chr11 : 116,215,638 – 116,216,056
418 bp 20 TFs 0 linked genes
This 418 bp open chromatin element has no linked target genes and is bound by 20 transcription factors.
Linked Genes
No linked genes
Chromatin Accessibility

Genome Browser

Genomic view centred on this element. Tracks include ATAC-seq accessibility, RNA-seq expression, TF binding peaks, and element-to-gene loop arcs across the ESC → DE time course.

View limited to ±500 kb around this element.
For the full data, please see the UCSC track linked below.
chr11:116,210,638 – 116,221,056
This element Other elements ATAC-seq RNA-seq

Transcription factors with ChIP-seq peak overlap or TOBIAS motif footprint evidence at this element.

TF Binding
20 transcription factors
Source
Cell type
ARNTL 1 dataset
ChIP U2OS_DMOG GSE85096.ARNTL.U2OS_DMOG 294 bp overlap
ATF2 1 dataset
ChIP WA01 ENCSR000BQU.ATF2.WA01 158 bp overlap
BRD1 1 dataset
ChIP HUES-64 GSE104059.BRD1.HUES-64 229 bp overlap
CREB1 1 dataset
ChIP WA01 ENCSR000BSN.CREB1.WA01 160 bp overlap
CTCF 444 datasets
ChIP 22Rv1 ENCFF466OXN 154 bp overlap
ChIP 22Rv1 ENCSR857PBV.CTCF.22Rv1 350 bp overlap
ChIP 22Rv1_hydroxy_10nM_4h ENCSR847XGE.CTCF.22Rv1_hydroxy_10nM_4h 324 bp overlap
ChIP 786-O_NORMOXIA GSE78113.CTCF.786-O_NORMOXIA 368 bp overlap
ChIP 81-3 ERP002246.CTCF.81-3 147 bp overlap
ChIP A-549 ENCSR000AUF.CTCF.A-549 375 bp overlap
ChIP A-549 ENCSR000AUE.CTCF.A-549 351 bp overlap
ChIP A-549 ENCSR000DPF.CTCF.A-549 203 bp overlap
ChIP A-549 ENCSR000DNA.CTCF.A-549 156 bp overlap
ChIP A-549 ENCSR000BHW.CTCF.A-549 119 bp overlap
ChIP A-673 ENCSR611JJS.CTCF.A-673 363 bp overlap
ChIP A549 ENCFF034FVO 276 bp overlap
ChIP A549 ENCFF182TCQ 217 bp overlap
ChIP A673 ENCFF123WOM 318 bp overlap
ChIP AG09309 ENCFF478XPS 244 bp overlap
ChIP AG10803 ENCFF549AQK 224 bp overlap
ChIP BE2C ENCFF757SRF 286 bp overlap
ChIP BE2C ENCSR000DQD.CTCF.BE2C 368 bp overlap
ChIP BJ ENCFF434HEC 293 bp overlap
ChIP BJ ENCSR000DQI.CTCF.BJ 162 bp overlap
ChIP COLO-829 GSE81945.CTCF.COLO-829 246 bp overlap
ChIP CUTLL1_DMSO GSE130140.CTCF.CUTLL1_DMSO 127 bp overlap
ChIP CUTLL1_GSI GSE130140.CTCF.CUTLL1_GSI 170 bp overlap
ChIP Caco-2 ENCFF753NZV 337 bp overlap
ChIP Caco-2 ENCFF753NZV 78 bp overlap
ChIP Caco-2 ENCFF934QYS 215 bp overlap
ChIP Caco-2 ENCSR000DQN.CTCF.Caco-2 136 bp overlap
ChIP D54 ENCSR000DKN.CTCF.D54 209 bp overlap
ChIP D721Med ENCFF513FYD 211 bp overlap
ChIP DND-41 ENCFF913MRA 158 bp overlap
ChIP DND41 ENCSR000AQU.CTCF.DND41 328 bp overlap
ChIP DOHH2 ENCFF637WNW 222 bp overlap
ChIP DOHH2 ENCFF637WNW 180 bp overlap
ChIP DOHH2 ENCSR084RDK.CTCF.DOHH2 418 bp overlap
ChIP FT282 GSE131931.CTCF.FT282 203 bp overlap
ChIP FT282_EV GSE131931.CTCF.FT282_EV 185 bp overlap
ChIP GM04025_B-lymphocytes GSE111170.CTCF.GM04025_B-lymphocytes 248 bp overlap
ChIP GM04604 GSE148179.CTCF.GM04604 269 bp overlap
ChIP GM04648 GSE148179.CTCF.GM04648 400 bp overlap
ChIP GM06077 GSE148179.CTCF.GM06077 298 bp overlap
ChIP GM06990 ENCFF471OQT 271 bp overlap
ChIP GM06990 ENCSR000DQW.CTCF.GM06990 250 bp overlap
ChIP GM09236__B-lymphocytes GSE111170.CTCF.GM09236__B-lymphocytes 418 bp overlap
ChIP GM09237_B-lymphocytes GSE111170.CTCF.GM09237_B-lymphocytes 351 bp overlap
ChIP GM12864 ENCFF357DQE 250 bp overlap
ChIP GM12864 ENCSR000DRB.CTCF.GM12864 203 bp overlap
ChIP GM12865 ENCFF067GFI 244 bp overlap
ChIP GM12865 ENCSR000DRE.CTCF.GM12865 188 bp overlap
ChIP GM12866 ENCSR000DRF.CTCF.GM12866 176 bp overlap
ChIP GM12867 ENCSR000DRH.CTCF.GM12867 169 bp overlap
ChIP GM12868 ENCSR000DRI.CTCF.GM12868 198 bp overlap
ChIP GM12869 ENCSR000DRJ.CTCF.GM12869 182 bp overlap
ChIP GM12870 ENCSR000DRK.CTCF.GM12870 243 bp overlap
ChIP GM12871 ENCSR000DRL.CTCF.GM12871 192 bp overlap
ChIP GM12872 ENCFF697BYI 247 bp overlap
ChIP GM12872 ENCSR000DRN.CTCF.GM12872 169 bp overlap
ChIP GM12873 ENCFF711LOS 264 bp overlap
ChIP GM12873 ENCSR000DRP.CTCF.GM12873 245 bp overlap
ChIP GM12874 ENCFF942MTD 254 bp overlap
ChIP GM12874 ENCSR000DRR.CTCF.GM12874 209 bp overlap
ChIP GM12875 ENCFF081UCQ 227 bp overlap
ChIP GM12875 ENCSR000DRU.CTCF.GM12875 254 bp overlap
ChIP GM12878 ENCFF485TGR 249 bp overlap
ChIP GM12878 ENCFF511URZ 202 bp overlap
ChIP GM12878 ENCFF635MMB 217 bp overlap
ChIP GM12878 GSE76922.CTCF.GM12878 286 bp overlap
ChIP GM12878 ENCSR000DKV.CTCF.GM12878 192 bp overlap
ChIP GM12878 ERP002246.CTCF.GM12878 154 bp overlap
ChIP GM12878 ENCSR000DRZ.CTCF.GM12878 152 bp overlap
ChIP GM12878 ENCSR000DZN.CTCF.GM12878 112 bp overlap
ChIP GM12878 ENCSR000AKB.CTCF.GM12878 167 bp overlap
ChIP GM13977 ENCFF528ESQ 171 bp overlap
ChIP GM13977 ENCSR000DLB.CTCF.GM13977 127 bp overlap
ChIP GM17942 GSE76922.CTCF.GM17942 327 bp overlap
ChIP GM20000 ENCFF217HWJ 165 bp overlap
ChIP GM20000 ENCFF218HKS 165 bp overlap
ChIP GM20000 ENCSR000DLG.CTCF.GM20000 109 bp overlap
ChIP GM23338 ENCFF531QOI 266 bp overlap
ChIP GM23338 ENCFF772DML 185 bp overlap
ChIP GM23338 ENCFF832KWE 387 bp overlap
ChIP GM23338 ENCFF832KWE 187 bp overlap
ChIP GP5D GSE51234.CTCF.GP5D 371 bp overlap
ChIP GP5D_SIRAD21 GSE51234.CTCF.GP5D_SIRAD21 286 bp overlap
ChIP GSC23 GSE139416.CTCF.GSC23 164 bp overlap
ChIP H1 ENCFF230QSV 165 bp overlap
ChIP H1 ENCFF414GZI 217 bp overlap
ChIP H1 ENCFF764RHO 220 bp overlap
ChIP H54 ENCFF255TVO 225 bp overlap
ChIP H9 ENCFF152GTF 306 bp overlap
ChIP HAP1 GSE94992.CTCF.HAP1 285 bp overlap
ChIP HAP1 GSE152721.CTCF.HAP1 239 bp overlap
ChIP HAP1_2kb GSE152721.CTCF.HAP1_2kb 222 bp overlap
ChIP HAP1_WT GSE126634.CTCF.HAP1_WT 266 bp overlap
ChIP HAP1_WaplKO-33 GSE94992.CTCF.HAP1_WaplKO-33 220 bp overlap
ChIP HAP1_WaplKO-33_SCC4KO GSE94992.CTCF.HAP1_WaplKO-33_SCC4KO 219 bp overlap
ChIP HAP1_clone21 GSE152721.CTCF.HAP1_clone21 239 bp overlap
ChIP HAP1_clone21_CTCF-CRISPR GSE152721.CTCF.HAP1_clone21_CTCF-CRISPR 267 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_1 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_1 253 bp overlap
ChIP HAP1_clone21_CTCF-TSS-CRISPR_2 GSE152721.CTCF.HAP1_clone21_CTCF-TSS-CRISPR_2 251 bp overlap
ChIP HAP1_clone21_TSS-CRISPR GSE152721.CTCF.HAP1_clone21_TSS-CRISPR 280 bp overlap
ChIP HAP1_clone25 GSE152721.CTCF.HAP1_clone25 249 bp overlap
ChIP HCT-116 ENCSR240PRQ.CTCF.HCT-116 350 bp overlap
ChIP HCT-116 GSE92879.CTCF.HCT-116 275 bp overlap
ChIP HCT-116 ENCSR000DTO.CTCF.HCT-116 236 bp overlap
ChIP HCT-116 ENCSR000BSE.CTCF.HCT-116 142 bp overlap
ChIP HCT-116_RAD21-mAC GSE104888.CTCF.HCT-116_RAD21-mAC 278 bp overlap
ChIP HCT-116_RAD21-mAC_500uM_auxin GSE104888.CTCF.HCT-116_RAD21-mAC_500uM_auxin 274 bp overlap
ChIP HCT116 ENCFF003KHP 179 bp overlap
ChIP HCT116 ENCFF209YMI 276 bp overlap
ChIP HEC-1-B_R1-mutation GSE140868.CTCF.HEC-1-B_R1-mutation 112 bp overlap
ChIP HEC-1-B_RF-insertion GSE140868.CTCF.HEC-1-B_RF-insertion 145 bp overlap
ChIP HEC-1-B_RF-mutation GSE140868.CTCF.HEC-1-B_RF-mutation 195 bp overlap
ChIP HEC-1-B_WT GSE140868.CTCF.HEC-1-B_WT 188 bp overlap
ChIP HEK293 ENCFF498RMM 105 bp overlap
ChIP HEK293 ENCSR617IFZ.CTCF.HEK293 213 bp overlap
ChIP HEK293 ENCSR000DTW.CTCF.HEK293 276 bp overlap
ChIP HEK293T GSE115862.CTCF.HEK293T 262 bp overlap
ChIP HFF ENCSR000DUM.CTCF.HFF 233 bp overlap
ChIP HFF-Myc ENCFF680WYR 293 bp overlap
ChIP HL-60 GSE131577.CTCF.HL-60 217 bp overlap
ChIP HL-60 ENCSR000DUP.CTCF.HL-60 163 bp overlap
ChIP HT-1080 GSE135580.CTCF.HT-1080 319 bp overlap
ChIP HT-1080_CBL0137-6h GSE122462.CTCF.HT-1080_CBL0137-6h 391 bp overlap
ChIP HUDEP-2_30min GSE104676.CTCF.HUDEP-2_30min 97 bp overlap
ChIP HUES-64 GSE97394.CTCF.HUES-64 216 bp overlap
ChIP HUES-64_DNMT-KO GSE97394.CTCF.HUES-64_DNMT-KO 300 bp overlap
ChIP HeLa GSE126990.CTCF.HeLa 199 bp overlap
ChIP HeLa-Kyoto GSE138405.CTCF.HeLa-Kyoto 199 bp overlap
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.CTCF.HeLa-Kyoto_PDS5-depleted 195 bp overlap
ChIP HeLa-Kyoto_WAPL-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL-depleted 243 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.CTCF.HeLa-Kyoto_WAPL_PDS-depleted 238 bp overlap
ChIP HeLa-S3 ENCFF565UFR 161 bp overlap
ChIP HeLa-S3 ENCSR000AOA.CTCF.HeLa-S3 272 bp overlap
ChIP HeLa-S3 ENCSR000DUB.CTCF.HeLa-S3 113 bp overlap
ChIP HeLa-S3 ENCSR000DLO.CTCF.HeLa-S3 88 bp overlap
ChIP HeLa-S3_biotin GSE108869.CTCF.HeLa-S3_biotin 172 bp overlap
ChIP HeLa-S3_shCtrl GSE137848.CTCF.HeLa-S3_shCtrl 280 bp overlap
ChIP HeLa-S3_unsynchro GSE108173.CTCF.HeLa-S3_unsynchro 191 bp overlap
ChIP HeLa_dC9Sun-D3AMut_MIR152 GSE107607.CTCF.HeLa_dC9Sun-D3AMut_MIR152 131 bp overlap
ChIP HeLa_dC9Sun-D3A_CDCC85C GSE107607.CTCF.HeLa_dC9Sun-D3A_CDCC85C 192 bp overlap
ChIP Hep-G2 ENCSR000AMA.CTCF.Hep-G2 248 bp overlap
ChIP Hep-G2 ERP000209.CTCF.Hep-G2 203 bp overlap
ChIP Hep-G2 ENCSR000DUG.CTCF.Hep-G2 193 bp overlap
ChIP Hep-G2 ENCSR000BIE.CTCF.Hep-G2 206 bp overlap
ChIP Hep-G2 ENCSR000DLS.CTCF.Hep-G2 151 bp overlap
ChIP HepG2 ENCFF127KUP 231 bp overlap
ChIP HepG2 ENCFF194VBQ 241 bp overlap
ChIP HepG2 ENCFF348BUL 196 bp overlap
ChIP HepG2 ENCFF668CTD 145 bp overlap
ChIP HepG2 ENCFF757EKU 289 bp overlap
ChIP ID00014 GSE76922.CTCF.ID00014 308 bp overlap
ChIP ID00016 GSE76922.CTCF.ID00016 309 bp overlap
ChIP IMR-90 ENCSR000EFI.CTCF.IMR-90 126 bp overlap
ChIP Jurkat GSE68976.CTCF.Jurkat 204 bp overlap
ChIP Jurkat_GSI3d GSE130140.CTCF.Jurkat_GSI3d 170 bp overlap
ChIP K-562 ENCSR000AKO.CTCF.K-562 330 bp overlap
ChIP K-562 GSE70764.CTCF.K-562 278 bp overlap
ChIP K-562 GSE70482.CTCF.K-562 149 bp overlap
ChIP K-562 ENCSR000DMA.CTCF.K-562 209 bp overlap
ChIP K-562 ENCSR000EGM.CTCF.K-562 159 bp overlap
ChIP K-562 ENCSR000DWE.CTCF.K-562 175 bp overlap
ChIP K-562 ENCSR000BPJ.CTCF.K-562 169 bp overlap
ChIP K-562_11698 GSE131349.CTCF.K-562_11698 107 bp overlap
ChIP K-562_13189 GSE131349.CTCF.K-562_13189 103 bp overlap
ChIP K-562_14259 GSE131349.CTCF.K-562_14259 127 bp overlap
ChIP K-562_15173 GSE131349.CTCF.K-562_15173 157 bp overlap
ChIP K-562_15776 GSE131349.CTCF.K-562_15776 121 bp overlap
ChIP K-562_7140 GSE131349.CTCF.K-562_7140 145 bp overlap
ChIP K-562_8005 GSE131349.CTCF.K-562_8005 112 bp overlap
ChIP K-562_CRISPRa_N4293 GSE131349.CTCF.K-562_CRISPRa_N4293 180 bp overlap
ChIP K-562_MonoC GSE137216.CTCF.K-562_MonoC 354 bp overlap
ChIP K-562_MonoN GSE137216.CTCF.K-562_MonoN 216 bp overlap
ChIP K-562_RF2-insertion GSE140868.CTCF.K-562_RF2-insertion 180 bp overlap
ChIP K-562_RabbitC GSE137216.CTCF.K-562_RabbitC 322 bp overlap
ChIP K-562_WT GSE140868.CTCF.K-562_WT 130 bp overlap
ChIP K-562_enCRISPRi-KL GSE132212.CTCF.K-562_enCRISPRi-KL 265 bp overlap
ChIP K562 ENCFF082GOI 185 bp overlap
ChIP K562 ENCFF111MGE 208 bp overlap
ChIP K562 ENCFF400DFR 225 bp overlap
ChIP K562 ENCFF430KTH 135 bp overlap
ChIP K562 ENCFF598YSU 240 bp overlap
ChIP KARPAS422 ENCSR113REG.CTCF.KARPAS422 296 bp overlap
ChIP KMS-11 ENCFF853JKX 403 bp overlap
ChIP Kelly_resistant GSE103030.CTCF.Kelly_resistant 244 bp overlap
ChIP Kelly_resistant_JQ1 GSE103030.CTCF.Kelly_resistant_JQ1 203 bp overlap
ChIP Kelly_sensitive GSE103030.CTCF.Kelly_sensitive 197 bp overlap
ChIP Kelly_shB4-res GSE115248.CTCF.Kelly_shB4-res 162 bp overlap
ChIP Kelly_shLUC-res GSE115248.CTCF.Kelly_shLUC-res 186 bp overlap
ChIP LNCAP ENCFF223HIG 339 bp overlap
ChIP LNCAP ENCFF700QXT 343 bp overlap
ChIP LNCaP ENCSR315NAC.CTCF.LNCaP 248 bp overlap
ChIP LNCaP ENCSR000DMF.CTCF.LNCaP 98 bp overlap
ChIP LNCaP clone FGC ENCFF519YVI 181 bp overlap
ChIP LNCaP clone FGC ENCFF957KCI 181 bp overlap
ChIP LNCaP-C4-2B ENCSR460LGH.CTCF.LNCaP-C4-2B 327 bp overlap
ChIP LNCaP_Ctrl GSE125639.CTCF.LNCaP_Ctrl 226 bp overlap
ChIP Loucy ENCFF359TVQ 339 bp overlap
ChIP Loucy ENCSR464DKE.CTCF.Loucy 272 bp overlap
ChIP MCF-10AT1 GSE98551.CTCF.MCF-10AT1 224 bp overlap
ChIP MCF-7 ENCFF139NQI 245 bp overlap
ChIP MCF-7 ENCFF162GNE 235 bp overlap
ChIP MCF-7 ENCFF198DQX 109 bp overlap
ChIP MCF-7 ENCFF210JUZ 126 bp overlap
ChIP MCF-7 ENCFF414SZG 191 bp overlap
ChIP MCF-7 ENCFF424NQR 191 bp overlap
ChIP MCF-7 ENCFF494VXA 109 bp overlap
ChIP MCF-7 ENCFF844STM 191 bp overlap
ChIP MCF-7 ENCFF954TUV 205 bp overlap
ChIP MCF-7 ENCSR560BUE.CTCF.MCF-7 336 bp overlap
ChIP MCF-7 GSE137216.CTCF.MCF-7 232 bp overlap
ChIP MCF-7 ENCSR000DMR.CTCF.MCF-7 226 bp overlap
ChIP MCF-7 ENCSR000DWH.CTCF.MCF-7 217 bp overlap
ChIP MCF-7 ENCSR000AHD.CTCF.MCF-7 212 bp overlap
ChIP MCF-7 ENCSR000DMS.CTCF.MCF-7 175 bp overlap
ChIP MCF-7 ERP000380.CTCF.MCF-7 174 bp overlap
ChIP MCF-7 GSE70764.CTCF.MCF-7 162 bp overlap
ChIP MCF-7L_t0 GSE108787.CTCF.MCF-7L_t0 249 bp overlap
ChIP MCF-7L_t1 GSE108787.CTCF.MCF-7L_t1 275 bp overlap
ChIP MCF-7L_t16 GSE108787.CTCF.MCF-7L_t16 220 bp overlap
ChIP MCF-7L_t24 GSE108787.CTCF.MCF-7L_t24 292 bp overlap
ChIP MCF-7_E2 ERP000380.CTCF.MCF-7_E2 202 bp overlap
ChIP MCF-7_HYPOXIA GSE78113.CTCF.MCF-7_HYPOXIA 294 bp overlap
ChIP MCF-7_NORMOXIA GSE78113.CTCF.MCF-7_NORMOXIA 311 bp overlap
ChIP MCF-7_TAM ERP000380.CTCF.MCF-7_TAM 192 bp overlap
ChIP MCF-7_TAM ERP000783.CTCF.MCF-7_TAM 132 bp overlap
ChIP MDM_IFNb GSE103477.CTCF.MDM_IFNb 150 bp overlap
ChIP MM.1S ENCFF869JMQ 320 bp overlap
ChIP MM1-S ENCSR402IDP.CTCF.MM1-S 269 bp overlap
ChIP MM1-S GSE43743.CTCF.MM1-S 175 bp overlap
ChIP NB4 ENCFF155DNY 225 bp overlap
ChIP NB4 ENCSR000DWN.CTCF.NB4 209 bp overlap
ChIP NB69 GSE101295.CTCF.NB69 341 bp overlap
ChIP NCI-H929 ENCFF305JAB 202 bp overlap
ChIP NCI-H929 ENCFF305JAB 154 bp overlap
ChIP NCI-H929 ENCSR634OAQ.CTCF.NCI-H929 418 bp overlap
ChIP OCI-LY3 ENCFF939BYJ 208 bp overlap
ChIP OCI-LY7 ENCFF086AXQ 183 bp overlap
ChIP OCI-Ly1 ENCSR072EUE.CTCF.OCI-Ly1 293 bp overlap
ChIP OCI-Ly3 ENCSR756ZKG.CTCF.OCI-Ly3 341 bp overlap
ChIP OCI-Ly7 ENCSR027HML.CTCF.OCI-Ly7 332 bp overlap
ChIP OVCAR-8 GSE70764.CTCF.OVCAR-8 209 bp overlap
ChIP PC-3 ENCFF487TUI 368 bp overlap
ChIP PC-3 ENCSR359LOD.CTCF.PC-3 281 bp overlap
ChIP Peyers-patch ENCSR375VXU.CTCF.Peyers-patch 253 bp overlap
ChIP Peyers-patch ENCSR799WDT.CTCF.Peyers-patch 176 bp overlap
ChIP RH4 GSE83726.CTCF.RH4 249 bp overlap
ChIP RWPE-2 ENCSR856JJB.CTCF.RWPE-2 282 bp overlap
ChIP RWPE2 ENCFF911IEE 418 bp overlap
ChIP SH-SY5Y GSE101295.CTCF.SH-SY5Y 217 bp overlap
ChIP SK-N-SH ENCFF575DMG 169 bp overlap
ChIP SK-N-SH ENCFF575DMG 153 bp overlap
ChIP SK-N-SH ENCFF731NJX 224 bp overlap
ChIP SK-N-SH ENCSR541AMF.CTCF.SK-N-SH 312 bp overlap
ChIP SK-N-SH GSE76815.CTCF.SK-N-SH 261 bp overlap
ChIP SK-N-SH ENCSR000DXQ.CTCF.SK-N-SH 237 bp overlap
ChIP SK-N-SH ENCSR000EIC.CTCF.SK-N-SH 211 bp overlap
ChIP SK-N-SH ENCSR000BLX.CTCF.SK-N-SH 168 bp overlap
ChIP SLK_Ctrl GSE138105.CTCF.SLK_Ctrl 345 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 213 bp overlap
ChIP SU-DHL-6 ENCFF116KKR 180 bp overlap
ChIP SU-DHL-6 ENCSR125DKL.CTCF.SU-DHL-6 418 bp overlap
ChIP T-47D GSE111923.CTCF.T-47D 262 bp overlap
ChIP T-47D GSE120162.CTCF.T-47D 132 bp overlap
ChIP T-47D_NaCl-isotonic GSE111923.CTCF.T-47D_NaCl-isotonic 254 bp overlap
ChIP T-47D_NaCl-isotonic-triptolide GSE111923.CTCF.T-47D_NaCl-isotonic-triptolide 253 bp overlap
ChIP T-47D_triptolide GSE111923.CTCF.T-47D_triptolide 289 bp overlap
ChIP TALL-1 GSE115893.CTCF.TALL-1 269 bp overlap
ChIP THP-1_PMA_Dex-6h GSE103477.CTCF.THP-1_PMA_Dex-6h 201 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-000m-Flavo-000m 318 bp overlap
ChIP THP-1_Pam3csk-000m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-000m-Flavo-240m 303 bp overlap
ChIP THP-1_Pam3csk-020m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-020m-Flavo-000m 291 bp overlap
ChIP THP-1_Pam3csk-025m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-025m-Flavo-000m 262 bp overlap
ChIP THP-1_Pam3csk-030m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-030m-Flavo-000m 211 bp overlap
ChIP THP-1_Pam3csk-045m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-045m-Flavo-000m 325 bp overlap
ChIP THP-1_Pam3csk-060m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-060m-Flavo-000m 277 bp overlap
ChIP THP-1_Pam3csk-120m-Flavo-000m GSE103477.CTCF.THP-1_Pam3csk-120m-Flavo-000m 229 bp overlap
ChIP THP-1_Pam3csk-150m-Flavo-030m GSE103477.CTCF.THP-1_Pam3csk-150m-Flavo-030m 252 bp overlap
ChIP THP-1_Pam3csk-180m-Flavo-060m GSE103477.CTCF.THP-1_Pam3csk-180m-Flavo-060m 295 bp overlap
ChIP THP-1_Pam3csk-360m-Flavo-240m GSE103477.CTCF.THP-1_Pam3csk-360m-Flavo-240m 350 bp overlap
ChIP THP-1_siCtrl-NS1-Pam3csk-0h GSE103477.CTCF.THP-1_siCtrl-NS1-Pam3csk-0h 207 bp overlap
ChIP THP-1_siCtrl-NS1-Pam3csk-4h GSE103477.CTCF.THP-1_siCtrl-NS1-Pam3csk-4h 230 bp overlap
ChIP THP-1_siCtrl-eGFP-Pam3csk-0h GSE103477.CTCF.THP-1_siCtrl-eGFP-Pam3csk-0h 349 bp overlap
ChIP THP-1_siCtrl-eGFP-Pam3csk-4h GSE103477.CTCF.THP-1_siCtrl-eGFP-Pam3csk-4h 346 bp overlap
ChIP THP-1_siNIPBL-NS1-Pam3csk-4h GSE103477.CTCF.THP-1_siNIPBL-NS1-Pam3csk-4h 341 bp overlap
ChIP THP-1_siNIPBL-eGFP-Pam3csk-4h GSE103477.CTCF.THP-1_siNIPBL-eGFP-Pam3csk-4h 193 bp overlap
ChIP THP-1_siWAPL-NS1-Pam3csk-4h GSE103477.CTCF.THP-1_siWAPL-NS1-Pam3csk-4h 246 bp overlap
ChIP THP-1_siWAPL-eGFP-Pam3csk-4h GSE103477.CTCF.THP-1_siWAPL-eGFP-Pam3csk-4h 319 bp overlap
ChIP U2OS_interphase GSE141081.CTCF.U2OS_interphase 236 bp overlap
ChIP VCaP ENCFF858YQT 402 bp overlap
ChIP VCaP ENCSR265ARE.CTCF.VCaP 362 bp overlap
ChIP WA01 ENCSR000BNH.CTCF.WA01 204 bp overlap
ChIP WA01 ENCSR000DLK.CTCF.WA01 196 bp overlap
ChIP WA01_NEUR GSE52457.CTCF.WA01_NEUR 250 bp overlap
ChIP WERI-Rb-1 ENCFF349QKF 150 bp overlap
ChIP WERI-Rb-1 ENCSR000DXW.CTCF.WERI-Rb-1 253 bp overlap
ChIP WTC11 ENCFF658QVH 371 bp overlap
ChIP ZR751 ERP000783.CTCF.ZR751 103 bp overlap
ChIP activated CD8-positive, alpha-beta T cell ENCFF006MHW 418 bp overlap
ChIP astrocyte ENCFF042YJV 342 bp overlap
ChIP astrocyte ENCSR000AOO.CTCF.astrocyte 213 bp overlap
ChIP astrocyte of the spinal cord ENCFF213GKL 241 bp overlap
ChIP astrocyte_cerebellum ENCSR000DSZ.CTCF.astrocyte_cerebellum 225 bp overlap
ChIP astrocyte_spinal_cord ENCSR000DSU.CTCF.astrocyte_spinal_cord 219 bp overlap
ChIP body of pancreas ENCFF798MEO 230 bp overlap
ChIP bonchial_epithelial ENCSR000DXI.CTCF.bonchial_epithelial 267 bp overlap
ChIP brain ENCFF163BBN 390 bp overlap
ChIP brain ENCFF685VRG 418 bp overlap
ChIP brain microvascular endothelial cell ENCFF526VHE 239 bp overlap
ChIP bronchial epithelial cell ENCFF500SEA 260 bp overlap
ChIP cardiac muscle cell ENCFF728JSA 323 bp overlap
ChIP cardiac muscle cell ENCFF777TNC 157 bp overlap
ChIP cardiac-muscle ENCSR713SXF.CTCF.cardiac-muscle 314 bp overlap
ChIP cardiac_muscle ENCSR000DTI.CTCF.cardiac_muscle 235 bp overlap
ChIP chondrocyte ENCFF134ORZ 204 bp overlap
ChIP choroid-plexus_epithelial ENCSR000DTL.CTCF.choroid-plexus_epithelial 150 bp overlap
ChIP cortical-interneuron_KCl-neg_JQ1-neg GSE117508.CTCF.cortical-interneuron_KCl-neg_JQ1-neg 284 bp overlap
ChIP cortical-interneuron_KCl-neg_JQ1-pos GSE117508.CTCF.cortical-interneuron_KCl-neg_JQ1-pos 204 bp overlap
ChIP cortical-interneuron_KCl-pos_JQ1-neg GSE117508.CTCF.cortical-interneuron_KCl-pos_JQ1-neg 235 bp overlap
ChIP cortical-interneuron_KCl-pos_JQ1-pos GSE117508.CTCF.cortical-interneuron_KCl-pos_JQ1-pos 177 bp overlap
ChIP delta-47 GSE70764.CTCF.delta-47 285 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF277CZQ 297 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF373BUI 301 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF403LNW 368 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF628TCI 364 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF628TCI 162 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF631JNO 331 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF756TDJ 332 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF835ZSJ 372 bp overlap
ChIP dorsolateral prefrontal cortex ENCFF883PFA 348 bp overlap
ChIP endodermal cell ENCFF471YCZ 309 bp overlap
ChIP endothelial cell ENCFF663LIE 418 bp overlap
ChIP endothelial cell ENCFF663LIE 290 bp overlap
ChIP endothelial cell of umbilical vein ENCFF455OQM 197 bp overlap
ChIP endothelial_brain-microvascular ENCSR000DTA.CTCF.endothelial_brain-microvascular 185 bp overlap
ChIP endothelial_umbilical-vein ENCSR000ALA.CTCF.endothelial_umbilical-vein 194 bp overlap
ChIP endothelial_umbilical-vein ENCSR000DLW.CTCF.endothelial_umbilical-vein 146 bp overlap
ChIP endothelial_umbilical-vein ENCSR000DVP.CTCF.endothelial_umbilical-vein 121 bp overlap
ChIP epithelial cell of prostate ENCFF086GTI 329 bp overlap
ChIP epithelial cell of proximal tubule ENCFF763ZKS 191 bp overlap
ChIP epithelial_kidney ENCSR000DVH.CTCF.epithelial_kidney 257 bp overlap
ChIP epithelial_mammary ENCSR000ALV.CTCF.epithelial_mammary 210 bp overlap
ChIP epithelial_mammary ENCSR000DUS.CTCF.epithelial_mammary 181 bp overlap
ChIP epithelial_proximal_tubule ENCSR000DXD.CTCF.epithelial_proximal_tubule 319 bp overlap
ChIP epithelial_retinal_pigment ENCSR000DVI.CTCF.epithelial_retinal_pigment 195 bp overlap
ChIP erythroblast_adult_erythroblasts GSE102184.CTCF.erythroblast_adult_erythroblasts 237 bp overlap
ChIP erythroid_Don001 GSE137982.CTCF.erythroid_Don001 199 bp overlap
ChIP esophagus squamous epithelium ENCFF037IYT 280 bp overlap
ChIP esophagus_squamous-epithelium ENCSR450FRI.CTCF.esophagus_squamous-epithelium 200 bp overlap
ChIP fetal_erythroblasts GSE102184.CTCF.fetal_erythroblasts 221 bp overlap
ChIP fibroblast_FORESKIN ENCSR000DUH.CTCF.fibroblast_FORESKIN 158 bp overlap
ChIP fibroblast_VILLOUS_MESENCHYME ENCSR000DVQ.CTCF.fibroblast_VILLOUS_MESENCHYME 137 bp overlap
ChIP foreskin keratinocyte ENCFF980OWR 225 bp overlap
ChIP glutamatergic neuron ENCFF816BTR 418 bp overlap
ChIP hESC GSE20650.CTCF.hESC 129 bp overlap
ChIP hESC_D0 GSE116862.CTCF.hESC_D0 243 bp overlap
ChIP hESC_D15 GSE116862.CTCF.hESC_D15 293 bp overlap
ChIP hESC_D2 GSE116862.CTCF.hESC_D2 352 bp overlap
ChIP hESC_D5 GSE116862.CTCF.hESC_D5 418 bp overlap
ChIP hESC_D7 GSE116862.CTCF.hESC_D7 246 bp overlap
ChIP hESC_NAIVE GSE69646.CTCF.hESC_NAIVE 287 bp overlap
ChIP hESC_PRIMED GSE69646.CTCF.hESC_PRIMED 218 bp overlap
ChIP heart left ventricle ENCFF244ZHV 308 bp overlap
ChIP heart left ventricle ENCFF354HOQ 331 bp overlap
ChIP heart left ventricle ENCFF548XHH 320 bp overlap
ChIP heart left ventricle ENCFF548XHH 73 bp overlap
ChIP heart right ventricle ENCFF577TID 272 bp overlap
ChIP heart right ventricle ENCFF767XJQ 329 bp overlap
ChIP heart_left-ventricle ENCSR544APK.CTCF.heart_left-ventricle 201 bp overlap
ChIP hiPSC_Angelman GSE117283.CTCF.hiPSC_Angelman 294 bp overlap
ChIP hiPSC_CVI GSE106870.CTCF.hiPSC_CVI 244 bp overlap
ChIP hiPSC_HUES9 GSE106870.CTCF.hiPSC_HUES9 179 bp overlap
ChIP hiPSC_IB12 GSE106870.CTCF.hiPSC_IB12 207 bp overlap
ChIP hiPSC_IB7 GSE106870.CTCF.hiPSC_IB7 213 bp overlap
ChIP hiPSC_IIA11 GSE106870.CTCF.hiPSC_IIA11 256 bp overlap
ChIP hiPSC_IID12 GSE106870.CTCF.hiPSC_IID12 246 bp overlap
ChIP hiPSC_IIIB12 GSE106870.CTCF.hiPSC_IIIB12 130 bp overlap
ChIP hiPSC_IV8 GSE106870.CTCF.hiPSC_IV8 245 bp overlap
ChIP hiPSC_Prader-Willi GSE117283.CTCF.hiPSC_Prader-Willi 243 bp overlap
ChIP hiPSC_failed-genome-editing GSE106870.CTCF.hiPSC_failed-genome-editing 256 bp overlap
ChIP hiPSC_parental-line GSE106870.CTCF.hiPSC_parental-line 231 bp overlap
ChIP islet ERP004003.CTCF.islet 234 bp overlap
ChIP keratinocyte ENCFF046PBT 128 bp overlap
ChIP keratinocyte ENCFF291YDC 124 bp overlap
ChIP keratinocyte ENCFF667ULX 281 bp overlap
ChIP keratinocyte ENCFF805QIE 276 bp overlap
ChIP keratinocyte ENCSR000ALJ.CTCF.keratinocyte 271 bp overlap
ChIP keratinocyte ENCSR000DWX.CTCF.keratinocyte 239 bp overlap
ChIP keratinocyte ENCSR000DNC.CTCF.keratinocyte 216 bp overlap
ChIP kidney ENCFF335EKK 163 bp overlap
ChIP kidney epithelial cell ENCFF173LWY 260 bp overlap
ChIP left ventricle myocardium inferior ENCFF161DPW 367 bp overlap
ChIP leukemia GSE142161.CTCF.leukemia 255 bp overlap
ChIP leukemia_CEBPA-24h GSE142161.CTCF.leukemia_CEBPA-24h 192 bp overlap
ChIP leukemia_CEBPA-96h GSE142161.CTCF.leukemia_CEBPA-96h 248 bp overlap
ChIP liver ENCFF895ERR 116 bp overlap
ChIP lower lobe of left lung ENCFF906NCV 320 bp overlap
ChIP lower lobe of right lung ENCFF092XHT 351 bp overlap
ChIP lung_left_upper-lobe ENCSR799TJD.CTCF.lung_left_upper-lobe 220 bp overlap
ChIP lymphoblast GSE155324.CTCF.lymphoblast 418 bp overlap
ChIP lymphoblast_mut GSE155324.CTCF.lymphoblast_mut 418 bp overlap
ChIP mammary epithelial cell ENCFF164SPU 243 bp overlap
ChIP mammary epithelial cell ENCFF873ERE 243 bp overlap
ChIP medulloblastoma ENCSR000DMY.CTCF.medulloblastoma 139 bp overlap
ChIP mesothelial cell of epicardium ENCFF427RFE 336 bp overlap
ChIP myoblast_skeletal_muscle ENCSR000ANE.CTCF.myoblast_skeletal_muscle 140 bp overlap
ChIP nephron ENCFF411ACD 338 bp overlap
ChIP nephron ENCFF411ACD 138 bp overlap
ChIP nephron ENCFF589HXU 389 bp overlap
ChIP nephron ENCFF972IQB 349 bp overlap
ChIP nephron progenitor cell ENCFF455DMI 353 bp overlap
ChIP neural ENCSR822CEA.CTCF.neural 365 bp overlap
ChIP neural cell ENCFF335ADI 129 bp overlap
ChIP neural crest cell ENCFF182LWK 345 bp overlap
ChIP neural progenitor cell ENCFF420RBO 259 bp overlap
ChIP neural_progenitor ENCSR125NBL.CTCF.neural_progenitor 418 bp overlap
ChIP neuroblastoma GSE115862.CTCF.neuroblastoma 210 bp overlap
ChIP neuron_bipolar_doxy_4d ENCSR619IUE.CTCF.neuron_bipolar_doxy_4d 257 bp overlap
ChIP osteoblast ENCFF491ZJZ 306 bp overlap
ChIP osteoblast ENCSR000APF.CTCF.osteoblast 273 bp overlap
ChIP osteocyte ENCFF929FPD 351 bp overlap
ChIP pancreas ENCSR000DND.CTCF.pancreas 167 bp overlap
ChIP pancreas_body ENCSR408XTO.CTCF.pancreas_body 170 bp overlap
ChIP placenta ENCFF029PHY 335 bp overlap
ChIP plasmablast GSE142493.CTCF.plasmablast 138 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d0 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d0 238 bp overlap
ChIP primary-epidermal-keratinocyte_diff_d6 GSE84657.CTCF.primary-epidermal-keratinocyte_diff_d6 230 bp overlap
ChIP primary-glioblastoma_G523 GSE121601.CTCF.primary-glioblastoma_G523 236 bp overlap
ChIP primary-glioblastoma_G583 GSE121601.CTCF.primary-glioblastoma_G583 229 bp overlap
ChIP progenitor cell of endocrine pancreas ENCFF492KHV 221 bp overlap
ChIP prostate_epithelial ENCSR196HOM.CTCF.prostate_epithelial 418 bp overlap
ChIP retina_AB1-FW14 GSE86981.CTCF.retina_AB1-FW14 312 bp overlap
ChIP retina_AB1-FW16 GSE86981.CTCF.retina_AB1-FW16 355 bp overlap
ChIP retina_AB1-FW23 GSE86981.CTCF.retina_AB1-FW23 270 bp overlap
ChIP retina_AB1-RB GSE86981.CTCF.retina_AB1-RB 266 bp overlap
ChIP retinal pigment epithelial cell ENCFF810AAG 245 bp overlap
ChIP right lobe of liver ENCFF011NDG 343 bp overlap
ChIP skeletal muscle myoblast ENCFF813BQI 247 bp overlap
ChIP smooth-muscle-cell ENCSR261VAS.CTCF.smooth-muscle-cell 217 bp overlap
ChIP transverse colon ENCFF653EYS 315 bp overlap
ChIP type B pancreatic cell ENCFF910FNQ 198 bp overlap
ChIP type B pancreatic cell ENCFF910FNQ 178 bp overlap
ChIP upper lobe of left lung ENCFF108BCY 322 bp overlap
ChIP upper lobe of left lung ENCFF277NLT 298 bp overlap
ChIP upper lobe of right lung ENCFF065JCM 327 bp overlap
E2F6 2 datasets
ChIP H1 ENCFF785DWK 403 bp overlap
ChIP WA01 ENCSR000BSI.E2F6.WA01 186 bp overlap
ESR1 10 datasets
ChIP MCF-7 GSE119057.ESR1.MCF-7 257 bp overlap
ChIP MCF-7_ER_t10 GSE119057.ESR1.MCF-7_ER_t10 263 bp overlap
ChIP MCF-7_ER_t20 GSE119057.ESR1.MCF-7_ER_t20 237 bp overlap
ChIP MCF-7_ER_t30 GSE119057.ESR1.MCF-7_ER_t30 249 bp overlap
ChIP MCF-7_ER_t40 GSE119057.ESR1.MCF-7_ER_t40 248 bp overlap
ChIP MCF-7_ER_t50 GSE119057.ESR1.MCF-7_ER_t50 255 bp overlap
ChIP MCF-7_ER_t60 GSE119057.ESR1.MCF-7_ER_t60 253 bp overlap
ChIP MCF-7_ER_t70 GSE119057.ESR1.MCF-7_ER_t70 247 bp overlap
ChIP MCF-7_ER_t80 GSE119057.ESR1.MCF-7_ER_t80 250 bp overlap
ChIP MCF-7_ER_t90 GSE119057.ESR1.MCF-7_ER_t90 232 bp overlap
MAX 1 dataset
ChIP H1 ENCFF914VQY 313 bp overlap
MYC 1 dataset
ChIP MCF-7 ENCSR000DMQ.MYC.MCF-7 117 bp overlap
NANOG 1 dataset
ChIP hESC GSE18292.NANOG.hESC 93 bp overlap
RAD21 30 datasets
ChIP GM12878 ENCFF101UQZ 191 bp overlap
ChIP GM12878 ENCSR000BMY.RAD21.GM12878 147 bp overlap
ChIP H1 ENCFF698EWO 165 bp overlap
ChIP H1 ENCFF967OJF 109 bp overlap
ChIP HAP1_clone25 GSE152721.RAD21.HAP1_clone25 202 bp overlap
ChIP HCT-116 GSE131606.RAD21.HCT-116 418 bp overlap
ChIP HEK293_siE1A GSE130135.RAD21.HEK293_siE1A 334 bp overlap
ChIP HUVEC-C_hypoxia GSE94872.RAD21.HUVEC-C_hypoxia 144 bp overlap
ChIP Hep-G2 ENCSR054FKH.RAD21.Hep-G2 120 bp overlap
ChIP HepG2 ENCFF906QIS 225 bp overlap
ChIP K-562 ENCSR000BKV.RAD21.K-562 87 bp overlap
ChIP MCF-7 ENCFF694KOM 291 bp overlap
ChIP MCF-7 ENCSR703TNG.RAD21.MCF-7 197 bp overlap
ChIP MCF-7 ERP000209.RAD21.MCF-7 121 bp overlap
ChIP MCF-7 ENCSR000BTQ.RAD21.MCF-7 121 bp overlap
ChIP MCF-7 GSE72082.RAD21.MCF-7 134 bp overlap
ChIP MCF-7_E2 ERP000209.RAD21.MCF-7_E2 137 bp overlap
ChIP MDM_-dNS1 GSE103477.RAD21.MDM_-dNS1 219 bp overlap
ChIP SK-N-SH ENCFF747MAS 211 bp overlap
ChIP SK-N-SH ENCSR000EHX.RAD21.SK-N-SH 210 bp overlap
ChIP THP-1_Pam3csk-025m-Flavo-000m GSE103477.RAD21.THP-1_Pam3csk-025m-Flavo-000m 183 bp overlap
ChIP THP-1_Pam3csk-180m-Flavo-060m GSE103477.RAD21.THP-1_Pam3csk-180m-Flavo-060m 279 bp overlap
ChIP WA09 GSE105028.RAD21.WA09 231 bp overlap
ChIP WA09_heat-shock GSE105028.RAD21.WA09_heat-shock 256 bp overlap
ChIP hiPSC_CVI GSE106870.RAD21.hiPSC_CVI 127 bp overlap
ChIP hiPSC_IB12 GSE106870.RAD21.hiPSC_IB12 163 bp overlap
ChIP hiPSC_IV8 GSE106870.RAD21.hiPSC_IV8 227 bp overlap
ChIP hiPSC_parental-line GSE106870.RAD21.hiPSC_parental-line 149 bp overlap
ChIP lymphoblast_mut GSE155324.RAD21.lymphoblast_mut 331 bp overlap
ChIP neural ENCSR198ZYJ.RAD21.neural 270 bp overlap
SMC1 1 dataset
ChIP DKO GSE131606.SMC1.DKO 263 bp overlap
SMC3 3 datasets
ChIP HeLa-Kyoto_PDS5-depleted GSE102884.SMC3.HeLa-Kyoto_PDS5-depleted 310 bp overlap
ChIP HeLa-Kyoto_WAPL_PDS-depleted GSE102884.SMC3.HeLa-Kyoto_WAPL_PDS-depleted 199 bp overlap
ChIP neural ENCSR404BPV.SMC3.neural 326 bp overlap
STAG1 6 datasets
ChIP HeLa GSE126990.STAG1.HeLa 285 bp overlap
ChIP HeLa-Kyoto GSE138405.STAG1.HeLa-Kyoto 285 bp overlap
ChIP Hep-G2 ERP000209.STAG1.Hep-G2 161 bp overlap
ChIP Hep-G2 ENCSR167MTG.STAG1.Hep-G2 132 bp overlap
ChIP MCF-7 ERP000209.STAG1.MCF-7 187 bp overlap
ChIP MCF-7_E2 ERP000209.STAG1.MCF-7_E2 195 bp overlap
USF1 1 dataset
ChIP WA01 ENCSR000BIU.USF1.WA01 133 bp overlap
YY1 1 dataset
ChIP WA01 ENCSR000BKD.YY1.WA01 134 bp overlap
ZBTB2 1 dataset
ChIP GM12878 GSE97661.ZBTB2.GM12878 132 bp overlap
ZEB1 1 dataset
ChIP GM12878 ENCSR000BND.ZEB1.GM12878 131 bp overlap
ZNF143 1 dataset
ChIP WA01 ENCSR000EBW.ZNF143.WA01 148 bp overlap
ZNF257 1 dataset
ChIP HEK293T GSE78099.ZNF257.HEK293T 157 bp overlap
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