GSE133122.NUTM1.NUT_DMSO 161 bp overlap
RAD21 18 datasets
ChIP GP5D GSE51234.RAD21.GP5D 119 bp overlap
ChIP H1 ENCFF698EWO 181 bp overlap
ChIP HCT-116 GSE121355.RAD21.HCT-116 279 bp overlap
ChIP HCT-116 GSE131606.RAD21.HCT-116 201 bp overlap
ChIP HCT-116 ENCSR000BSB.RAD21.HCT-116 125 bp overlap
ChIP HCT116 ENCFF568PEO 196 bp overlap
ChIP HCT116 ENCFF568PEO 263 bp overlap
ChIP Hep-G2 ENCSR000EEG.RAD21.Hep-G2 151 bp overlap
ChIP Ishikawa ENCFF570JVV 173 bp overlap
ChIP Ishikawa ENCSR000BTU.RAD21.Ishikawa 140 bp overlap
ChIP MCF-7 ENCFF694KOM 232 bp overlap
ChIP MCF-7 ENCFF724VCQ 204 bp overlap
ChIP MCF-7 ENCSR703TNG.RAD21.MCF-7 156 bp overlap
ChIP MCF-7 ERP000209.RAD21.MCF-7 203 bp overlap
ChIP MCF-7 GSE72082.RAD21.MCF-7 108 bp overlap
ChIP MCF-7 ENCSR000BTQ.RAD21.MCF-7 128 bp overlap
ChIP MCF-7_E2 ERP000209.RAD21.MCF-7_E2 151 bp overlap
ChIP SLK_Ctrl GSE138105.RAD21.SLK_Ctrl 227 bp overlap
SMC1 1 dataset
ChIP HCT-116_RAD21-mAC GSE104888.SMC1.HCT-116_RAD21-mAC 123 bp overlap
SMC1A 1 dataset
ChIP MCF-7 GSE76893.SMC1A.MCF-7 156 bp overlap
STAG1 2 datasets
ChIP MCF-7 ERP000209.STAG1.MCF-7 157 bp overlap
ChIP MCF-7_E2 ERP000209.STAG1.MCF-7_E2 161 bp overlap
STAT1 1 dataset
ChIP HeLa-S3 ENCSR000EZK.STAT1.HeLa-S3 156 bp overlap
STAT3 7 datasets
ChIP HCC70 GSE152203.STAT3.HCC70 179 bp overlap
ChIP HCC70_EtOH GSE85579.STAT3.HCC70_EtOH 96 bp overlap
ChIP MCF-7_jc5836 GSE126004.STAT3.MCF-7_jc5836 107 bp overlap
ChIP MCF-7_jc5838 GSE126004.STAT3.MCF-7_jc5838 279 bp overlap
ChIP MCF-7_jc5840 GSE126004.STAT3.MCF-7_jc5840 195 bp overlap
ChIP T-47D_JC5056 GSE126004.STAT3.T-47D_JC5056 73 bp overlap
ChIP T-47D_JC5062 GSE126004.STAT3.T-47D_JC5062 128 bp overlap
TFAP4 1 dataset
ChIP DLD-1 GSE46935.TFAP4.DLD-1 157 bp overlap
ZFP64 3 datasets
ChIP HEK293 GSE76494.ZFP64.HEK293 205 bp overlap
ChIP Hep-G2 ENCSR487CPI.ZFP64.Hep-G2 364 bp overlap
ChIP HepG2 ENCFF873EPM 146 bp overlap
ZNF143 2 datasets
ChIP MCF-7 GSE76454.ZNF143.MCF-7 191 bp overlap
ChIP MCF-7_E2 GSE76454.ZNF143.MCF-7_E2 209 bp overlap