AK4
adenylate kinase 4 | AK3, AK3L1

This gene encodes a member of the adenylate kinase family of enzymes. The encoded protein is localized to the mitochondrial matrix. Adenylate kinases regulate the adenine and guanine nucleotide compositions within a cell by catalyzing the reversible transfer of phosphate group among these nucleotides. Five isozymes of adenylate kinase have been identified in vertebrates. Expression of these isozymes is tissue-specific and developmentally regulated. A pseudogene for this gene has been located on chromosome 17. Three transcript variants encoding the same protein have been identified for this gene. Sequence alignment suggests that the gene defined by NM_013410, NM_203464, and NM_001005353 is located on chromosome 1. [provided by RefSeq, Jul 2008]

Member of: DE-4 Developmental clusters: GC5
Biological processes 40 terms
ADP biosynthetic process (GO:0006172)AMP kinase activity (GO:0004017)AMP kinase activity (GO:0004017)AMP metabolic process (GO:0046033)AMP metabolic process (GO:0046033)AMP metabolic process (GO:0046033)ATP binding (GO:0005524)ATP metabolic process (GO:0046034)ATP metabolic process (GO:0046034)CMP kinase activity (GO:0036430)GTP metabolic process (GO:0046039)GTP metabolic process (GO:0046039)cellular response to hypoxia (GO:0071456)cellular response to hypoxia (GO:0071456)cytoplasm (GO:0005737)dAMP kinase activity (GO:0047506)dCMP kinase activity (GO:0036431)mitochondrial matrix (GO:0005759)mitochondrial matrix (GO:0005759)mitochondrial matrix (GO:0005759)mitochondrial matrix (GO:0005759)mitochondrion (GO:0005739)mitochondrion (GO:0005739)mitochondrion (GO:0005739)nucleobase-containing compound kinase activity (GO:0019205)nucleobase-containing small molecule interconversion (GO:0015949)nucleoside diphosphate kinase activity (GO:0004550)nucleoside diphosphate kinase activity (GO:0004550)nucleoside diphosphate kinase activity (GO:0004550)nucleoside monophosphate kinase activity (GO:0050145)nucleoside monophosphate kinase activity (GO:0050145)nucleoside monophosphate metabolic process (GO:0009123)nucleoside triphosphate adenylate kinase activity (GO:0046899)nucleoside triphosphate adenylate kinase activity (GO:0046899)nucleoside triphosphate biosynthetic process (GO:0009142)nucleotide metabolic process (GO:0009117)phosphotransferase activity, phosphate group as acceptor (GO:0016776)protein binding (GO:0005515)regulation of oxidative phosphorylation (GO:0002082)ribonucleoside diphosphate biosynthetic process (GO:0009188)
Expression (TPM)
AK4 — as a Regulated Gene

TFs regulating AK4 0 TFs

Transcription factors with Perturb-seq knockdown data for AK4. The Binding column indicates whether any binding evidence exists for this TF–gene pair (ChIP-seq or motif footprint peaks). The Mean coef is the average Perturb-seq regression coefficient across active gRNAs (positive = AK4 upregulated upon KD; negative = downregulated). The Outlier column indicates whether this gene is in the top or bottom 5% of all TF knockdown effects.

Data: Effect:
TF Mean coef Binding Outlier TF→Gene link

Elements linked to AK4

Open chromatin peaks (ATAC-seq) in the genomic neighbourhood of AK4, linked by TSS proximity or chromatin conformation (Multiome / HiCAR). Each element overlaps at least one TF ChIP-seq binding site — the TFs column shows how many distinct TFs bind that element.

Accessibility Element Dist. to TSS Link type TFs
chr1:63,574,025–63,574,937 1573.2 kb Distal (>10kb) Multiome HiCAR 44
chr1:64,965,270–64,967,021 181.1 kb Distal (>10kb) Multiome 764
chr1:65,002,269–65,003,167 144.9 kb Distal (>10kb) Multiome 584
chr1:65,065,952–65,068,343 81.1 kb Distal (>10kb) Multiome 865
chr1:65,139,896–65,140,272 7.4 kb Proximal (<10kb) 15
chr1:65,147,108–65,150,062 93 bp At TSS Multiome 912
chr1:65,150,597–65,151,062 2.9 kb Proximal (<10kb) 93
chr1:65,253,859–65,255,712 106.8 kb Distal (>10kb) Multiome 756
chr1:65,265,365–65,266,424 118.2 kb Distal (>10kb) Multiome 602
chr1:65,308,757–65,310,566 162.3 kb Distal (>10kb) Multiome 950
chr1:65,419,304–65,421,455 272.7 kb Distal (>10kb) Multiome 961
chr1:66,923,845–66,926,268 1777.5 kb Distal (>10kb) Multiome HiCAR 945
chr1:66,929,624–66,931,089 1782.6 kb Distal (>10kb) Multiome HiCAR 922

Genome Browser

Genomic view of the AK4 locus showing ATAC-seq accessibility and RNA-seq expression across the ESC → DE time course, together with TF binding peaks and element-to-TSS loop connections.

chr1:63,564,025 – 66,941,089
Proximal 1 kb Distal 10 kb Multiome HiCAR ATAC-seq RNA-seq