TSKU
tsukushi, small leucine rich proteoglycan | E2IG4, TSK, LRRC54

Predicted to enable transforming growth factor beta binding activity. Predicted to be involved in several processes, including cholesterol efflux; cholesterol homeostasis; and nervous system development. Predicted to act upstream of or within several processes, including ciliary body morphogenesis; negative regulation of Wnt signaling pathway; and telencephalon development. Predicted to be located in extracellular region. Predicted to be active in extracellular space. [provided by Alliance of Genome Resources, Apr 2025]

Member of: DE-10 DE-10.8 Developmental clusters: GC6
Biological processes 36 terms
anatomical structure morphogenesis (GO:0009653)anterior commissure morphogenesis (GO:0021960)bone growth (GO:0098868)camera-type eye development (GO:0043010)cholesterol efflux (GO:0033344)cholesterol efflux (GO:0033344)cholesterol homeostasis (GO:0042632)cholesterol homeostasis (GO:0042632)cholesterol metabolic process (GO:0008203)cholesterol metabolic process (GO:0008203)ciliary body morphogenesis (GO:0061073)cytokine binding (GO:0019955)energy homeostasis (GO:0097009)energy homeostasis (GO:0097009)extracellular region (GO:0005576)extracellular region (GO:0005576)gel phase of interstitial matrix (GO:0140150)growth plate cartilage chondrocyte development (GO:0003431)hippocampus development (GO:0021766)hippocampus development (GO:0021766)inner ear receptor cell stereocilium organization (GO:0060122)inner ear receptor cell stereocilium organization (GO:0060122)negative regulation of Wnt signaling pathway (GO:0030178)negative regulation of myofibroblast differentiation (GO:1904761)negative regulation of myofibroblast differentiation (GO:1904761)negative regulation of neuron projection development (GO:0010977)negative regulation of neuron projection development (GO:0010977)negative regulation of transforming growth factor beta1 production (GO:0032911)negative regulation of transforming growth factor beta1 production (GO:0032911)positive regulation of hair cycle (GO:0042635)positive regulation of hair cycle (GO:0042635)regulation of gene expression (GO:0010468)transforming growth factor beta binding (GO:0050431)transforming growth factor beta binding (GO:0050431)wound healing (GO:0042060)wound healing (GO:0042060)
Expression (TPM)
TSKU — as a Regulated Gene

TFs regulating TSKU 0 TFs

Transcription factors with Perturb-seq knockdown data for TSKU. The Binding column indicates whether any binding evidence exists for this TF–gene pair (ChIP-seq or motif footprint peaks). The Mean coef is the average Perturb-seq regression coefficient across active gRNAs (positive = TSKU upregulated upon KD; negative = downregulated). The Outlier column indicates whether this gene is in the top or bottom 5% of all TF knockdown effects.

Data: Effect:
TF Mean coef Binding Outlier TF→Gene link

Elements linked to TSKU

Open chromatin peaks (ATAC-seq) in the genomic neighbourhood of TSKU, linked by TSS proximity or chromatin conformation (Multiome / HiCAR). Each element overlaps at least one TF ChIP-seq binding site — the TFs column shows how many distinct TFs bind that element.

Accessibility Element Dist. to TSS Link type TFs
chr11:76,554,474–76,555,062 229.4 kb Distal (>10kb) Multiome 45
chr11:76,624,568–76,625,315 159.1 kb Distal (>10kb) Multiome 105
chr11:76,670,486–76,671,227 113.3 kb Distal (>10kb) Multiome 303
chr11:76,733,962–76,736,172 50.0 kb Distal (>10kb) Multiome HiCAR 353
chr11:76,758,769–76,759,574 24.9 kb Distal (>10kb) Multiome 99
chr11:76,770,356–76,771,447 13.3 kb Distal (>10kb) Multiome 371
chr11:76,781,571–76,782,180 1.2 kb Proximal (<10kb) 189
chr11:76,782,794–76,784,887 63 bp At TSS Multiome 929
chr11:76,785,535–76,785,870 2.2 kb Proximal (<10kb) 52
chr11:76,787,865–76,789,130 4.4 kb Proximal (<10kb) Multiome 462
chr11:76,793,114–76,793,711 9.8 kb Proximal (<10kb) 181
chr11:76,798,746–76,799,610 15.2 kb Distal (>10kb) Multiome 661
chr11:76,807,112–76,807,736 23.4 kb Distal (>10kb) Multiome 373
chr11:76,860,430–76,861,513 76.7 kb Distal (>10kb) Multiome 706
chr11:77,039,527–77,040,269 255.7 kb Distal (>10kb) Multiome 214
chr11:77,066,090–77,067,711 282.7 kb Distal (>10kb) Multiome 664

Genome Browser

Genomic view of the TSKU locus showing ATAC-seq accessibility and RNA-seq expression across the ESC → DE time course, together with TF binding peaks and element-to-TSS loop connections.

chr11:76,544,474 – 77,077,711
Proximal 1 kb Distal 10 kb Multiome HiCAR ATAC-seq RNA-seq