SLC15A4
solute carrier family 15 member 4 | PHT1, PTR4

Enables several functions, including L-histidine transmembrane transporter activity; peptide:proton symporter activity; and peptidoglycan transmembrane transporter activity. Involved in dipeptide import across plasma membrane; peptidoglycan transport; and positive regulation of pattern recognition receptor signaling pathway. Located in endolysosome membrane. [provided by Alliance of Genome Resources, Jul 2025]

Biological processes 52 terms
L-histidine transmembrane export from vacuole (GO:0089708)L-histidine transmembrane transporter activity (GO:0005290)L-histidine transmembrane transporter activity (GO:0005290)L-histidine transmembrane transporter activity (GO:0005290)dipeptide import across plasma membrane (GO:0140206)dipeptide import across plasma membrane (GO:0140206)dipeptide transmembrane transporter activity (GO:0071916)dipeptide transmembrane transporter activity (GO:0071916)early endosome membrane (GO:0031901)endolysosome membrane (GO:0036020)endolysosome membrane (GO:0036020)endosome membrane (GO:0010008)histidine transport (GO:0015817)lysosomal membrane (GO:0005765)lysosomal membrane (GO:0005765)lysosomal membrane (GO:0005765)lysosomal membrane (GO:0005765)mast cell homeostasis (GO:0033023)mast cell homeostasis (GO:0033023)membrane (GO:0016020)membrane (GO:0016020)oligopeptide transmembrane transport (GO:0035672)oligopeptide transport (GO:0006857)peptide:proton symporter activity (GO:0015333)peptide:proton symporter activity (GO:0015333)peptide:proton symporter activity (GO:0015333)peptidoglycan transmembrane transporter activity (GO:0015647)peptidoglycan transport (GO:0015835)peptidoglycan transport (GO:0015835)plasma membrane (GO:0005886)positive regulation of innate immune response (GO:0045089)positive regulation of nucleotide-binding oligomerization domain containing 1 signaling pathway (GO:0070430)positive regulation of nucleotide-binding oligomerization domain containing 1 signaling pathway (GO:0070430)positive regulation of nucleotide-binding oligomerization domain containing 2 signaling pathway (GO:0070434)positive regulation of nucleotide-binding oligomerization domain containing 2 signaling pathway (GO:0070434)positive regulation of toll-like receptor 7 signaling pathway (GO:0034157)positive regulation of toll-like receptor 7 signaling pathway (GO:0034157)positive regulation of toll-like receptor 7 signaling pathway (GO:0034157)positive regulation of toll-like receptor 8 signaling pathway (GO:0034161)positive regulation of toll-like receptor 8 signaling pathway (GO:0034161)positive regulation of toll-like receptor 9 signaling pathway (GO:0034165)positive regulation of toll-like receptor 9 signaling pathway (GO:0034165)positive regulation of toll-like receptor 9 signaling pathway (GO:0034165)protein binding (GO:0005515)proton transmembrane transport (GO:1902600)regulation of isotype switching to IgG isotypes (GO:0048302)regulation of isotype switching to IgG isotypes (GO:0048302)regulation of nucleotide-binding domain, leucine rich repeat containing receptor signaling pathway (GO:0070424)regulation of nucleotide-binding domain, leucine rich repeat containing receptor signaling pathway (GO:0070424)specific granule membrane (GO:0035579)transmembrane transport (GO:0055085)transmembrane transporter activity (GO:0022857)
Expression (TPM)
SLC15A4 — as a Regulated Gene

TFs regulating SLC15A4 0 TFs

Transcription factors with Perturb-seq knockdown data for SLC15A4. The Binding column indicates whether any binding evidence exists for this TF–gene pair (ChIP-seq or motif footprint peaks). The Mean coef is the average Perturb-seq regression coefficient across active gRNAs (positive = SLC15A4 upregulated upon KD; negative = downregulated). The Outlier column indicates whether this gene is in the top or bottom 5% of all TF knockdown effects.

Data: Effect:
TF Mean coef Binding Outlier TF→Gene link

Elements linked to SLC15A4

Open chromatin peaks (ATAC-seq) in the genomic neighbourhood of SLC15A4, linked by TSS proximity or chromatin conformation (Multiome / HiCAR). Each element overlaps at least one TF ChIP-seq binding site — the TFs column shows how many distinct TFs bind that element.

Accessibility Element Dist. to TSS Link type TFs
chr12:128,823,117–128,824,931 22 bp At TSS Multiome 840
chr12:128,853,166–128,854,788 30.3 kb Distal (>10kb) Multiome HiCAR 293
chr12:128,856,024–128,856,722 32.4 kb Distal (>10kb) Multiome HiCAR 116
chr12:128,858,379–128,859,438 35.0 kb Distal (>10kb) Multiome HiCAR 60
chr12:128,969,259–128,970,072 145.6 kb Distal (>10kb) Multiome 192
chr12:129,871,727–129,873,060 1048.3 kb Distal (>10kb) Multiome HiCAR 258
chr12:129,902,870–129,904,853 1080.3 kb Distal (>10kb) Multiome HiCAR 260
chr12:130,013,812–130,014,266 1190.2 kb Distal (>10kb) Multiome HiCAR 158
chr12:130,716,209–130,717,026 1892.6 kb Distal (>10kb) Multiome HiCAR 196

Genome Browser

Genomic view of the SLC15A4 locus showing ATAC-seq accessibility and RNA-seq expression across the ESC → DE time course, together with TF binding peaks and element-to-TSS loop connections.

chr12:128,813,117 – 130,727,026
Proximal 1 kb Distal 10 kb Multiome HiCAR ATAC-seq RNA-seq