IL4
interleukin 4 | BCGF-1, BCGF1, BSF1, IL-4, MGC79402

The protein encoded by this gene is a pleiotropic cytokine produced by activated T cells. This cytokine is a ligand for interleukin 4 receptor. The interleukin 4 receptor also binds to IL13, which may contribute to many overlapping functions of this cytokine and IL13. STAT6, a signal transducer and activator of transcription, has been shown to play a central role in mediating the immune regulatory signal of this cytokine. This gene, IL3, IL5, IL13, and CSF2 form a cytokine gene cluster on chromosome 5q, with this gene particularly close to IL13. This gene, IL13 and IL5 are found to be regulated coordinately by several long-range regulatory elements in an over 120 kilobase range on the chromosome. IL4 is considered an important cytokine for tissue repair, counterbalancing the effects of proinflammatory type 1 cytokines, however, it also promotes allergic airway inflammation. Moreover, IL-4, a type 2 cytokine, mediates and regulates a variety of human host responses such as allergic, anti-parasitic, wound healing, and acute inflammation. This cytokine has been reported to promote resolution of neutrophil-mediated acute lung injury. In an allergic response, IL-4 has an essential role in the production of allergen-specific immunoglobin (Ig) E. This pro-inflammatory cytokine has been observed to be increased in COVID-19 (Coronavirus disease 2019) patients, but is not necessarily associated with severe COVID-19 pathology. Two alternatively spliced transcript variants of this gene encoding distinct isoforms have been reported. [provided by RefSeq, Aug 2020]

Biological processes 68 terms
B cell differentiation (GO:0030183)T cell activation (GO:0042110)cell surface receptor signaling pathway via JAK-STAT (GO:0007259)cell surface receptor signaling pathway via JAK-STAT (GO:0007259)cholesterol metabolic process (GO:0008203)cytokine activity (GO:0005125)cytokine receptor binding (GO:0005126)dendritic cell differentiation (GO:0097028)extracellular region (GO:0005576)extracellular region (GO:0005576)extracellular region (GO:0005576)growth factor activity (GO:0008083)growth factor activity (GO:0008083)immune response (GO:0006955)immune response (GO:0006955)interleukin-4 receptor binding (GO:0005136)interleukin-4 receptor binding (GO:0005136)interleukin-4-mediated signaling pathway (GO:0035771)interleukin-4-mediated signaling pathway (GO:0035771)interleukin-4-mediated signaling pathway (GO:0035771)leukocyte activation (GO:0045321)macrophage activation (GO:0042116)myeloid dendritic cell differentiation (GO:0043011)negative regulation of DNA-templated transcription (GO:0045892)negative regulation of apoptotic process (GO:0043066)negative regulation of cellular response to transforming growth factor beta stimulus (GO:1903845)negative regulation of complement-dependent cytotoxicity (GO:1903660)negative regulation of endothelial cell apoptotic process (GO:2000352)negative regulation of epithelial cell migration (GO:0010633)negative regulation of inflammatory response (GO:0050728)negative regulation of inflammatory response (GO:0050728)negative regulation of neuroinflammatory response (GO:0150079)negative regulation of osteoclast differentiation (GO:0045671)negative regulation of transcription by RNA polymerase II (GO:0000122)negative regulation of tumor necrosis factor production (GO:0032720)neuroinflammatory response (GO:0150076)positive regulation of ATP biosynthetic process (GO:2001171)positive regulation of B cell proliferation (GO:0030890)positive regulation of DNA-templated transcription (GO:0045893)positive regulation of DNA-templated transcription (GO:0045893)positive regulation of MHC class II biosynthetic process (GO:0045348)positive regulation of T cell differentiation (GO:0045582)positive regulation of T cell proliferation (GO:0042102)positive regulation of T-helper 2 cell cytokine production (GO:2000553)positive regulation of amyloid-beta clearance (GO:1900223)positive regulation of cell migration (GO:0030335)positive regulation of cell migration (GO:0030335)positive regulation of cell population proliferation (GO:0008284)positive regulation of cellular respiration (GO:1901857)positive regulation of cold-induced thermogenesis (GO:0120162)positive regulation of gene expression (GO:0010628)positive regulation of gene expression (GO:0010628)positive regulation of interleukin-10 production (GO:0032733)positive regulation of interleukin-13 production (GO:0032736)positive regulation of isotype switching to IgE isotypes (GO:0048295)positive regulation of isotype switching to IgG isotypes (GO:0048304)positive regulation of leukocyte differentiation (GO:1902107)positive regulation of macroautophagy (GO:0016239)positive regulation of receptor-mediated endocytosis (GO:0048260)positive regulation of transcription by RNA polymerase II (GO:0045944)positive regulation of transcription by RNA polymerase II (GO:0045944)positive regulation of tyrosine phosphorylation of STAT protein (GO:0042531)protein binding (GO:0005515)regulation of immune response (GO:0050776)regulation of immune response (GO:0050776)regulation of isotype switching (GO:0045191)regulation of multicellular organismal process (GO:0051239)type 2 immune response (GO:0042092)
Expression (TPM)
IL4 — as a Regulated Gene

TFs regulating IL4 0 TFs

Transcription factors with Perturb-seq knockdown data for IL4. The Binding column indicates whether any binding evidence exists for this TF–gene pair (ChIP-seq or motif footprint peaks). The Mean coef is the average Perturb-seq regression coefficient across active gRNAs (positive = IL4 upregulated upon KD; negative = downregulated). The Outlier column indicates whether this gene is in the top or bottom 5% of all TF knockdown effects.

Data: Effect:
TF Mean coef Binding Outlier TF→Gene link

Elements linked to IL4

Open chromatin peaks (ATAC-seq) in the genomic neighbourhood of IL4, linked by TSS proximity or chromatin conformation (Multiome / HiCAR). Each element overlaps at least one TF ChIP-seq binding site — the TFs column shows how many distinct TFs bind that element.

Accessibility Element Dist. to TSS Link type TFs
chr5:132,673,915–132,674,420 at TSS At TSS 62

Genome Browser

Genomic view of the IL4 locus showing ATAC-seq accessibility and RNA-seq expression across the ESC → DE time course, together with TF binding peaks and element-to-TSS loop connections.

chr5:132,663,915 – 132,684,420
Proximal 1 kb Distal 10 kb Multiome HiCAR ATAC-seq RNA-seq