CDKN2A
cyclin dependent kinase inhibitor 2A | ARF, CAI2, CDK4I, CMM2, INK4, INK4a, MTS1, P16-INK4A, p14, p14ARF, p16, p16INK4a, p19, p19Arf, CDKN2, MLM

This gene generates several transcript variants which differ in their first exons. At least three alternatively spliced variants encoding distinct proteins have been reported, two of which encode structurally related isoforms known to function as inhibitors of CDK4 kinase. The remaining transcript includes an alternate first exon located 20 Kb upstream of the remainder of the gene; this transcript contains an alternate open reading frame (ARF) that specifies a protein which is structurally unrelated to the products of the other variants. This ARF product functions as a stabilizer of the tumor suppressor protein p53 as it can interact with, and sequester, the E3 ubiquitin-protein ligase MDM2, a protein responsible for the degradation of p53. In spite of the structural and functional differences, the CDK inhibitor isoforms and the ARF product encoded by this gene, through the regulatory roles of CDK4 and p53 in cell cycle G1 progression, share a common functionality in cell cycle G1 control. This gene is frequently mutated or deleted in a wide variety of tumors, and is known to be an important tumor suppressor gene. [provided by RefSeq, Sep 2012]

Biological processes 88 terms
MDM2/MDM4 family protein binding (GO:0097371)NF-kappaB binding (GO:0051059)RNA binding (GO:0003723)RNA polymerase II-specific DNA-binding transcription factor binding (GO:0061629)Ras protein signal transduction (GO:0007265)SUMO transferase activity (GO:0019789)amyloid fibril formation (GO:1990000)amyloid fibril formation (GO:1990000)apoptotic process (GO:0006915)apoptotic signaling pathway (GO:0097190)autophagy of mitochondrion (GO:0000422)cellular senescence (GO:0090398)cellular senescence (GO:0090398)cyclin-dependent protein serine/threonine kinase inhibitor activity (GO:0004861)cyclin-dependent protein serine/threonine kinase inhibitor activity (GO:0004861)cyclin-dependent protein serine/threonine kinase inhibitor activity (GO:0004861)cyclin-dependent protein serine/threonine kinase inhibitor activity (GO:0004861)cytoplasm (GO:0005737)cytoplasm (GO:0005737)cytoplasm (GO:0005737)cytosol (GO:0005829)disordered domain specific binding (GO:0097718)ligase inhibitor activity (GO:0055104)mitochondrial depolarization (GO:0051882)mitochondrial matrix (GO:0005759)mitochondrion (GO:0005739)mitochondrion (GO:0005739)negative regulation of B cell proliferation (GO:0030889)negative regulation of DNA-templated transcription (GO:0045892)negative regulation of cell growth (GO:0030308)negative regulation of cell population proliferation (GO:0008285)negative regulation of cell population proliferation (GO:0008285)negative regulation of cell population proliferation (GO:0008285)negative regulation of cell population proliferation (GO:0008285)negative regulation of cell-matrix adhesion (GO:0001953)negative regulation of cyclin-dependent protein serine/threonine kinase activity (GO:0045736)negative regulation of immature T cell proliferation in thymus (GO:0033088)negative regulation of transcription by RNA polymerase II (GO:0000122)negative regulation of ubiquitin-dependent protein catabolic process (GO:2000059)nuclear body (GO:0016604)nuclear body organization (GO:0030575)nucleolus (GO:0005730)nucleolus (GO:0005730)nucleoplasm (GO:0005654)nucleoplasm (GO:0005654)nucleoplasm (GO:0005654)nucleus (GO:0005634)nucleus (GO:0005634)nucleus (GO:0005634)p53 binding (GO:0002039)positive regulation of DNA damage response, signal transduction by p53 class mediator (GO:0043517)positive regulation of DNA-templated transcription (GO:0045893)positive regulation of apoptotic process (GO:0043065)positive regulation of macrophage apoptotic process (GO:2000111)positive regulation of macrophage apoptotic process (GO:2000111)positive regulation of protein localization to nucleus (GO:1900182)positive regulation of protein sumoylation (GO:0033235)positive regulation of signal transduction by p53 class mediator (GO:1901798)positive regulation of smooth muscle cell apoptotic process (GO:0034393)positive regulation of smooth muscle cell apoptotic process (GO:0034393)positive regulation of transcription by RNA polymerase II (GO:0045944)protein K63-linked ubiquitination (GO:0070534)protein binding (GO:0005515)protein destabilization (GO:0031648)protein kinase binding (GO:0019901)protein localization to nucleolus (GO:1902570)protein localization to nucleus (GO:0034504)protein polyubiquitination (GO:0000209)protein stabilization (GO:0050821)protein stabilization (GO:0050821)protein sumoylation (GO:0016925)protein-containing complex (GO:0032991)regulation of G1/S transition of mitotic cell cycle (GO:2000045)regulation of G1/S transition of mitotic cell cycle (GO:2000045)regulation of G1/S transition of mitotic cell cycle (GO:2000045)regulation of cell cycle (GO:0051726)regulation of cell cycle (GO:0051726)regulation of cell cycle (GO:0051726)regulation of protein export from nucleus (GO:0046825)regulation of protein localization to mitochondrion (GO:1903747)regulation of protein stability (GO:0031647)regulation of protein stability (GO:0031647)regulation of ubiquitin protein ligase activity (GO:1904666)replicative senescence (GO:0090399)senescence-associated heterochromatin focus (GO:0035985)somatic stem cell division (GO:0048103)ubiquitin ligase inhibitor activity (GO:1990948)ubiquitin-protein transferase inhibitor activity (GO:0055105)
Expression (TPM)
CDKN2A — as a Regulated Gene

TFs regulating CDKN2A 0 TFs

Transcription factors with Perturb-seq knockdown data for CDKN2A. The Binding column indicates whether any binding evidence exists for this TF–gene pair (ChIP-seq or motif footprint peaks). The Mean coef is the average Perturb-seq regression coefficient across active gRNAs (positive = CDKN2A upregulated upon KD; negative = downregulated). The Outlier column indicates whether this gene is in the top or bottom 5% of all TF knockdown effects.

Data: Effect:
TF Mean coef Binding Outlier TF→Gene link

Elements linked to CDKN2A

Open chromatin peaks (ATAC-seq) in the genomic neighbourhood of CDKN2A, linked by TSS proximity or chromatin conformation (Multiome / HiCAR). Each element overlaps at least one TF ChIP-seq binding site — the TFs column shows how many distinct TFs bind that element.

Accessibility Element Dist. to TSS Link type TFs
chr9:21,964,802–21,966,024 8.8 kb Proximal (<10kb) 308
chr9:21,966,199–21,966,526 8.3 kb Proximal (<10kb) 38
chr9:21,968,166–21,969,501 5.4 kb Proximal (<10kb) 181
chr9:21,969,634–21,969,867 5.0 kb Proximal (<10kb) 49
chr9:21,970,447–21,971,494 3.4 kb Proximal (<10kb) 93
chr9:21,974,429–21,975,677 at TSS At TSS 502

Genome Browser

Genomic view of the CDKN2A locus showing ATAC-seq accessibility and RNA-seq expression across the ESC → DE time course, together with TF binding peaks and element-to-TSS loop connections.

chr9:21,954,802 – 21,985,677
Proximal 1 kb Distal 10 kb Multiome HiCAR ATAC-seq RNA-seq