chr1 : 220,685,948 220,686,221
273 bp 25 TFs 1 linked gene
This 273 bp open chromatin element is linked to C1orf115 and is bound by 25 transcription factors.
Linked Genes
1 gene
Gene Expression Dist. to TSS Distance Link type
C1orf115 4.1 kb Proximal Proximity
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr1:220,680,948 – 220,691,221
This element Other elements Gene links ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
25 transcription factors
Source
Cell type
BCL11A 1 dataset
ChIP CD34_Day9_90min GSE104676.BCL11A.CD34_Day9_90min 56 bp overlap
BCL6B 1 dataset
Motif DE_36h DE_36h-BCL6B_MA0731.1 17 bp overlap
CTCF 181 datasets
ChIP A-549 ENCSR000AUF.CTCF.A-549 230 bp overlap
ChIP A-549 ENCSR000DPF.CTCF.A-549 246 bp overlap
ChIP A-549 ENCSR000DYD.CTCF.A-549 191 bp overlap
ChIP A-549 ENCSR000DNA.CTCF.A-549 109 bp overlap
ChIP A549 ENCFF034FVO 270 bp overlap
ChIP A549 ENCFF434LUY 245 bp overlap
ChIP C4-2B ENCFF821XVN 273 bp overlap
ChIP Caco-2 ENCFF753NZV 273 bp overlap
ChIP Caco-2 ENCFF934QYS 217 bp overlap
ChIP Caco-2 ENCSR000DQN.CTCF.Caco-2 124 bp overlap
ChIP GM04648 GSE148179.CTCF.GM04648 226 bp overlap
ChIP GM23338 ENCFF531QOI 148 bp overlap
ChIP GM23338 ENCFF772DML 201 bp overlap
ChIP H1 ENCFF230QSV 165 bp overlap
ChIP H1 ENCFF414GZI 217 bp overlap
ChIP H1 ENCFF764RHO 111 bp overlap
ChIP H9 ENCFF152GTF 244 bp overlap
ChIP HAP1_WaplKO-33_SCC4KO GSE94992.CTCF.HAP1_WaplKO-33_SCC4KO 187 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 273 bp overlap
ChIP HCT-116 GSE131606.CTCF.HCT-116 266 bp overlap
ChIP HCT-116 ENCSR000DTO.CTCF.HCT-116 261 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.CTCF.HCT-116_RAD21-mAC 273 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.CTCF.HCT-116_RAD21-mAC_500uM_auxin 247 bp overlap
ChIP HCT116 ENCFF003KHP 144 bp overlap
ChIP HCT116 ENCFF003KHP 135 bp overlap
ChIP HCT116 ENCFF209YMI 268 bp overlap
ChIP HEC-1-B GSE139679.CTCF.HEC-1-B 180 bp overlap
ChIP HEC-1-B_F-insertion GSE140868.CTCF.HEC-1-B_F-insertion 185 bp overlap
ChIP HEC-1-B_R1-insertion GSE140868.CTCF.HEC-1-B_R1-insertion 125 bp overlap
ChIP HEC-1-B_R1-mutation GSE140868.CTCF.HEC-1-B_R1-mutation 75 bp overlap
ChIP HEC-1-B_RF-mutation GSE140868.CTCF.HEC-1-B_RF-mutation 166 bp overlap
ChIP HEC-1-B_WT GSE140868.CTCF.HEC-1-B_WT 229 bp overlap
ChIP HEC-1-B_a12-CBS-deletion GSE140868.CTCF.HEC-1-B_a12-CBS-deletion 126 bp overlap
ChIP HEC-1-B_ac1-CBS-deletion GSE140868.CTCF.HEC-1-B_ac1-CBS-deletion 58 bp overlap
ChIP HEK293 ENCFF498RMM 261 bp overlap
ChIP HEK293 ENCSR617IFZ.CTCF.HEK293 273 bp overlap
ChIP HEK293 ENCSR000DTW.CTCF.HEK293 247 bp overlap
ChIP HFFc6 ENCFF005CJI 273 bp overlap
ChIP HFFc6 ENCFF005CJI 273 bp overlap
ChIP HL-60 GSE131577.CTCF.HL-60 143 bp overlap
ChIP HMS001 GSE143026.CTCF.HMS001 92 bp overlap
ChIP HUDEP-2_30min GSE104676.CTCF.HUDEP-2_30min 95 bp overlap
ChIP HeLa GSE126990.CTCF.HeLa 153 bp overlap
ChIP HeLa-Kyoto GSE138405.CTCF.HeLa-Kyoto 153 bp overlap
ChIP HeLa-Kyoto GSE102884.CTCF.HeLa-Kyoto 158 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 247 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 206 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 270 bp overlap
ChIP HeLa-S3 ENCFF255ASZ 273 bp overlap
ChIP HeLa-S3 ENCFF565UFR 161 bp overlap
ChIP HeLa-S3 ENCFF626XQK 251 bp overlap
ChIP HeLa-S3 ENCSR000AOA.CTCF.HeLa-S3 273 bp overlap
ChIP HeLa-S3 ENCSR000DUB.CTCF.HeLa-S3 223 bp overlap
ChIP HeLa-S3 ENCSR000DLO.CTCF.HeLa-S3 160 bp overlap
ChIP HeLa-S3_biotin GSE108869.CTCF.HeLa-S3_biotin 273 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 273 bp overlap
ChIP HeLa-S3_unsynchro GSE108173.CTCF.HeLa-S3_unsynchro 187 bp overlap
ChIP HeLa_dC9Sun-D3A_CDCC85C GSE107607.CTCF.HeLa_dC9Sun-D3A_CDCC85C 126 bp overlap
ChIP HeLa_dC9Sun-D3A_MIR152 GSE107607.CTCF.HeLa_dC9Sun-D3A_MIR152 136 bp overlap
ChIP Hep-G2 ERP000209.CTCF.Hep-G2 244 bp overlap
ChIP Hep-G2 ENCSR000BIE.CTCF.Hep-G2 240 bp overlap
ChIP Hep-G2 ENCSR000AMA.CTCF.Hep-G2 250 bp overlap
ChIP Hep-G2 ENCSR000DUG.CTCF.Hep-G2 218 bp overlap
ChIP Hep-G2 ENCSR000DLS.CTCF.Hep-G2 120 bp overlap
ChIP HepG2 ENCFF127KUP 231 bp overlap
ChIP HepG2 ENCFF194VBQ 267 bp overlap
ChIP HepG2 ENCFF348BUL 197 bp overlap
ChIP IMR-90_Ctrl GSE125639.CTCF.IMR-90_Ctrl 202 bp overlap
ChIP K-562 ENCSR000BPJ.CTCF.K-562 213 bp overlap
ChIP K-562_CRISPRa_N4293 GSE131349.CTCF.K-562_CRISPRa_N4293 115 bp overlap
ChIP K562 ENCFF111MGE 241 bp overlap
ChIP LNCaP clone FGC ENCFF519YVI 181 bp overlap
ChIP LNCaP-C4-2B ENCSR460LGH.CTCF.LNCaP-C4-2B 273 bp overlap
ChIP LNCaP_Ctrl GSE125639.CTCF.LNCaP_Ctrl 245 bp overlap
ChIP MCF-7 ENCFF139NQI 270 bp overlap
ChIP MCF-7 ENCFF162GNE 128 bp overlap
ChIP MCF-7 ENCFF198DQX 77 bp overlap
ChIP MCF-7 ENCFF210JUZ 167 bp overlap
ChIP MCF-7 ENCFF414SZG 191 bp overlap
ChIP MCF-7 ENCFF424NQR 191 bp overlap
ChIP MCF-7 ENCFF494VXA 77 bp overlap
ChIP MCF-7 ENCFF844STM 191 bp overlap
ChIP MCF-7 ENCFF954TUV 205 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 273 bp overlap
ChIP MCF-7 ENCSR000AHD.CTCF.MCF-7 245 bp overlap
ChIP MCF-7 ENCSR000DWH.CTCF.MCF-7 256 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 245 bp overlap
ChIP MCF-7 ENCSR000DMS.CTCF.MCF-7 211 bp overlap
ChIP MCF-7 ERP000380.CTCF.MCF-7 167 bp overlap
ChIP MCF-7 GSE70764.CTCF.MCF-7 91 bp overlap
ChIP MCF-7L_t0 GSE108787.CTCF.MCF-7L_t0 273 bp overlap
ChIP MCF-7L_t1 GSE108787.CTCF.MCF-7L_t1 273 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 273 bp overlap
ChIP MCF-7_E2 ERP000380.CTCF.MCF-7_E2 156 bp overlap
ChIP MCF-7_TAM ERP000380.CTCF.MCF-7_TAM 186 bp overlap
ChIP MDA-MB-157 GSE116868.CTCF.MDA-MB-157 269 bp overlap
ChIP MM1-S ENCSR402IDP.CTCF.MM1-S 258 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 121 bp overlap
ChIP NCI-H929 ENCSR634OAQ.CTCF.NCI-H929 273 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 189 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 273 bp overlap
ChIP Panc1 ENCFF056JQX 200 bp overlap
ChIP Peyer's patch ENCFF701KWW 264 bp overlap
ChIP Peyer's patch ENCFF746TCR 273 bp overlap
ChIP Peyers-patch ENCSR375VXU.CTCF.Peyers-patch 273 bp overlap
ChIP Peyers-patch ENCSR419ANE.CTCF.Peyers-patch 273 bp overlap
ChIP Peyers-patch ENCSR799WDT.CTCF.Peyers-patch 205 bp overlap
ChIP SK-MEL-147 GSE94488.CTCF.SK-MEL-147 119 bp overlap
ChIP SK-N-AS GSE101295.CTCF.SK-N-AS 233 bp overlap
ChIP SK-N-SH ENCFF575DMG 273 bp overlap
ChIP SK-N-SH ENCSR541AMF.CTCF.SK-N-SH 232 bp overlap
ChIP SK-N-SH ENCSR000DXQ.CTCF.SK-N-SH 116 bp overlap
ChIP SLK_Ctrl GSE138105.CTCF.SLK_Ctrl 273 bp overlap
ChIP SLK_RAD21-KD GSE138105.CTCF.SLK_RAD21-KD 273 bp overlap
ChIP T-47D GSE111923.CTCF.T-47D 273 bp overlap
ChIP T-47D ENCSR000BNO.CTCF.T-47D 212 bp overlap
ChIP T-47D GSE120162.CTCF.T-47D 176 bp overlap
ChIP T-47D_Y537S GSE148277.CTCF.T-47D_Y537S 273 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-000m-Flavo-240m 149 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-020m-Flavo-000m 174 bp overlap
ChIP THP-1_Pam3csk-025m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-025m-Flavo-000m 273 bp overlap
ChIP THP-1_Pam3csk-030m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-030m-Flavo-000m 177 bp overlap
ChIP THP-1_Pam3csk-045m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-045m-Flavo-000m 246 bp overlap
ChIP THP-1_Pam3csk-120m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-120m-Flavo-000m 208 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.CTCF.THP-1_Pam3csk-150m-Flavo-030m 201 bp overlap
ChIP THP-1_siWAPL-NS1-Pam3csk-4h GSE103477.CTCF.THP-1_siWAPL-NS1-Pam3csk-4h 258 bp overlap
ChIP VCaP ENCSR265ARE.CTCF.VCaP 273 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 204 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 131 bp overlap
ChIP WERI-Rb-1 ENCFF349QKF 231 bp overlap
ChIP WERI-Rb-1 ENCSR000DXW.CTCF.WERI-Rb-1 207 bp overlap
ChIP ZR751 ERP000783.CTCF.ZR751 198 bp overlap
ChIP adrenal gland ENCFF678WUB 236 bp overlap
ChIP adrenal-gland ENCSR770IWO.CTCF.adrenal-gland 174 bp overlap
ChIP brain microvascular endothelial cell ENCFF526VHE 262 bp overlap
ChIP chondrocyte ENCFF134ORZ 273 bp overlap
ChIP colon_transverse ENCSR102CSD.CTCF.colon_transverse 273 bp overlap
ChIP colon_transverse ENCSR833FWC.CTCF.colon_transverse 241 bp overlap
ChIP endodermal cell ENCFF471YCZ 273 bp overlap
ChIP endothelial_brain-microvascular ENCSR000DTA.CTCF.endothelial_brain-microvascular 216 bp overlap
ChIP endothelial_umbilical-vein ENCSR000DVP.CTCF.endothelial_umbilical-vein 172 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 273 bp overlap
ChIP epithelial cell of proximal tubule ENCFF763ZKS 273 bp overlap
ChIP epithelial_proximal_tubule ENCSR000DXD.CTCF.epithelial_proximal_tubule 271 bp overlap
ChIP esophagus_squamous-epithelium ENCSR838RUX.CTCF.esophagus_squamous-epithelium 254 bp overlap
ChIP esophagus_squamous-epithelium ENCSR450FRI.CTCF.esophagus_squamous-epithelium 219 bp overlap
ChIP fibroblast of pulmonary artery ENCFF742RSV 264 bp overlap
ChIP fibroblast_CARDIAC ENCSR000DTF.CTCF.fibroblast_CARDIAC 203 bp overlap
ChIP fibroblast_FORESKIN ENCSR000DWQ.CTCF.fibroblast_FORESKIN 140 bp overlap
ChIP fibroblast_PULMONARY_ARTERY ENCSR000DUX.CTCF.fibroblast_PULMONARY_ARTERY 123 bp overlap
ChIP fibroblast_VILLOUS_MESENCHYME ENCSR000DVQ.CTCF.fibroblast_VILLOUS_MESENCHYME 137 bp overlap
ChIP glutamatergic neuron ENCFF816BTR 273 bp overlap
ChIP hESC_D0 GSE116862.CTCF.hESC_D0 216 bp overlap
ChIP hESC_D2 GSE116862.CTCF.hESC_D2 251 bp overlap
ChIP hESC_D5 GSE116862.CTCF.hESC_D5 236 bp overlap
ChIP hepatocyte ERP000395.CTCF.hepatocyte 167 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 125 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 127 bp overlap
ChIP hiPSC_Prader-Willi GSE117283.CTCF.hiPSC_Prader-Willi 140 bp overlap
ChIP keratinocyte ENCSR000DWX.CTCF.keratinocyte 161 bp overlap
ChIP liver ENCFF895ERR 222 bp overlap
ChIP liver ENCSR254YRM.CTCF.liver 213 bp overlap
ChIP liver_right-lobe-of ENCSR911GFJ.CTCF.liver_right-lobe-of 273 bp overlap
ChIP lung_left_upper-lobe ENCSR027FSZ.CTCF.lung_left_upper-lobe 207 bp overlap
ChIP myoblast_skeletal_muscle ENCSR000ANE.CTCF.myoblast_skeletal_muscle 137 bp overlap
ChIP myotube ENCSR000ANS.CTCF.myotube 174 bp overlap
ChIP neural crest cell ENCFF182LWK 273 bp overlap
ChIP neural progenitor cell ENCFF420RBO 273 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 243 bp overlap
ChIP pancreas_body ENCSR265PFQ.CTCF.pancreas_body 193 bp overlap
ChIP placenta ENCFF029PHY 273 bp overlap
ChIP progenitor cell of endocrine pancreas ENCFF492KHV 273 bp overlap
ChIP retinal pigment epithelial cell ENCFF810AAG 257 bp overlap
ChIP right lobe of liver ENCFF956UTA 273 bp overlap
ChIP thyroid gland ENCFF631QRY 217 bp overlap
ChIP thyroid-gland ENCSR492ZIW.CTCF.thyroid-gland 273 bp overlap
ChIP thyroid-gland ENCSR744YJR.CTCF.thyroid-gland 273 bp overlap
ChIP thyroid-gland ENCSR955BIB.CTCF.thyroid-gland 207 bp overlap
ChIP transverse colon ENCFF749DPF 273 bp overlap
ChIP type B pancreatic cell ENCFF910FNQ 273 bp overlap
ChIP upper lobe of left lung ENCFF277NLT 273 bp overlap
CTCF_s 1 dataset
ChIP HeLa-S3_biotin GSE108869.CTCF_s.HeLa-S3_biotin 273 bp overlap
DMRTA2 1 dataset
Motif DE_36h DE_36h-DMRTA2_MA1478.2 6 bp overlap
ELF1 1 dataset
ChIP MCF-7 ENCSR000BSS.ELF1.MCF-7 185 bp overlap
ESR1 10 datasets
ChIP MCF-7 GSE119057.ESR1.MCF-7 238 bp overlap
ChIP MCF-7_ER_t10 GSE119057.ESR1.MCF-7_ER_t10 255 bp overlap
ChIP MCF-7_ER_t20 GSE119057.ESR1.MCF-7_ER_t20 263 bp overlap
ChIP MCF-7_ER_t30 GSE119057.ESR1.MCF-7_ER_t30 255 bp overlap
ChIP MCF-7_ER_t40 GSE119057.ESR1.MCF-7_ER_t40 255 bp overlap
ChIP MCF-7_ER_t50 GSE119057.ESR1.MCF-7_ER_t50 273 bp overlap
ChIP MCF-7_ER_t60 GSE119057.ESR1.MCF-7_ER_t60 252 bp overlap
ChIP MCF-7_ER_t70 GSE119057.ESR1.MCF-7_ER_t70 260 bp overlap
ChIP MCF-7_ER_t80 GSE119057.ESR1.MCF-7_ER_t80 272 bp overlap
ChIP MCF-7_ER_t90 GSE119057.ESR1.MCF-7_ER_t90 265 bp overlap
FOXA1 1 dataset
ChIP T-47D_shRNF2 GSE137579.FOXA1.T-47D_shRNF2 180 bp overlap
MYC 1 dataset
ChIP MCF-7 ENCSR000DMQ.MYC.MCF-7 131 bp overlap
RAD21 17 datasets
ChIP H1 ENCFF698EWO 241 bp overlap
ChIP H1 ENCFF967OJF 245 bp overlap
ChIP HCT-116 GSE121355.RAD21.HCT-116 273 bp overlap
ChIP HEC-1-B_F-insertion GSE140868.RAD21.HEC-1-B_F-insertion 176 bp overlap
ChIP HeLa-S3_CTCF-s-OE GSE108869.RAD21.HeLa-S3_CTCF-s-OE 208 bp overlap
ChIP Hep-G2 ENCSR000EEG.RAD21.Hep-G2 200 bp overlap
ChIP HepG2 ENCFF906QIS 225 bp overlap
ChIP HepG2 ENCFF916QGM 273 bp overlap
ChIP HepG2 ENCFF963UBJ 243 bp overlap
ChIP MCF-7 ENCFF694KOM 273 bp overlap
ChIP MCF-7 ERP000209.RAD21.MCF-7 178 bp overlap
ChIP MCF-7 ENCSR000BTQ.RAD21.MCF-7 151 bp overlap
ChIP MCF-7 GSE72082.RAD21.MCF-7 129 bp overlap
ChIP MCF-7 ENCSR703TNG.RAD21.MCF-7 121 bp overlap
ChIP MCF-7_E2 ERP000209.RAD21.MCF-7_E2 166 bp overlap
ChIP SLK_Ctrl GSE138105.RAD21.SLK_Ctrl 273 bp overlap
ChIP liver ENCFF522JHE 133 bp overlap
SIN3A 1 dataset
ChIP MCF-7 ENCSR468LUO.SIN3A.MCF-7 196 bp overlap
SMC1 1 dataset
ChIP HCT-116_RAD21-mAC GSE104888.SMC1.HCT-116_RAD21-mAC 237 bp overlap
SMC1A 2 datasets
ChIP Hep-G2 GSE76893.SMC1A.Hep-G2 196 bp overlap
ChIP MCF-7 GSE76893.SMC1A.MCF-7 172 bp overlap
SMC3 7 datasets
ChIP HeLa GSE126990.SMC3.HeLa 221 bp overlap
ChIP HeLa-Kyoto GSE102884.SMC3.HeLa-Kyoto 221 bp overlap
ChIP HeLa-Kyoto GSE138405.SMC3.HeLa-Kyoto 221 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.SMC3.HeLa-Kyoto_PDS5-depleted 233 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.SMC3.HeLa-Kyoto_WAPL_PDS-depleted 272 bp overlap
ChIP Hep-G2 ENCSR000EDW.SMC3.Hep-G2 174 bp overlap
ChIP HepG2 ENCFF745UAV 238 bp overlap
STAG1 5 datasets
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 187 bp overlap
ChIP Hep-G2 ERP000209.STAG1.Hep-G2 197 bp overlap
ChIP HepG2 ENCFF843EBZ 261 bp overlap
ChIP MCF-7 ERP000209.STAG1.MCF-7 233 bp overlap
ChIP MCF-7_E2 ERP000209.STAG1.MCF-7_E2 196 bp overlap
Stat5a 1 dataset
Motif DE_36h DE_36h-Stat5a_MA1624.2 9 bp overlap
TP53 1 dataset
ChIP SW480_shp53_16h_TNF-a GSE115985.TP53.SW480_shp53_16h_TNF-a 204 bp overlap
XRCC5 2 datasets
ChIP HepG2 ENCFF330PDO 273 bp overlap
ChIP HepG2 ENCFF680LVJ 273 bp overlap
ZNF12 1 dataset
ChIP Hep-G2 ENCSR711KBM.ZNF12.Hep-G2 151 bp overlap
ZNF143 2 datasets
ChIP Hep-G2 ENCSR101FJT.ZNF143.Hep-G2 118 bp overlap
ChIP WA01 ENCSR000EBW.ZNF143.WA01 131 bp overlap
ZNF41 1 dataset
ChIP HEK293 GSE76494.ZNF41.HEK293 171 bp overlap
ZNF667 1 dataset
Motif DE_36h DE_36h-ZNF667_MA1984.2 11 bp overlap
ZNF93 1 dataset
ChIP HEK293T GSE78099.ZNF93.HEK293T 191 bp overlap